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<document id="A73E2E1810D0D9D2DCE6E3670897C574" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A30BB16FEFEE432E55B2D9F" docLanguage="en" docName="J_NAT_HIST.53.37–38.2231–2351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) arnaudi Nunes &amp; Vaz-de-Mello 2019, new species" docType="treatment" docVersion="18" lastPageNumber="2313" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2312" updateTime="1698782751744" updateUser="ExternalLinkService">
<mods:mods id="70BC2989E2CB8EAB4579B636105A8929" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2B1F5588C6D64632E06B8F1133D888C9">
<mods:title id="7A215958A7B6D43801800470EB6EFAE8">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
</mods:titleInfo>
<mods:name id="7B89D195C9AD8B128E75062DB0110CD5" type="personal">
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<mods:namePart id="171384799E6ABD8BCAFA86A542B18D9F">Nunes, Rafael V.</mods:namePart>
</mods:name>
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<mods:namePart id="BBB12D160035BB6C989086468D831046">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="4D7F7B6EAC9D507A3D93651C8656B035">text</mods:typeOfResource>
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<mods:title id="2264E59B1C8BFC959FDAD367E2B12977">Journal of Natural History</mods:title>
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<mods:part id="56BE590D4E26D067F2B99128208E592C">
<mods:date id="450713E5000558C3C4A4D472C18E34D0">2019</mods:date>
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<mods:number id="555C2A8C32B323C1247B632D07BDFFC2">2020-01-13</mods:number>
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<mods:number id="E06B74A4AB4BE1F334B94FD972D2BAC0">53</mods:number>
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<mods:number id="04E802D760B03D509A1C63452470A772">37</mods:number>
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<mods:classification id="12D131254F292DE64B63EE54035C994E">journal article</mods:classification>
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<mods:identifier id="5A8E686055F3C100192B65A1693BC5D6" type="DOI">10.1080/00222933.2019.1692088</mods:identifier>
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<treatment id="C740D6092A30BB16FEFEE432E55B2D9F" ID-DOI="http://doi.org/10.5281/zenodo.3671908" ID-GBIF-Taxon="162116281" ID-Zenodo-Dep="3671908" LSID="urn:lsid:plazi:treatment:C740D6092A30BB16FEFEE432E55B2D9F" httpUri="http://treatment.plazi.org/id/C740D6092A30BB16FEFEE432E55B2D9F" lastPageId="83" lastPageNumber="2313" pageId="82" pageNumber="2312">
<subSubSection id="07F334942A30BB17FEFEE432E6802DAB" pageId="82" pageNumber="2312" type="nomenclature">
<paragraph id="4F56671F2A30BB17FEFEE432E7AB2D46" blockId="82.[348,967,332,392]" box="[348,967,332,357]" pageId="82" pageNumber="2312">
<heading id="141ED0732A30BB17FEFEE432E7AB2D46" box="[348,967,332,357]" centered="true" fontSize="11" level="2" pageId="82" pageNumber="2312" reason="2">
3.7.:
<taxonomicName id="88E91C9C2A30BB17FE30E432E7442D46" ID-CoL="35N9J" authority="Nunes &amp; Vaz-de-Mello, 2019" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[402,808,332,357]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="2312" phylum="Arthropoda" rank="species" species="arnaudi" status="sp. nov." subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A30BB17FE30E432E7442D46" box="[402,808,332,357]" italics="true" pageId="82" pageNumber="2312">Dichotomius (Cephagonus) arnaudi</emphasis>
</taxonomicName>
<emphasis id="7D9DBB0D2A30BB17FC92E432E7AB2D46" bold="true" box="[816,967,332,357]" pageId="82" pageNumber="2312">
