175 lines
19 KiB
XML
175 lines
19 KiB
XML
<document id="A6AD53F0D7295AC760639ECC20028011" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. & Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A0ABB2CFE94E741E5722E65" docLanguage="en" docName="J_NAT_HIST.53.37–38.2231–2351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) lucianomourai Nunes & Vaz-de-Mello 2019, new species" docType="treatment" docVersion="18" lastPageNumber="2335" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2334" updateTime="1698782751744" updateUser="ExternalLinkService">
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<mods:titleInfo id="8C8C0DD0C6D56252645FC131CB2D9E2F">
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<mods:title id="760B8CF55C3BA1D379529FBFAEA249F3">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
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<mods:name id="A99E7BA8FF8320F6865DC5537CC1B5C7" type="personal">
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<mods:roleTerm id="493E2CF34A8433DC8DEF11052FE087BC">Author</mods:roleTerm>
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<mods:namePart id="61CCBA7B6921B907A4638A2E5D6BAE9C">Nunes, Rafael V.</mods:namePart>
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<mods:name id="F59051447406A6B16DE057D2B045901E" type="personal">
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<mods:roleTerm id="49609CAD3D4220461F6F0A0C05990588">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AEA8C6590E13175C4DEC2B9FB4E41317">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="405D6DFC6714B08A0D26DDD78A6EAC29">text</mods:typeOfResource>
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<mods:title id="C42374A19707CFAF4EE65F2A7CA6F4EA">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="68912CD130E3E84893AB945E4F60E515">
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<mods:date id="006D15FBC5E147B5D67A954D4FC405B1">2019</mods:date>
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<mods:number id="F06E244534EDBAEBABB91A474AADBC1D">2020-01-13</mods:number>
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<mods:detail id="3C059620C8A32FCEC06140835949B325" type="volume">
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<mods:number id="F38A3B267EEA4E33BF7547DE49FEA4DB">53</mods:number>
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<mods:detail id="8736FDF11F1255A320D69EB8F6D83855" type="issue">
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<mods:number id="52C1736BA09A11A38A3DC43A2ADE0CFA">37</mods:number>
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<mods:classification id="FAB674BC7AAE7EBBACAE2E87D39322CD">journal article</mods:classification>
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<mods:identifier id="53A5C00A244268C4472BF11DD2059694" type="DOI">10.1080/00222933.2019.1692088</mods:identifier>
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<mods:identifier id="BCA3F5B5E8852B04F1AAAE67E727B3AA" type="ISSN">1464-5262</mods:identifier>
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<treatment id="C740D6092A0ABB2CFE94E741E5722E65" ID-DOI="http://doi.org/10.5281/zenodo.3671879" ID-GBIF-Taxon="162253023" ID-Zenodo-Dep="3671879" LSID="urn:lsid:plazi:treatment:C740D6092A0ABB2CFE94E741E5722E65" httpUri="http://treatment.plazi.org/id/C740D6092A0ABB2CFE94E741E5722E65" lastPageId="105" lastPageNumber="2335" pageId="104" pageNumber="2334">
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<subSubSection id="07F334942A0ABB2DFE94E741E7812E7B" box="[310,1005,575,600]" pageId="104" pageNumber="2334" type="nomenclature">
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<paragraph id="4F56671F2A0ABB2DFE94E741E7812E7B" blockId="104.[310,1005,575,635]" box="[310,1005,575,600]" pageId="104" pageNumber="2334">
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<heading id="141ED0732A0ABB2DFE94E741E7812E7B" box="[310,1005,575,600]" pageId="104" pageNumber="2334">
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5.3.:
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<taxonomicName id="88E91C9C2A0ABB2DFECEE741E7222E7B" ID-CoL="6CS3L" authority="Nunes & Vaz-de-Mello, 2019" authorityName="Nunes & Vaz-de-Mello" authorityYear="2019" box="[364,846,575,600]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="2334" phylum="Arthropoda" rank="species" species="lucianomourai" status="sp. nov." subGenus="Cephagonus">
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<emphasis id="7D9DBB0D2A0ABB2DFECEE741E7222E7B" box="[364,846,575,600]" italics="true" pageId="104" pageNumber="2334">Dichotomius (Cephagonus) lucianomourai</emphasis>
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</taxonomicName>