<taxonomicNameLabel id="66AE06762A30BB17FC92E432E7AB2D46" box="[816,967,332,357]" pageId="82" pageNumber="2312" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4F56671F2A30BB17FD95E411E6802DAB" blockId="82.[348,967,332,392]" box="[567,748,367,392]" pageId="82" pageNumber="2312">
<heading id="141ED0732A30BB17FD95E411E6802DAB" box="[567,748,367,392]" centered="true" fontSize="11" level="2" pageId="82" pageNumber="2312" reason="5">
(
<figureCitation id="D7D27B9A2A30BB17FD9DE411E6C32DAB" box="[575,687,367,392]" captionStart="Figure 33" captionStartId="48.[160,225,1500,1523]" captionTargetBox="[163,1153,142,1469]" captionTargetId="figure@48.[163,1153,142,1469]" captionTargetPageId="48" captionText="Figure 33. D. arnaudi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view (grey arrow indicates the position of longitudinal excavation on lateral view). D. asenjoi: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. barbarae (i) male dorsal habitus; (j) paramera dorsal view (black arrow indicating slight medial constriction); (k) paramera ventral view; (l) paramera lateral view (black arrow indicates the surface of longitudinal excavation, which is smooth with fine punctures (see D. paschoali) and the differentiation of both species)." figureDoi="http://doi.org/10.5281/zenodo.3665581" httpUri="https://zenodo.org/record/3665581/files/figure.png" pageId="82" pageNumber="2312">Figure 33</figureCitation>
(a-d))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07F334942A30BB17FF02E4B8E06D2E8C" pageId="82" pageNumber="2312" type="diagnosis">
<paragraph id="4F56671F2A30BB17FF02E4B8E06D2E8C" blockId="82.[160,1156,454,687]" pageId="82" pageNumber="2312">
<emphasis id="7D9DBB0D2A30BB17FF02E4B8E5762DFC" bold="true" box="[160,282,454,479]" italics="true" pageId="82" pageNumber="2312">Diagnosis.</emphasis>
this species is separated from other in the group by the following combined characters: males pronotal disc simply convex with a most central knob on anterior portion (as
<taxonomicName id="88E91C9C2A30BB17FE8EE772E5E52E07" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[300,393,524,548]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="2312" phylum="Arthropoda" rank="species" species="emas">
<emphasis id="7D9DBB0D2A30BB17FE8EE772E5E52E07" box="[300,393,524,548]" italics="true" pageId="82" pageNumber="2312">D. emas</emphasis>
</taxonomicName>
); ventrite glabrous and lacking distinct ocellate puncture along anterior margins; males cephalic process produced in a single tubercle (as
<taxonomicName id="88E91C9C2A30BB17FC5BE751E0EF2E64" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[1017,1155,559,583]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="2312" phylum="Arthropoda" rank="species" species="edmondsi">
<emphasis id="7D9DBB0D2A30BB17FC5BE751E0EF2E64" box="[1017,1155,559,583]" italics="true" pageId="82" pageNumber="2312">D. edmondsi</emphasis>
</taxonomicName>
and smaller specimens of
<emphasis id="7D9DBB0D2A30BB17FE63E72FE6D62E4A" box="[449,698,593,617]" italics="true" pageId="82" pageNumber="2312">
<taxonomicName id="88E91C9C2A30BB17FE63E72FE65A2E4A" box="[449,566,593,617]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="2312" phylum="Arthropoda" rank="species" species="bicuspis">D. bicuspis</taxonomicName>
,
<taxonomicName id="88E91C9C2A30BB17FDE2E72FE6D62E4A" authorityName="Haroldi" authorityYear="1869" box="[576,698,593,617]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="2312" phylum="Arthropoda" rank="species" species="ascanius">D. ascanius</taxonomicName>
</emphasis>
and
<taxonomicName id="88E91C9C2A30BB17FD50E72FE7392E4A" authorityName="Harold" authorityYear="1869" box="[754,853,593,617]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="2312" phylum="Arthropoda" rank="species" species="reichei">
<emphasis id="7D9DBB0D2A30BB17FD50E72FE7392E4A" box="[754,853,593,617]" italics="true" pageId="82" pageNumber="2312">D. reichei</emphasis>
</taxonomicName>
); genae borders divergentı posterior angle rounded; paramera having fine carina delimiting basal and longitudinal excavations and sides of elytra sub-parallel in dorsal view (as in
<taxonomicName id="88E91C9C2A30BB17FCD3E7E9E7982E8C" box="[881,1012,663,687]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="2312" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="7D9DBB0D2A30BB17FCD3E7E9E7982E8C" box="[881,1012,663,687]" italics="true" pageId="82" pageNumber="2312">D. clypeatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A30BB17FF1CE7A3E5792F5C" pageId="82" pageNumber="2312" type="materials_examined">
<paragraph id="4F56671F2A30BB17FF1CE7A3E5412F34" blockId="82.[190,1156,732,895]" pageId="82" pageNumber="2312">
<emphasis id="7D9DBB0D2A30BB17FF1CE7A3E4A02ED2" bold="true" box="[190,204,733,753]" pageId="82" pageNumber="2312">●</emphasis>
<materialsCitation id="FF816D422A30BB17FF7AE7A2E5412F34" ID-GBIF-Occurrence="2570430616" collectingDate="1991-12" collectionCode="CEMT" country="Bolivia" location="Arnaud" municipality="Nor Yungas. Caranavari / Carrasco" pageId="82" pageNumber="2312" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="9052D9BD2A30BB17FF7AE7A2E53E2ED7" box="[216,338,732,756]" pageId="82" pageNumber="2312" type="holotype">HOLOTYPE</typeStatus>
(
<specimenCount id="59EFAC962A30BB17FEC1E7A2E5F22ED7" box="[355,414,732,756]" pageId="82" pageNumber="2312" type="male">male</specimenCount>
):
<collectingCountry id="37FE278F2A30BB17FE16E7A2E67C2ED7" box="[436,528,732,756]" name="Bolivia" pageId="82" pageNumber="2312">BOLIVIA</collectingCountry>
:
<collectingMunicipality id="AF32FD652A30BB17FDBCE7A2E7CD2ED7" box="[542,929,732,756]" pageId="82" pageNumber="2312">
<emphasis id="7D9DBB0D2A30BB17FDBCE7A2E6C12ED7" bold="true" box="[542,685,732,756]" pageId="82" pageNumber="2312">Nor Yungas</emphasis>
. Caranavari/Carrasco
</collectingMunicipality>
.
<date id="3B5741DF2A30BB17FC0DE7A2E0642ED7" box="[943,1032,732,756]" pageId="82" pageNumber="2312" value="1991-12">
<collectingDate id="2B13B8372A30BB17FC0DE7A2E0642ED7" box="[943,1032,732,756]" pageId="82" pageNumber="2312" value="1991-12">xii-1991</collectingDate>
</date>
. P
<location id="4A3631C42A30BB17FB93E7A2E0EF2ED7" LSID="urn:lsid:plazi:treatment:C740D6092A30BB16FEFEE432E55B2D9F:4A3631C42A30BB17FB93E7A2E0EF2ED7" box="[1073,1155,732,756]" country="Bolivia" municipality="Nor Yungas. Caranavari / Carrasco" name="Arnaud" pageId="82" pageNumber="2312">Arnaud</location>
[
<collectionCode id="29F8FFDA2A30BB17FF7FE781E5492F34" box="[221,293,767,791]" pageId="82" pageNumber="2312">CEMT</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph id="4F56671F2A30BB17FF1CE65CE5792F5C" blockId="82.[190,1156,732,895]" pageId="82" pageNumber="2312">
<emphasis id="7D9DBB0D2A30BB17FF1CE65CE4A02F15" bold="true" box="[190,204,802,822]" pageId="82" pageNumber="2312">●</emphasis>
<typeStatus id="9052D9BD2A30BB17FF7AE65FE5302F1A" box="[216,348,801,825]" pageId="82" pageNumber="2312" type="paratype">PARATYPES</typeStatus>
: same data of
<typeStatus id="9052D9BD2A30BB17FDB4E65FE6112F1A" box="[534,637,801,825]" pageId="82" pageNumber="2312" type="holotype">Holotype</typeStatus>
[
<specimenCount id="59EFAC962A30BB17FD32E65FE6B92F1A" box="[656,725,801,825]" pageId="82" pageNumber="2312" type="female">2 ♀♀</specimenCount>
at
<collectionCode id="29F8FFDA2A30BB17FCA7E65FE7262F1A" box="[773,842,801,825]" pageId="82" pageNumber="2312">CEMT</collectionCode>
].