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<emphasis id="7D9DBB0D2A0ABB2DFCF4E741E7812E7B" bold="true" box="[854,1005,575,600]" pageId="104" pageNumber="2334">
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<taxonomicNameLabel id="66AE06762A0ABB2DFCF4E741E7812E7B" box="[854,1005,575,600]" pageId="104" pageNumber="2334" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="07F334942A0ABB2DFD9AE71CE6872E58" box="[568,747,610,635]" pageId="104" pageNumber="2334" type="description">
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<paragraph id="4F56671F2A0ABB2DFD9AE71CE6872E58" blockId="104.[310,1005,575,635]" box="[568,747,610,635]" pageId="104" pageNumber="2334">
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(
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<figureCitation id="D7D27B9A2A0ABB2DFDE2E71CE6C32E58" box="[576,687,610,635]" captionStart="Figure 46" captionStartId="61.[160,225,1252,1275]" captionTargetBox="[163,1153,142,1220]" captionTargetId="figure@61.[163,1153,142,1220]" captionTargetPageId="61" captionText="Figure 46. D. lucianomourai: (a) male dorsal habitus; (b) female ventral habitus; (c) paramera dorsal view; (d) paramera ventral view; (e) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3666754" httpUri="https://zenodo.org/record/3666754/files/figure.png" pageId="104" pageNumber="2334">Figure 46</figureCitation>
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(a-e))
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</paragraph>
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</subSubSection>
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<subSubSection id="07F334942A0ABB2DFF02E7C7E6152F7F" pageId="104" pageNumber="2334" type="diagnosis">
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<paragraph id="4F56671F2A0ABB2DFF02E7C7E6152F7F" blockId="104.[160,1155,697,860]" pageId="104" pageNumber="2334">
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<emphasis id="7D9DBB0D2A0ABB2DFF02E7C7E5762EF1" bold="true" box="[160,282,697,722]" italics="true" pageId="104" pageNumber="2334">Diagnosis.</emphasis>
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this species is differentiated among the group by the following combined characters: males pronotal disc having at most two weak knobs on anterior portion (as
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<taxonomicName id="88E91C9C2A0ABB2DFF02E781E5342F34" authorityName="Nunes & Vaz-de-Mello" authorityYear="2019" box="[160,344,767,791]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="2334" phylum="Arthropoda" rank="species" species="fernandosilvai">
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<emphasis id="7D9DBB0D2A0ABB2DFF02E781E5342F34" box="[160,344,767,791]" italics="true" pageId="104" pageNumber="2334">D. fernandosilvai</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="88E91C9C2A0ABB2DFE31E781E67B2F34" box="[403,535,767,791]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="2334" phylum="Arthropoda" rank="species" species="clypeatus">
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<emphasis id="7D9DBB0D2A0ABB2DFE31E781E67B2F34" box="[403,535,767,791]" italics="true" pageId="104" pageNumber="2334">D. clypeatus</emphasis>
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</taxonomicName>
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); paramera very longı having more than half of phalobasis length and distance between triangular projections on female sixth ventrite equal to 1/3 of ventrite length (as on
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<taxonomicName id="88E91C9C2A0ABB2DFE46E63AE6072F7F" box="[484,619,836,860]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="104" pageNumber="2334" phylum="Arthropoda" rank="species" species="spadiceus">
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<emphasis id="7D9DBB0D2A0ABB2DFE46E63AE6072F7F" box="[484,619,836,860]" italics="true" pageId="104" pageNumber="2334">D. spadiceus</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="07F334942A0ABB2DFF1CE6F4E7752FC4" pageId="104" pageNumber="2334" type="materials_examined">
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<paragraph id="4F56671F2A0ABB2DFF1CE6F4E5EC2FE7" blockId="104.[190,1155,906,999]" pageId="104" pageNumber="2334">
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<emphasis id="7D9DBB0D2A0ABB2DFF1CE6F4E4A02FBD" bold="true" box="[190,204,906,926]" pageId="104" pageNumber="2334">●</emphasis>
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<materialsCitation id="FF816D422A0ABB2DFF7AE6F4E51B2FE7" ID-GBIF-Occurrence="2570409565" collectingDate="2003-10-31" collectionCode="CEMT" country="Brazil" location="Turvo. Pitfall" municipality="Derrubadas" pageId="104" pageNumber="2334" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul" typeStatus="holotype">
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<typeStatus id="9052D9BD2A0ABB2DFF7AE6F4E53E2F81" box="[216,338,906,930]" pageId="104" pageNumber="2334" type="holotype">HOLOTYPE</typeStatus>
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(
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<specimenCount id="59EFAC962A0ABB2DFEC1E6F4E5F22F81" box="[355,414,906,930]" pageId="104" pageNumber="2334" type="male">male</specimenCount>
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):
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<collectingCountry id="37FE278F2A0ABB2DFE11E6F4E66F2F81" box="[435,515,906,930]" name="Brazil" pageId="104" pageNumber="2334">BRASIL</collectingCountry>
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:
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<emphasis id="7D9DBB0D2A0ABB2DFDB3E6F4E69A2F81" bold="true" box="[529,758,906,930]" pageId="104" pageNumber="2334">
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||
<collectingRegion id="8D2DA9FD2A0ABB2DFDB3E6F4E69A2F81" box="[529,758,906,930]" country="Brazil" name="Rio Grande do Sul" pageId="104" pageNumber="2334">Rio Grande do Sul</collectingRegion>
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</emphasis>
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.
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<collectingMunicipality id="AF32FD652A0ABB2DFCA5E6F4E7E72F81" box="[775,907,906,930]" pageId="104" pageNumber="2334">Derrubadas</collectingMunicipality>
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. PE
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<location id="4A3631C42A0ABB2DFC62E6F4E03D2F81" LSID="urn:lsid:plazi:treatment:C740D6092A0ABB2CFE94E741E5722E65:4A3631C42A0ABB2DFC62E6F4E03D2F81" box="[960,1105,906,930]" country="Brazil" municipality="Derrubadas" name="Turvo. Pitfall" pageId="104" pageNumber="2334" stateProvince="Rio Grande do Sul">Turvo. Pitfall</location>
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.
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<date id="3B5741DF2A0ABB2DFBC2E6F4E5482FE7" pageId="104" pageNumber="2334" value="2003-10-31">
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<collectingDate id="2B13B8372A0ABB2DFBC2E6F4E5482FE7" pageId="104" pageNumber="2334" value="2003-10-31">31- x-2003</collectingDate>
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</date>
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[
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<collectionCode id="29F8FFDA2A0ABB2DFE92E6D2E51B2FE7" box="[304,375,940,964]" pageId="104" pageNumber="2334">CEMT</collectionCode>
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</materialsCitation>
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];
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</paragraph>
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<paragraph id="4F56671F2A0ABB2DFF1CE6B1E7752FC4" blockId="104.[190,1155,906,999]" box="[190,793,975,999]" pageId="104" pageNumber="2334">
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<emphasis id="7D9DBB0D2A0ABB2DFF1CE6B1E4A02FC0" bold="true" box="[190,204,975,995]" pageId="104" pageNumber="2334">●</emphasis>
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<typeStatus id="9052D9BD2A0ABB2DFF7AE6B1E5202FC4" box="[216,332,975,999]" pageId="104" pageNumber="2334" type="paratype">PARATYPE</typeStatus>
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(female): same data of
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<typeStatus id="9052D9BD2A0ABB2DFDF7E6B1E6D02FC4" box="[597,700,975,999]" pageId="104" pageNumber="2334" type="holotype">Holotype</typeStatus>
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[
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<collectionCode id="29F8FFDA2A0ABB2DFD6AE6B1E77C2FC4" box="[712,784,975,999]" pageId="104" pageNumber="2334">CEMT</collectionCode>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="07F334942A0ABB2CFF02E16DE5AE2D77" lastPageId="105" lastPageNumber="2335" pageId="104" pageNumber="2334" type="description">
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<paragraph id="4F56671F2A0ABB2DFF02E16DE7CC2A17" blockId="104.[160,1156,1043,1588]" pageId="104" pageNumber="2334">
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<emphasis id="7D9DBB0D2A0ABB2DFF02E16DE5DD280F" bold="true" box="[160,433,1043,1068]" italics="true" pageId="104" pageNumber="2334">
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Description (
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<typeStatus id="9052D9BD2A0ABB2DFE95E16DE5CE280F" box="[311,418,1043,1068]" pageId="104" pageNumber="2334" type="holotype">Holotype</typeStatus>
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).