<collectingCountry id="37FE278F2A30BB17FCFCE65FE7F02F1A" box="[862,924,801,825]" name="Peru" pageId="82" pageNumber="2312">PERU</collectingCountry>
:
<emphasis id="7D9DBB0D2A30BB17FC0EE65FE79F2F1A" bold="true" box="[940,1011,801,825]" pageId="82" pageNumber="2312">
<collectingRegion id="8D2DA9FD2A30BB17FC0EE65FE79F2F1A" box="[940,1011,801,825]" country="Peru" name="Cusco" pageId="82" pageNumber="2312">Cuzco</collectingRegion>
</emphasis>
. San Pedro. 13³03
<emphasis id="7D9DBB0D2A30BB17FEBAE63AE5722F7F" bold="true" box="[280,286,836,860]" italics="true" pageId="82" pageNumber="2312">ʹ</emphasis>
15
<emphasis id="7D9DBB0D2A30BB17FE9BE63AE5292F7F" box="[313,325,836,860]" italics="true" pageId="82" pageNumber="2312">
<emphasis id="7D9DBB0D2A30BB17FE9BE63AE5532F7F" bold="true" box="[313,319,836,860]" italics="true" pageId="82" pageNumber="2312">ʹ</emphasis>
’
</emphasis>
Sı 71³32
<emphasis id="7D9DBB0D2A30BB17FE02E63AE5CA2F7F" bold="true" box="[416,422,836,860]" italics="true" pageId="82" pageNumber="2312">ʹ</emphasis>
54
<emphasis id="7D9DBB0D2A30BB17FE63E63AE5A12F7F" box="[449,461,836,860]" italics="true" pageId="82" pageNumber="2312">
<emphasis id="7D9DBB0D2A30BB17FE63E63AE5AB2F7F" bold="true" box="[449,455,836,860]" italics="true" pageId="82" pageNumber="2312">ʹ</emphasis>
’
</emphasis>
Wı
<quantity id="8811CAFA2A30BB17FE53E63AE6242F7F" box="[497,584,836,860]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="82" pageNumber="2312" unit="m" value="1500.0">1500 m</quantity>
. Cloub Forest. FIT.
<date id="3B5741DF2A30BB17FC83E63AE7192F7F" box="[801,885,836,860]" pageId="82" pageNumber="2312" value="1999-11">xi-1999</date>
.
<collectionCode id="29F8FFDA2A30BB17FC22E63AE7E12F7F" box="[896,909,836,860]" name="Tavera, Department of Geology and Geophysics" pageId="82" pageNumber="2312">T</collectionCode>
Larsen [1 Ƌı
<specimenCount id="59EFAC962A30BB17FB85E63AE0092F7F" box="[1063,1125,836,860]" pageId="82" pageNumber="2312" type="female">2 ♀♀</specimenCount>
at TLpc]
</paragraph>
</subSubSection>
<subSubSection id="07F334942A30BB16FF02E6D5E6912D2D" lastPageId="83" lastPageNumber="2313" pageId="82" pageNumber="2312" type="description">
<paragraph id="4F56671F2A30BB17FF02E6D5E5ED2A31" blockId="82.[160,1156,939,1554]" pageId="82" pageNumber="2312">
<emphasis id="7D9DBB0D2A30BB17FF02E6D5E5C12FE7" bold="true" box="[160,429,939,964]" italics="true" pageId="82" pageNumber="2312">
Description (
<typeStatus id="9052D9BD2A30BB17FE95E6D5E5F32FE7" box="[311,415,939,964]" pageId="82" pageNumber="2312" type="holotype">holotype</typeStatus>
).
</emphasis>
male: BL: 15 mmı PW:
<quantity id="8811CAFA2A30BB17FD10E6D2E69A2FE7" box="[690,758,940,964]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="82" pageNumber="2312" unit="mm" value="8.0">8 mm</quantity>
. Colour: dark brown to blackı shiny.
<emphasis id="7D9DBB0D2A30BB17FF02E6B1E4B22FC4" bold="true" box="[160,222,975,999]" pageId="82" pageNumber="2312">Head</emphasis>
: fronto-clypeal surface smooth with few lateral striations along clypeo-genal groove. Cephalic horn produced in a single central tubercle.
<emphasis id="7D9DBB0D2A30BB17FCF4E68CE7A22829" bold="true" box="[854,974,1010,1034]" pageId="82" pageNumber="2312">Pronotum</emphasis>
: simply convexı with a weak knob on anterior portion. Ocellate punctures present on posterior angles. Pronotal disc smoothı shinyı with very fine punctures (viewed under 20x magnification). Anterior portionı adjacent to eyes with coarse weakly impressed punctures.
<emphasis id="7D9DBB0D2A30BB17FF02E102E52C28B7" bold="true" box="[160,320,1148,1172]" pageId="82" pageNumber="2312">Metasternum</emphasis>
: sides and margins of anterior lobe covered by long hair. Anterior portion of anterior lobe glabrous. Setigerous punctures equally spaced at sides.
<emphasis id="7D9DBB0D2A30BB17FC53E1E1E05A2894" bold="true" box="[1009,1078,1183,1207]" pageId="82" pageNumber="2312">Elytra</emphasis>
: striae deeply impressed having fine punctures spaced by twice their diameter. Interstriae convexı disc smooth with very fine punctures (viewed under 20x magnification).
<emphasis id="7D9DBB0D2A30BB17FF02E079E579293C" bold="true" box="[160,277,1287,1311]" pageId="82" pageNumber="2312">Abdomen</emphasis>
: ventrites glabrous and lacking rows ocellate punctures along anterior margin. Groove along the posterior margin of sixth ventrite complete but not concealing the ventrite.
<emphasis id="7D9DBB0D2A30BB17FEABE032E5132947" bold="true" box="[265,383,1356,1380]" pageId="82" pageNumber="2312">Aedeagus</emphasis>
(
<figureCitation id="D7D27B9A2A30BB17FE30E032E66F2947" box="[402,515,1356,1380]" captionStart="Figure 33" captionStartId="48.[160,225,1500,1523]" captionTargetBox="[163,1153,142,1469]" captionTargetId="figure@48.[163,1153,142,1469]" captionTargetPageId="48" captionText="Figure 33. D. arnaudi: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view (grey arrow indicates the position of longitudinal excavation on lateral view). D. asenjoi: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view. D. barbarae (i) male dorsal habitus; (j) paramera dorsal view (black arrow indicating slight medial constriction); (k) paramera ventral view; (l) paramera lateral view (black arrow indicates the surface of longitudinal excavation, which is smooth with fine punctures (see D. paschoali) and the differentiation of both species)." figureDoi="http://doi.org/10.5281/zenodo.3665581" httpUri="https://zenodo.org/record/3665581/files/figure.png" pageId="82" pageNumber="2312">Figure 33</figureCitation>
(b-d)): Dorsallyı paramere apexes roundedı convergent; longitudinal excavation present taking the whole paramera length; basal-rounded excavation present; fine continuous carina present along paramera delimiting basal and longitudinal excavations. Laterallyı subtle declivity from basis towards apex. Ventrallyı basis with wide excavationı lacing sclerotised portions; subgenital plate emarginated both basally and apically.
</paragraph>
<paragraph id="4F56671F2A30BB16FF02E340E6912D2D" blockId="82.[160,1156,1598,1727]" lastBlockId="83.[160,1155,142,270]" lastPageId="83" lastPageNumber="2313" pageId="82" pageNumber="2312">
<emphasis id="7D9DBB0D2A30BB17FF02E340E5A92A74" bold="true" box="[160,453,1598,1623]" italics="true" pageId="82" pageNumber="2312">Morphological variation.</emphasis>
males BL: 13
<emphasis id="7D9DBB0D2A30BB17FDFCE341E6072A74" box="[606,619,1599,1623]" italics="true" pageId="82" pageNumber="2312">–</emphasis>
15 mmı PW: 7
<emphasis id="7D9DBB0D2A30BB17FCA1E341E77C2A74" box="[771,784,1599,1623]" italics="true" pageId="82" pageNumber="2312">–</emphasis>
8. Females BL: 15 mmı PW:
<quantity id="8811CAFA2A30BB17FB9FE341E0EC2A74" box="[1085,1152,1599,1623]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="82" pageNumber="2312" unit="mm" value="8.0">8 mm</quantity>
. Distinguished from males as follows: Head: fronto-clypeal surface striated. Few coarse puncture present posterior to cephalic carina near both eyes. Cephalic carina transversalı forming a weak pair of tubercles. Pronotum: simply convexı lacking lobes. Deeply impressed coarse punctures near anterior angles. Elytra: striae weakly impressedı punctures separated by one to twice their diameter. Interstriae convexı disc microsculpture chagrinated. Blue sheen present near basis. Abdomen: sixth ventrite three times larger than fifth and bearing a weakly central-rounded lobe.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A31BB16FF02E445E77A2D77" box="[160,790,315,340]" pageId="83" pageNumber="2313" type="distribution">
<paragraph id="4F56671F2A31BB16FF02E445E77A2D77" blockId="83.[160,790,315,340]" box="[160,790,315,340]" pageId="83" pageNumber="2313">
<emphasis id="7D9DBB0D2A31BB16FF02E445E55F2D77" bold="true" box="[160,307,315,340]" italics="true" pageId="83" pageNumber="2313">Distribution.</emphasis>
Peruvian and Bolivian Yungas (
<figureCitation id="D7D27B9A2A31BB16FD3DE442E7652D77" box="[671,777,316,340]" captionStart="Figure 34" captionStartId="49.[160,225,1006,1029]" captionTargetBox="[162,1151,142,975]" captionTargetId="figure@49.[162,1153,142,975]" captionTargetPageId="49" captionText="Figure 34. Distribution map of D. gardneriı D. aielloaeı D. asenjoiı D. arnaudiı D. Clypeatus and D. edmondsi." figureDoi="http://doi.org/10.5281/zenodo.3665583" httpUri="https://zenodo.org/record/3665583/files/figure.png" pageId="83" pageNumber="2313">Figure 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A31BB16FF02E4FEE55B2D9F" pageId="83" pageNumber="2313" type="etymology">
<paragraph id="4F56671F2A31BB16FF02E4FEE55B2D9F" blockId="83.[160,1155,384,444]" pageId="83" pageNumber="2313">
<emphasis id="7D9DBB0D2A31BB16FF02E4FEE54F2DBA" bold="true" box="[160,291,384,409]" italics="true" pageId="83" pageNumber="2313">Etymology.</emphasis>
named after Patrick Arnaudı entomologist from
<collectingCountry id="37FE278F2A31BB16FCF9E4FFE7CA2DBA" box="[859,934,385,409]" name="France" pageId="83" pageNumber="2313">France</collectingCountry>
and who collected the
<typeStatus id="9052D9BD2A31BB16FF6EE4DAE55F2D9F" box="[204,307,420,444]" pageId="83" pageNumber="2313" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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