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</emphasis>
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male BL: 13 mmı PW:
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<quantity id="8811CAFA2A0ABB2DFD16E16AE695280F" box="[692,761,1044,1068]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="104" pageNumber="2334" unit="mm" value="7.0">7 mm</quantity>
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. Colour: blackı shiny.
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<emphasis id="7D9DBB0D2A0ABB2DFC48E16AE044280F" bold="true" box="[1002,1064,1044,1068]" pageId="104" pageNumber="2334">Head</emphasis>
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: frontoclypeal surface smooth. Inter-ocular space-lacking puncturesı at most very fine ones (viewed under 30x magnification) near eyes. Gena borders are straight. Cephalic horn rounded at apex.
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<emphasis id="7D9DBB0D2A0ABB2DFECAE102E58C28B7" bold="true" box="[360,480,1148,1172]" pageId="104" pageNumber="2334">Pronotum</emphasis>
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: anterior portion having a subtle and narrow emargination. Pronotal disc smoothı shiny having a pair of central knobs on anterior portion. Posterior margin with ocellate punctures restricted to posterior angles. Posterior angle excavated and with groups of 5 ocellate punctures.
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<emphasis id="7D9DBB0D2A0ABB2DFD24E19AE74A28DF" bold="true" box="[646,806,1252,1276]" pageId="104" pageNumber="2334">Metasternum</emphasis>
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: sides and anterior portion of anterior lobe with ocellate setigerous puncturesı setae shortı with similar size to those on legs.
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<emphasis id="7D9DBB0D2A0ABB2DFF78E054E54C2961" bold="true" box="[218,288,1322,1346]" pageId="104" pageNumber="2334">Elytra</emphasis>
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: striae weakly impressed with weak punctures. Punctures separated by twice their diameter. Puncture on striae basis expanded forming a small fovea. Interstriae flatı lacking microsculpture.
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<emphasis id="7D9DBB0D2A0ABB2DFE0AE011E67129A4" bold="true" box="[424,541,1391,1415]" pageId="104" pageNumber="2334">Abdomen</emphasis>
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: punctures of each ventrite anterior margin interrupted medially. ventrites lacking setigerous punctures.
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<emphasis id="7D9DBB0D2A0ABB2DFD7AE0ECE7222989" bold="true" box="[728,846,1426,1450]" pageId="104" pageNumber="2334">Aedeagus</emphasis>
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(
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<figureCitation id="D7D27B9A2A0ABB2DFCC0E0ECE7BD2989" box="[866,977,1426,1450]" captionStart="Figure 46" captionStartId="61.[160,225,1252,1275]" captionTargetBox="[163,1153,142,1220]" captionTargetId="figure@61.[163,1153,142,1220]" captionTargetPageId="61" captionText="Figure 46. D. lucianomourai: (a) male dorsal habitus; (b) female ventral habitus; (c) paramera dorsal view; (d) paramera ventral view; (e) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3666754" httpUri="https://zenodo.org/record/3666754/files/figure.png" pageId="104" pageNumber="2334">Figure 46</figureCitation>
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(c-e)): paramera having twice the length of phalobasisı very long. Dorsallyı lacking basal and longitudinal excavation; apexes roundedı convergent. Laterallyı subtle declivity. Posterior margin with wide emargination near to phalobasis junction. Ventrallyı basis with a wide central excavation. Subgenital plate emarginated apicallyı rounded at basis.
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</paragraph>
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<paragraph id="4F56671F2A0ABB2CFF02E31FE5AE2D77" blockId="104.[160,1156,1633,1727]" lastBlockId="105.[160,1155,142,340]" lastPageId="105" lastPageNumber="2335" pageId="104" pageNumber="2334">
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<emphasis id="7D9DBB0D2A0ABB2DFF02E31FE5A92A59" bold="true" box="[160,453,1633,1658]" italics="true" pageId="104" pageNumber="2334">Morphological variation.</emphasis>
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male and female BL: 13 mmı PW:
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<quantity id="8811CAFA2A0ABB2DFCE9E31CE7E32A59" box="[843,911,1634,1658]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="104" pageNumber="2334" unit="mm" value="7.0">7 mm</quantity>
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. and pronotum width
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<quantity id="8811CAFA2A0ABB2DFF02E3FBE48B2ABE" box="[160,231,1669,1693]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="104" pageNumber="2334" unit="mm" value="7.0">7 mm</quantity>
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. Female is distinguished from the male as follows: Head: fronto-clypeal surface transversely wrinkledı interocular space smooth. Cephalic carina low with two acute tubercles on apex. Pronotum: simply convexı punctures weaker than on males but following same distribution pattern. Elytra: striae deeply impressedı punctures separated by twice their diameter. Basis with blue sheen. Abdomen: sixth ventrite bearing a pair of triangular projections reaching pygidium apical margin; distance between triangular projections equal to 1/10 of ventrite length. Disc with very fine puncture medially (viewed under 20x magnification).
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</paragraph>
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</subSubSection>
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<subSubSection id="07F334942A0BBB2CFF02E4FEE5DC2DFD" pageId="105" pageNumber="2335" type="etymology">
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<paragraph id="4F56671F2A0BBB2CFF02E4FEE5DC2DFD" blockId="105.[160,1155,384,478]" pageId="105" pageNumber="2335">
|
||
<emphasis id="7D9DBB0D2A0BBB2CFF02E4FEE54F2DBA" bold="true" box="[160,291,384,409]" italics="true" pageId="105" pageNumber="2335">Etymology.</emphasis>
|
||
named after our friend Dr. Luciano Moura (Universidade Federal do Rio Grande do Sulı
|
||
<collectingCountry id="37FE278F2A0BBB2CFEEDE4DAE5E32D9F" box="[335,399,420,444]" name="Brazil" pageId="105" pageNumber="2335">Brasil</collectingCountry>
|
||
)ı coleopterologist specialist on Chrysomelidaeı friend and scientific partner of both authors.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07F334942A0BBB2CFF02E775E5722E65" pageId="105" pageNumber="2335" type="distribution">
|
||
<paragraph id="4F56671F2A0BBB2CFF02E775E5722E65" blockId="105.[160,1155,523,582]" pageId="105" pageNumber="2335">
|
||
<emphasis id="7D9DBB0D2A0BBB2CFF02E775E55F2E07" bold="true" box="[160,307,523,548]" italics="true" pageId="105" pageNumber="2335">Distribution.</emphasis>
|
||
municipality of Derrubadası west of the state of Rio Grande do Sulı
|
||
<collectingCountry id="37FE278F2A0BBB2CFBE5E772E0EF2E07" box="[1095,1155,524,548]" name="Brazil" pageId="105" pageNumber="2335">Brazil</collectingCountry>
|
||
(
|
||
<figureCitation id="D7D27B9A2A0BBB2CFF05E750E57D2E65" box="[167,273,558,582]" captionStart="Figure 44" captionStartId="59.[160,225,819,842]" captionTargetBox="[164,1152,142,787]" captionTargetId="figure@59.[163,1153,142,787]" captionTargetPageId="59" captionText="Figure 44. Distribution map of D. spadiceusı D. lucianomourai and D. itatiaiaensis." figureDoi="http://doi.org/10.5281/zenodo.3666750" httpUri="https://zenodo.org/record/3666750/files/figure.png" pageId="105" pageNumber="2335">Figure 44</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |