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<document id="259A147C7AD0803B66D4385BFE030E8A" ID-DOI="10.5252/zoosystema2018v40a12" ID-GBIF-Dataset="6c3d39df-7484-4878-8952-9c6a0d926465" ID-ISSN="1638-9387" ID-Zenodo-Dep="3741548" ID-ZooBank="urn:lsid:zoobank.org:pub:6CBF9039-7E4D-4938-909A-2DB5113C8D35" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586173524968" checkinUser="felipe" docAuthor="Salazar-Vallejo, Sergio I." docDate="2018" docId="C17687BDFFF9FFD5FC30F977DB32F804" docLanguage="en" docName="zoosystema.40.12.227-325.pdf.imf" docOrigin="Zoosystema 40 (12)" docStyle="DocumentStyle:AFAF64E709DD7C8340B70A24A5804402.4:Zoosystema.2018-2019.journal_article" docStyleId="AFAF64E709DD7C8340B70A24A5804402" docStyleName="Zoosystema.2018-2019.journal_article" docStyleVersion="4" docTitle="Hesione hartmanae Salazar-Vallejo 2018, n. sp." docType="treatment" docVersion="5" lastPageNumber="261" masterDocId="3D4FFFC5FFDBFFF1FFD4FFA3DA42FFE9" masterDocTitle="Revision of Hesione Savigny in Lamarck, 1818 (Annelida, Errantia, Hesionidae)" masterLastPageNumber="325" masterPageNumber="227" pageNumber="259" updateTime="1698735891220" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6BC3EEC23065DE2B9210E68C14FBD43F">Revision of Hesione Savigny in Lamarck, 1818 (Annelida, Errantia, Hesionidae)</mods:title>
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<mods:namePart id="8752BF74E4C5B34A486891B59DDAC784">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:title id="A78AC08022F1CE4CCB668F3561F33838">Zoosystema</mods:title>
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<mods:date id="7292D14EF5FCE15954FC4A7D7FA89A5B">2018</mods:date>
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<mods:number id="C3787A0523CE79F0EB337C60C0A77EEA">2018-06-19</mods:number>
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<treatment id="C17687BDFFF9FFD5FC30F977DB32F804" ID-DOI="http://doi.org/10.5281/zenodo.3810258" ID-GBIF-Taxon="163437898" ID-Zenodo-Dep="3810258" LSID="urn:lsid:plazi:treatment:C17687BDFFF9FFD5FC30F977DB32F804" httpUri="http://treatment.plazi.org/id/C17687BDFFF9FFD5FC30F977DB32F804" lastPageId="36" lastPageNumber="261" pageId="34" pageNumber="259">
<subSubSection id="01C56520FFF9FFD3FC30F977DEBAF907" box="[996,1272,1747,1774]" pageId="34" pageNumber="259" type="nomenclature">
<paragraph id="496036ABFFF9FFD3FC30F977DEBAF907" blockId="34.[996,1272,1747,1806]" box="[996,1272,1747,1774]" pageId="34" pageNumber="259">
<heading id="122881C7FFF9FFD3FC30F977DEBAF907" box="[996,1272,1747,1774]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="259" reason="2">
<taxonomicName id="8EDF4D28FFF9FFD3FC30F977DEF4F907" ID-CoL="3L3Y9" authority="Salazar-Vallejo, 2018" authorityName="Salazar-Vallejo" authorityYear="2018" box="[996,1206,1747,1774]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="34" pageNumber="259" phylum="Annelida" rank="species" species="hartmanae" status="sp. nov.">
<emphasis id="7BABEAB9FFF9FFD3FC30F977DEF4F907" bold="true" box="[996,1206,1747,1774]" italics="true" pageId="34" pageNumber="259">Hesione hartmanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="609857C2FFF9FFD3FB69F977DEBAF907" box="[1213,1272,1748,1774]" pageId="34" pageNumber="259" rank="species">n. sp.</taxonomicNameLabel>
</heading>
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<subSubSection id="01C56520FFF9FFD3FBFDF957DFD1F8AA" pageId="34" pageNumber="259" type="description">
<paragraph id="496036ABFFF9FFD3FBFDF957DEF1F8E7" blockId="34.[996,1272,1747,1806]" box="[1065,1203,1780,1806]" pageId="34" pageNumber="259">
(
<figureCitation id="D1E42A2EFFF9FFD3FBE4F957DEC0F8E7" box="[1072,1154,1780,1806]" captionStart="FIG" captionStartId="34.[133,144,1229,1247]" captionTargetBox="[172,1417,215,1184]" captionTargetId="figure@34.[172,1417,214,1184]" captionTargetPageId="34" captionText="FIG. 18. — Hesione hartmanae n. sp., holotype, LACM 10158: A, anterior region, dorsal view; B, prostomium, dorsal view (arrows point to antennae); C, chaetiger 8, right parapodium, anterior view (inset: acicular lobe); D, same, neurochaetal blades (insets: tips of blades); E, posterior region, dorsal view. Scale bars: A, 2.2 mm; B, 0.3 mm; C, 0.6 mm; D, 65 µm; E, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3741586" httpUri="https://zenodo.org/record/3741586/files/figure.png" pageId="34" pageNumber="259">Figs 18</figureCitation>
,
<figureCitation id="D1E42A2EFFF9FFD3FB5AF957DEE8F8E7" box="[1166,1194,1780,1806]" captionStart="FIG" captionStartId="36.[132,143,1151,1169]" captionTargetBox="[172,1416,216,1105]" captionTargetId="figure@36.[172,1416,216,1105]" captionTargetPageId="36" captionText="FIG. 19. — Hesione hartmanae n. sp., paratypes, LACM 10159: A, smallest paratype, anterior region, dorsal view; B, same, prostomium and everted pharynx (arrow points to dorsal papilla); C, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe); D, largest paratype, anterior region, slightly oblique dorsal view; E, same,prostomium slightly oblique dorsal view; F, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe). Scale bars: A, 1.3 mm; B, 0.7 mm; C, 0.4 mm; D, 1.8 mm; E, 0.3 mm; F, 0.6 mm." figureDoi="http://doi.org/10.5281/zenodo.3741588" httpUri="https://zenodo.org/record/3741588/files/figure.png" pageId="34" pageNumber="259">19</figureCitation>
)
</paragraph>
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<uri id="3D4E3AA9FFF9FFD3FC9CF892DFD1F8AA" box="[840,1427,1841,1859]" pageId="34" pageNumber="259">
urn:lsid:zoobank.org:act:
<uuid id="3D790C7EFFF9FFD3FBC5F892DFD1F8AA" box="[1041,1427,1841,1859]" pageId="34" pageNumber="259">F50811AA-A413-4219-AAF1-0B3DD68E2E4F</uuid>
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<subSubSection id="01C56520FFF9FFD3FCF9F8DAD9EAF805" pageId="34" pageNumber="259" type="reference_group">
<paragraph id="496036ABFFF9FFD3FCF9F8DADFF2F878" blockId="34.[813,1456,1913,1937]" box="[813,1456,1913,1937]" pageId="34" pageNumber="259">
<treatmentCitationGroup id="69CF1185FFF9FFD3FCF9F8DADFF2F878" box="[813,1456,1913,1937]" pageId="34" pageNumber="259">
<emphasis id="7BABEAB9FFF9FFD3FCF9F8DAD99EF878" box="[813,988,1913,1937]" italics="true" pageId="34" pageNumber="259">
<taxonomicName id="8EDF4D28FFF9FFD3FCF9F8DAD98EF878" ID-CoL="3L3YT" authorityName="Savigny in Lamarck" authorityYear="1818" box="[813,972,1913,1937]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="34" pageNumber="259" phylum="Annelida" rank="species" species="splendida">Hesione splendida</taxonomicName>
</emphasis>
<treatmentCitation id="C87E10BAFFF9FFD3FC35F8DADE3EF878" author="MONRO C. C. A." box="[993,1148,1913,1937]" page="26" pageId="34" pageNumber="259" year="1933">
<bibRefCitation id="2D4E4B5AFFF9FFD3FC35F8DADE3EF878" author="Monro" box="[993,1148,1913,1937]" firstAuthor="Monro" pageId="34" pageNumber="259" pagination="1 - 96" refId="ref96564" refString="MONRO C. C. A. 1933. - The Polychaeta Errantia collected by Dr C. Crossland at Colon, in the Panama Region, and the Galapagos Islands during the Expedition of the S. Y. &quot; St. George &quot;. Proceedings of the Zoological Society, London 1933 (1): 1 - 96. http: // onlinelibrary. wiley. com / doi / 10.1111 / j. 1096 - 3642.1933. tb 01578. x / full" type="journal article" year="1933">Monro 1933: 26</bibRefCitation>
</treatmentCitation>
(
<emphasis id="7BABEAB9FFF9FFD3FB5CF8D9DEE8F878" box="[1160,1194,1914,1937]" italics="true" pageId="34" pageNumber="259">non</emphasis>
Savigny
<emphasis id="7BABEAB9FFF9FFD3FB28F8DADF4DF879" box="[1276,1295,1913,1936]" italics="true" pageId="34" pageNumber="259">in</emphasis>
<bibRefCitation id="2D4E4B5AFFF9FFD3FAC7F8DADFE6F878" author="Lamarck" box="[1299,1444,1913,1937]" firstAuthor="Lamarck" pageId="34" pageNumber="259" refId="ref95809" refString="LAMARCK J. B. 1818. - Histoire Naturelle des Animaux sans Vertebres, presentant les Caracteres Generaux et Particuliers de ces Animaux, leur Distribution, leurs Classes, leurs Familles, leurs Genres, et la Citation des principales Especes qui s'y rapportent; Precedes d'une Introduction offrant la Determination des Caracteres Essentiels de l'Animal, sa distinction du Vegetal et des autres Corps Naturels, en fin, l'Exposition des Principes fondamentaux de la Zoologie, Deterville &amp; Verdiere, Paris, volume 5, 612 p. http: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 64280058" type="book" year="1818">Lamarck, 1818</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="496036ABFFF9FFD3FCF9F819D9EAF805" blockId="34.[813,1456,1977,2028]" pageId="34" pageNumber="259">
<treatmentCitationGroup id="69CF1185FFF9FFD3FCF9F819D9EAF805" pageId="34" pageNumber="259">
<emphasis id="7BABEAB9FFF9FFD3FCF9F819D9AEF838" box="[813,1004,1978,2001]" italics="true" pageId="34" pageNumber="259">
<taxonomicName id="8EDF4D28FFF9FFD3FCF9F819D995F838" ID-CoL="3L3YD" authorityName="Grube" authorityYear="1878" box="[813,983,1978,2001]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="34" pageNumber="259" phylum="Annelida" rank="species" species="intertexta">Hesione intertexta</taxonomicName>
</emphasis>
<treatmentCitation id="C87E10BAFFF9FFD3FC20F819DEFDF838" author="HARTMAN O." box="[1012,1215,1977,2001]" page="212" pageId="34" pageNumber="259" year="1940">
<bibRefCitation id="2D4E4B5AFFF9FFD3FC20F819DEFDF838" author="Hartman" box="[1012,1215,1977,2001]" firstAuthor="Hartman" pageId="34" pageNumber="259" pagination="173 - 287" refId="ref94489" refString="HARTMAN O. 1940. - Polychaetous annelids, 2. Chrysopetalidae to Goniadidae. Allan Hancock Pacific Expeditions 7: 173 - 287, pls 31 - 34. http: // cdm 15799. contentdm. oclc. org / cdm / ref / collection / p 15799 coll 82 / id / 5767" type="journal article" year="1940">Hartman 1940: 212</bibRefCitation>
</treatmentCitation>
, pl. 33, figs 30-31 (
<emphasis id="7BABEAB9FFF9FFD3FA5CF819D90CF802" italics="true" pageId="34" pageNumber="259">partim</emphasis>
, fig. 30).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="01C56520FFF8FFD2FF50FF7BD8ECFC04" pageId="35" pageNumber="260" type="materials_examined">
<paragraph id="496036ABFFF8FFD2FF50FF7BDBCFFE36" blockId="35.[132,776,216,479]" pageId="35" pageNumber="260">
TYPE MATERIAL. —
<materialsCitation id="F9B73CF6FFF8FFD2FE91FF7BDBCAFE36" ID-GBIF-Occurrence="2595817418" collectingDate="1934-01-19" collectingDateMax="1934-12-03" collectingDateMin="1934-01-19" collectionCode="LACM" collectorName="RV Velero &amp; Sta." county="Eastern Pacific." elevation="27" latitude="-1.2436111" location="Post Office Bay" longLatPrecision="21" longitude="-90.46889" municipality="Bindloe Island" pageId="35" pageNumber="260" specimenCode="LACM 10158, LACM 10159" specimenCount="3" stateProvince="Galapagos Islands" typeStatus="holotype">
<emphasis id="7BABEAB9FFF8FFD2FE91FF7BD8DFFF19" bold="true" box="[325,669,216,240]" pageId="35" pageNumber="260">
<collectingCounty id="A0014E27FFF8FFD2FE91FF7BDB9CFF19" box="[325,478,216,240]" pageId="35" pageNumber="260">Eastern Pacific.</collectingCounty>
<collectingRegion id="8B1BF849FFF8FFD2FE30FF7BD8DBFF19" box="[484,665,216,240]" country="Ecuador" name="Galapagos" pageId="35" pageNumber="260">Galápagos Islands</collectingRegion>
.
</emphasis>
<typeStatus id="96648809FFF8FFD2FD70FF7AD941FF18" box="[676,771,217,241]" pageId="35" pageNumber="260" type="holotype">Holotype</typeStatus>
,
<specimenCode id="19799ED0FFF8FFD2FF50FF50DB53FEE2" box="[132,273,243,267]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="35" pageNumber="260" type="Museum">LACM 10158</specimenCode>
,
<collectorName id="E42A537DFFF8FFD2FECDFF57DB34FEE2" box="[281,374,243,267]" pageId="35" pageNumber="260">
RV
<emphasis id="7BABEAB9FFF8FFD2FEEAFF50DB34FEE2" box="[318,374,243,267]" italics="true" pageId="35" pageNumber="260">Velero</emphasis>
</collectorName>
III,
<collectorName id="E42A537DFFF8FFD2FE4BFF50DB86FEE2" box="[415,452,243,267]" pageId="35" pageNumber="260">Sta.</collectorName>
310 (
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,
<geoCoordinate id="2CEB506CFFF8FFD2FDA9FF50D8BEFEE2" box="[637,764,243,267]" degrees="90" direction="west" minutes="31" orientation="longitude" pageId="35" pageNumber="260" precision="15" seconds="00" value="-90.51667">90°3100”W</geoCoordinate>
), off
<collectingMunicipality id="A904ACD1FFF8FFD2FF77FEADDB6CFECF" box="[163,302,270,294]" pageId="35" pageNumber="260">Bindloe Island</collectingMunicipality>
,
<elevation id="C2F2D198FFF8FFD2FEE3FEADDBE5FECF" box="[311,423,270,294]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="35" pageNumber="260" unit="m" value="27.0">
<quantity id="8E279B4EFFF8FFD2FEE3FEADDB2BFECC" box="[311,361,270,294]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="35" pageNumber="260" unit="m" value="27.0">27 m</quantity>
depth
</elevation>
, rock, tangles,
<date id="3D61106BFFF8FFD2FDE5FEADD8DDFECF" box="[561,671,270,294]" pageId="35" pageNumber="260" value="1934-12-03">
<collectingDate id="2D25E983FFF8FFD2FDE5FEADD8DDFECF" box="[561,671,270,294]" pageId="35" pageNumber="260" value="1934-12-03">3.XII.1934</collectingDate>
</date>
.
<typeStatus id="96648809FFF8FFD2FD73FEADD946FECF" box="[679,772,270,294]" pageId="35" pageNumber="260" type="paratype">Paratypes</typeStatus>
,
<specimenCode id="19799ED0FFF8FFD2FF50FE8BDB50FEA9" box="[132,274,296,320]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="35" pageNumber="260" type="Museum">LACM 10159</specimenCode>
,
<collectorName id="E42A537DFFF8FFD2FECFFE8ADB3AFEA9" box="[283,376,296,320]" pageId="35" pageNumber="260">
RV
<emphasis id="7BABEAB9FFF8FFD2FE94FE8BDB3AFEA9" box="[320,376,296,320]" italics="true" pageId="35" pageNumber="260">Velero</emphasis>
</collectorName>
III,
<collectorName id="E42A537DFFF8FFD2FE77FE8BDB8AFEA9" box="[419,456,296,320]" pageId="35" pageNumber="260">Sta.</collectorName>
167 (
<geoCoordinate id="2CEB506CFFF8FFD2FDD4FE8BD830FEA9" box="[512,626,296,320]" degrees="01" direction="south" minutes="14" orientation="latitude" pageId="35" pageNumber="260" precision="15" seconds="37" value="-1.2436111">01°1437”S</geoCoordinate>
,
<geoCoordinate id="2CEB506CFFF8FFD2FDA8FE8BD8B9FEA9" box="[636,763,296,320]" degrees="90" direction="west" minutes="28" orientation="longitude" pageId="35" pageNumber="260" precision="15" seconds="08" value="-90.46889">90°2808”W</geoCoordinate>
),
<location id="4C006070FFF8FFD2FF50FEE0DB5AFEB2" LSID="urn:lsid:plazi:treatment:C17687BDFFF9FFD5FC30F977DB32F804:4C006070FFF8FFD2FF50FEE0DB5AFEB2" box="[132,280,323,347]" county="Eastern Pacific." latitude="-1.2436111" longLatPrecision="21" longitude="-90.46889" municipality="Bindloe Island" name="Post Office Bay" pageId="35" pageNumber="260" stateProvince="Galapagos Islands">Post Office Bay</location>
,
<location id="4C006070FFF8FFD2FEF5FEE0DBECFEB2" LSID="urn:lsid:plazi:treatment:C17687BDFFF9FFD5FC30F977DB32F804:4C006070FFF8FFD2FEF5FEE0DBECFEB2" box="[289,430,323,347]" county="Eastern Pacific." latitude="-1.2436111" longLatPrecision="21" longitude="-90.46889" municipality="Bindloe Island" name="Charles Island" pageId="35" pageNumber="260" stateProvince="Galapagos Islands">Charles Island</location>
,
<elevation id="C2F2D198FFF8FFD2FE62FEE0D86AFEB2" box="[438,552,323,347]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="35" pageNumber="260" unit="m" value="27.0">
<quantity id="8E279B4EFFF8FFD2FE62FEE0DBAAFEB3" box="[438,488,323,347]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="35" pageNumber="260" unit="m" value="27.0">27 m</quantity>
depth
</elevation>
, rocks,
<date id="3D61106BFFF8FFD2FDBBFEE0D893FEB2" box="[623,721,323,347]" pageId="35" pageNumber="260" value="1934-01-19">
<collectingDate id="2D25E983FFF8FFD2FDBBFEE0D893FEB2" box="[623,721,323,347]" pageId="35" pageNumber="260" value="1934-01-19">19.I.1934</collectingDate>
</date>
[four complete
<typeStatus id="96648809FFF8FFD2FF30FEFDDB02FE9F" box="[228,320,350,374]" pageId="35" pageNumber="260" type="paratype">paratypes</typeStatus>
<quantity id="8E279B4EFFF8FFD2FE92FEFEDBF2FE9C" box="[326,432,349,373]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="4.0" metricValueMin="2.1" pageId="35" pageNumber="260" unit="mm" value="30.5" valueMax="40.0" valueMin="21.0">21-40 mm</quantity>
long,
<quantity id="8E279B4EFFF8FFD2FE3AFEFED87CFE9C" box="[494,574,349,373]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="35" pageNumber="260" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4-5 mm</quantity>
wide; some more or less macerated, most cirri and neurochaetal blades on site; chaetal lobes variably invaginated; right parapodium of chaetiger
<quantity id="8E279B4EFFF8FFD2FD09FE31D944FE43" box="[733,774,402,426]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="35" pageNumber="260" unit="in" value="9.0">9 in</quantity>
<specimenCount id="5FD9FD22FFF8FFD2FF50FE0EDAB9FE2D" box="[132,251,428,453]" pageId="35" pageNumber="260" type="generic">3 specimens</specimenCount>
removed (kept in vial); even the smallest one has oocytes, each about
<quantity id="8E279B4EFFF8FFD2FEE4FE64DBC1FE36" box="[304,387,455,479]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="35" pageNumber="260" unit="mm" value="100.0">100 µm</quantity>
]
</materialsCitation>
.
</paragraph>
<paragraph id="496036ABFFF8FFD2FF50FE5BD8ECFC04" blockId="35.[132,775,504,1005]" pageId="35" pageNumber="260">
ADDITIONAL MATERIAL. —
<materialsCitation id="F9B73CF6FFF8FFD2FE49FE5BDB66FCDA" ID-GBIF-Occurrence="2595817486" collectingDate="1923" collectingDateMax="1924" collectingDateMin="1923" collectionCode="BMNH" collectorName="RV St-George Pacific Expedition &amp; Sta. &amp; C. Crossland" location="Eastern Pacific." pageId="35" pageNumber="260" specimenCode="BMNH 1932.12" specimenCount="1" stateProvince="Galapagos Islands">
<emphasis id="7BABEAB9FFF8FFD2FE49FE5BD945FDF9" bold="true" box="[413,775,504,528]" pageId="35" pageNumber="260">
<location id="4C006070FFF8FFD2FE49FE5BD87FFDF9" LSID="urn:lsid:plazi:treatment:C17687BDFFF9FFD5FC30F977DB32F804:4C006070FFF8FFD2FE49FE5BD87FFDF9" box="[413,573,504,528]" name="Eastern Pacific." pageId="35" pageNumber="260" stateProvince="Galapagos Islands">Eastern Pacific.</location>
<collectingRegion id="8B1BF849FFF8FFD2FD91FE5BD941FDF9" box="[581,771,504,528]" country="Ecuador" name="Galapagos" pageId="35" pageNumber="260">Galápagos Islands</collectingRegion>
.
</emphasis>
<specimenCount id="5FD9FD22FFF8FFD2FF50FDB0DAB4FDC2" box="[132,246,531,555]" pageId="35" pageNumber="260" type="generic">1 specimen</specimenCount>
,
<specimenCode id="19799ED0FFF8FFD2FED4FDB0DBE8FDC2" box="[256,426,531,555]" collectionCode="BMNH" pageId="35" pageNumber="260">BMNH 1932.12</specimenCode>
.24.129,
<collectorName id="E42A537DFFF8FFD2FDD5FDB0DA80FDAC" pageId="35" pageNumber="260">
RV
<emphasis id="7BABEAB9FFF8FFD2FDFCFDB0D8C0FDC3" box="[552,642,531,554]" italics="true" pageId="35" pageNumber="260">St-George</emphasis>
Pacific Expedition
</collectorName>
,
<date id="3D61106BFFF8FFD2FF1AFD8EDB7CFDAC" box="[206,318,557,581]" pageId="35" pageNumber="260" value="1923" valueMax="1924">
<collectingDate id="2D25E983FFF8FFD2FF1AFD8EDB7CFDAC" box="[206,318,557,581]" pageId="35" pageNumber="260" value="1923" valueMax="1924">1923-1924</collectingDate>
</date>
,
<collectingRegion id="8B1BF849FFF8FFD2FE9DFD8EDBEFFDAC" box="[329,429,557,581]" country="Ecuador" name="Galapagos" pageId="35" pageNumber="260">Galápagos</collectingRegion>
<collectorName id="E42A537DFFF8FFD2FE61FD8EDB98FDAC" box="[437,474,557,581]" pageId="35" pageNumber="260">Sta.</collectorName>
9,
<collectorName id="E42A537DFFF8FFD2FE28FD8ED83FFDAC" box="[508,637,557,581]" pageId="35" pageNumber="260">C. Crossland</collectorName>
coll. [
<quantity id="8E279B4EFFF8FFD2FD68FD8ED945FDAD" box="[700,775,557,581]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="35" pageNumber="260" unit="mm" value="35.0">35 mm</quantity>
long,
<quantity id="8E279B4EFFF8FFD2FF68FDE4DAB4FDB6" box="[188,246,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="35" pageNumber="260" unit="mm" value="5.0">5 mm</quantity>
wide; almost colorless, a few poorly defined transverse bands along some anterior chaetigers; antennae twice as long as wide; anterior eyes slightly larger than posterior ones; right parapodium of chaetiger 12 previously removed; right parapodium of chaetiger 10 removed for observation (kept in vial); acicular lobe double, tines digitate, blunt, upper tine slightly longer than lower one; neurochaetal blades broken or without blades; blades bidentate, subdistal tooth smaller, guards most broken, if complete surpassing subdistal tooth]
</materialsCitation>
. —
<materialsCitation id="F9B73CF6FFF8FFD2FE85FCB8D8ECFC04" ID-GBIF-Occurrence="2595817463" collectingDate="1934-01-20" collectionCode="LACM" collectorName="RV Velero &amp; Sta." elevation="37" latitude="-0.77166665" location="Indefatigable Island" longLatPrecision="21" longitude="-90.324165" municipality="Academy Bay" pageId="35" pageNumber="260" specimenCode="LACM 10160" specimenCount="2" stateProvince="Galapagos">
<specimenCount id="5FD9FD22FFF8FFD2FE85FCB8DB8FFCDA" box="[337,461,795,819]" pageId="35" pageNumber="260" type="generic">2 specimens</specimenCount>
,
<specimenCode id="19799ED0FFF8FFD2FE0CFCB8D828FCDA" box="[472,618,795,819]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="35" pageNumber="260" type="Museum">LACM 10160</specimenCode>
),
<collectorName id="E42A537DFFF8FFD2FDAAFCBFD89DFCDA" box="[638,735,795,819]" pageId="35" pageNumber="260">
RV
<emphasis id="7BABEAB9FFF8FFD2FD72FCB8D89DFCDA" box="[678,735,795,819]" italics="true" pageId="35" pageNumber="260">Velero</emphasis>
</collectorName>
III,
<collectorName id="E42A537DFFF8FFD2FF50FC95DAEBFCA7" box="[132,169,822,846]" pageId="35" pageNumber="260">Sta.</collectorName>
169 (
<geoCoordinate id="2CEB506CFFF8FFD2FF31FC96DB1AFCA4" box="[229,344,821,845]" degrees="00" direction="south" minutes="46" orientation="latitude" pageId="35" pageNumber="260" precision="15" seconds="18" value="-0.77166665">00°4618”S</geoCoordinate>
,
<geoCoordinate id="2CEB506CFFF8FFD2FEB0FC95DBA6FCA7" box="[356,484,822,846]" degrees="90" direction="west" minutes="19" orientation="longitude" pageId="35" pageNumber="260" precision="15" seconds="27" value="-90.324165">90°1927”W</geoCoordinate>
),
<collectingMunicipality id="A904ACD1FFF8FFD2FE22FC95D83FFCA7" box="[502,637,822,846]" pageId="35" pageNumber="260">Academy Bay</collectingMunicipality>
,
<location id="4C006070FFF8FFD2FD5CFC95DA80FC81" LSID="urn:lsid:plazi:treatment:C17687BDFFF9FFD5FC30F977DB32F804:4C006070FFF8FFD2FD5CFC95DA80FC81" latitude="-0.77166665" longLatPrecision="21" longitude="-90.324165" municipality="Academy Bay" name="Indefatigable Island" pageId="35" pageNumber="260" stateProvince="Galapagos">Indefatigable Island</location>
,
<quantity id="8E279B4EFFF8FFD2FF1AFCF3DB64FC81" box="[206,294,848,872]" metricMagnitude="1" metricUnit="m" metricValue="3.65" metricValueMax="4.5" metricValueMin="2.8" pageId="35" pageNumber="260" unit="m" value="36.5" valueMax="45.0" valueMin="28.0">
<elevation id="C2F2D198FFF8FFD2FF1AFCF3DB64FC81" box="[206,294,848,872]" metricMagnitude="1" metricUnit="m" metricValue="3.65" metricValueMax="4.5" metricValueMin="2.8" pageId="35" pageNumber="260" unit="m" value="36.5" valueMax="45.0" valueMin="28.0">28-45 m</elevation>
</quantity>
depth, sand, rock, algae,
<date id="3D61106BFFF8FFD2FDFDFCF3D8CEFC81" box="[553,652,848,872]" pageId="35" pageNumber="260" value="1934-01-20">
<collectingDate id="2D25E983FFF8FFD2FDFDFCF3D8CEFC81" box="[553,652,848,872]" pageId="35" pageNumber="260" value="1934-01-20">20.I.1934</collectingDate>
</date>
[
<quantity id="8E279B4EFFF8FFD2FD4EFCF3D945FC81" box="[666,775,848,872]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="35" pageNumber="260" unit="mm" value="29.5" valueMax="31.0" valueMin="28.0">28-31 mm</quantity>
long,
<quantity id="8E279B4EFFF8FFD2FF68FCC8DB4EFC6B" box="[188,268,875,899]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="35" pageNumber="260" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3-4 mm</quantity>
wide; one macerated, the other in better condition; most cirri and neurochaetal blades on site; chaetal lobes variably invaginated; right parapodium of chaetiger 9 from both specimens removed (kept in vial); upper acicular lobe twice as long as lower one; largest specimen with oocytes, each about
<quantity id="8E279B4EFFF8FFD2FD85FC77D8E6FC04" box="[593,676,980,1005]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="35" pageNumber="260" unit="mm" value="100.0">100 µm</quantity>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="01C56520FFF8FFD2FF50FBAEDBC4FB2D" pageId="35" pageNumber="260" type="etymology">
<paragraph id="496036ABFFF8FFD2FF50FBAEDBC4FB2D" blockId="35.[132,776,1037,1220]" pageId="35" pageNumber="260">
ETYMOLOGY. — This species is named after Dr Olga Hartman, eminent taxonomist of polychaetes from the Allan Hancock Foundation, University of Southern
<collectingRegion id="8B1BF849FFF8FFD2FE6DFBE1D85DFBB3" box="[441,543,1090,1114]" country="United States of America" name="California" pageId="35" pageNumber="260">California</collectingRegion>
, to honor her immense legacy including lots of publications and new taxa, and because she provided some relevant details to recognise these specimens as a different, undescribed species. The name is a noun in genitive (
<bibRefCitation id="2D4E4B5AFFF8FFD2FF5FFB0FDB45FB2D" author="ICZN" box="[139,263,1196,1220]" firstAuthor="ICZN" pageId="35" pageNumber="260" refId="ref95314" refString="ICZN (INTERNATIONAL COMMISSION ON ZOOLOGICAL NOMENCLA- TURE). 1999. - International Code of Zoological Nomenclature, Fourth Edition. International Trust for Zoological Nomenclature in association with the British Museum (Natural History), London, 306 p. http: // www. iczn. org / iczn / index. jsp" type="book" year="1999">ICZN 1999</bibRefCitation>
: art. 31.1.2).
</paragraph>
</subSubSection>
<subSubSection id="01C56520FFF8FFD2FF50FB47DBCEFAFE" pageId="35" pageNumber="260" type="distribution">
<paragraph id="496036ABFFF8FFD2FF50FB47DBCEFAFE" blockId="35.[132,775,1252,1303]" pageId="35" pageNumber="260">
DISTRIBUTION. —
<collectingRegion id="8B1BF849FFF8FFD2FEEFFB47DBA9FB15" box="[315,491,1252,1276]" country="Ecuador" name="Galapagos" pageId="35" pageNumber="260">Galápagos Islands</collectingRegion>
,
<collectingCountry id="31C8763BFFF8FFD2FE20FB47D805FB15" box="[500,583,1252,1276]" name="Ecuador" pageId="35" pageNumber="260">Ecuador</collectingCountry>
, in subtidal to
<elevation id="C2F2D198FFF8FFD2FD00FB47DAFDFAFE" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="35" pageNumber="260" unit="m" value="45.0">
<quantity id="8E279B4EFFF8FFD2FD00FB47D945FB15" box="[724,775,1252,1276]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="35" pageNumber="260" unit="m" value="45.0">45 m</quantity>
depth
</elevation>
, in mixed substrates.
</paragraph>
</subSubSection>
<subSubSection id="01C56520FFF8FFD2FF50FA94D8BFFA50" pageId="35" pageNumber="260" type="diagnosis">
<paragraph id="496036ABFFF8FFD2FF50FA94D8BFFA50" blockId="35.[132,775,1335,1465]" pageId="35" pageNumber="260">
DIAGNOSIS. —
<taxonomicName id="8EDF4D28FFF8FFD2FEF4FA9BDB28FAA6" authorityName="Savigny in Lamarck" authorityYear="1818" box="[288,362,1336,1359]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="35" pageNumber="260" phylum="Annelida" rank="genus">
<emphasis id="7BABEAB9FFF8FFD2FEF4FA9BDB28FAA6" box="[288,362,1336,1359]" italics="true" pageId="35" pageNumber="260">Hesione</emphasis>
</taxonomicName>
with prostomium rectangular; parapodia with dorsal cirri basally cylindrical, dorsal cirrophore twice as long as wide; larger acicula blackish; acicular lobe double, tines long, digitate, subequal; neurochaetal blades bidentate, 3-6 times as long as wide; subdistal tooth smaller; guards approaching distal tooth.
</paragraph>
</subSubSection>
<subSubSection id="01C56520FFF8FFD2FF50FA57DE46FA86" pageId="35" pageNumber="260" type="description">
<paragraph id="496036ABFFF8FFD2FF50FA57DB57F9E4" blockId="35.[132,775,1524,2030]" box="[132,277,1524,1550]" pageId="35" pageNumber="260">DESCRIPTION</paragraph>
<paragraph id="496036ABFFF8FFD2FF50F9B7DB07F907" blockId="35.[132,775,1524,2030]" pageId="35" pageNumber="260">
<typeStatus id="96648809FFF8FFD2FF50F9B7DAAFF9C7" box="[132,237,1556,1582]" pageId="35" pageNumber="260" type="holotype">Holotype</typeStatus>
, LACM 10158, mature, complete, subcylindrical, tapered posteriorly, posterior region bent laterally, first four chaetigers with ill-defined brownish transverse bands, progressively paler (
<figureCitation id="D1E42A2EFFF8FFD2FEE7F9D7DBD1F967" box="[307,403,1652,1678]" captionStart="FIG" captionStartId="34.[133,144,1229,1247]" captionTargetBox="[172,1417,215,1184]" captionTargetId="figure@34.[172,1417,214,1184]" captionTargetPageId="34" captionText="FIG. 18. — Hesione hartmanae n. sp., holotype, LACM 10158: A, anterior region, dorsal view; B, prostomium, dorsal view (arrows point to antennae); C, chaetiger 8, right parapodium, anterior view (inset: acicular lobe); D, same, neurochaetal blades (insets: tips of blades); E, posterior region, dorsal view. Scale bars: A, 2.2 mm; B, 0.3 mm; C, 0.6 mm; D, 65 µm; E, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3741586" httpUri="https://zenodo.org/record/3741586/files/figure.png" pageId="35" pageNumber="260">Fig. 18A</figureCitation>
) in ethanol; some cirri lost, others in regeneration, most neurochaetal blades on site; right parapodium of chaetiger 8 removed (kept in vial). Body
<quantity id="8E279B4EFFF8FFD2FD6CF917D945F926" box="[696,775,1716,1743]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="35" pageNumber="260" unit="mm" value="41.0">41 mm</quantity>
long,
<quantity id="8E279B4EFFF8FFD2FF16F977DB41F907" box="[194,259,1748,1775]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="35" pageNumber="260" unit="mm" value="5.0">5 mm</quantity>
wide.
</paragraph>
<paragraph id="496036ABFFF8FFD2FF4FF957D8ECF807" blockId="35.[132,775,1524,2030]" pageId="35" pageNumber="260">
Prostomium slightly as long as wide, anterior margin truncate with a deep depression, lateral margins straight, slightly divergent posteriorly, posterior margin deeply cleft about ¼ prostomial length (reaching level of posterior eyes), longitudinal depression indistinct (
<figureCitation id="D1E42A2EFFF8FFD2FE1EF8D0D86FF864" box="[458,557,1907,1933]" captionStart="FIG" captionStartId="34.[133,144,1229,1247]" captionTargetBox="[172,1417,215,1184]" captionTargetId="figure@34.[172,1417,214,1184]" captionTargetPageId="34" captionText="FIG. 18. — Hesione hartmanae n. sp., holotype, LACM 10158: A, anterior region, dorsal view; B, prostomium, dorsal view (arrows point to antennae); C, chaetiger 8, right parapodium, anterior view (inset: acicular lobe); D, same, neurochaetal blades (insets: tips of blades); E, posterior region, dorsal view. Scale bars: A, 2.2 mm; B, 0.3 mm; C, 0.6 mm; D, 65 µm; E, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3741586" httpUri="https://zenodo.org/record/3741586/files/figure.png" pageId="35" pageNumber="260">Fig. 18B</figureCitation>
). Antennae minute, digitate, twice as long as wide, about as long as free interocular distance. Eyes brownish, circular; anterior eyes slightly larger and more distant to each other than posterior ones.
</paragraph>
<paragraph id="496036ABFFF8FFD2FC97FF7ADFDFFEDD" blockId="35.[813,1456,217,1040]" pageId="35" pageNumber="260">Tentacular cirri undulated, longest reaching chaetiger 4. Lateral cushions projected, with 2 sections in anterior to median chaetigers, and with 3 ones in posterior segments.</paragraph>
<paragraph id="496036ABFFF8FFD2FC97FE9AD9A0FE1B" blockId="35.[813,1456,217,1040]" pageId="35" pageNumber="260">
Parapodia with chaetal lobes tapered, truncate, dorsal cirri with cirrophores 3 times as long as wide (
<figureCitation id="D1E42A2EFFF8FFD2FAEEFEFADFE0FE9A" box="[1338,1442,345,371]" captionStart="FIG" captionStartId="34.[133,144,1229,1247]" captionTargetBox="[172,1417,215,1184]" captionTargetId="figure@34.[172,1417,214,1184]" captionTargetPageId="34" captionText="FIG. 18. — Hesione hartmanae n. sp., holotype, LACM 10158: A, anterior region, dorsal view; B, prostomium, dorsal view (arrows point to antennae); C, chaetiger 8, right parapodium, anterior view (inset: acicular lobe); D, same, neurochaetal blades (insets: tips of blades); E, posterior region, dorsal view. Scale bars: A, 2.2 mm; B, 0.3 mm; C, 0.6 mm; D, 65 µm; E, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3741586" httpUri="https://zenodo.org/record/3741586/files/figure.png" pageId="35" pageNumber="260">Fig. 18C</figureCitation>
); cirrostyle cylindrical, smooth basally, then annulated, become articulated in distal half, as long as half body width (without parapodia). Ventral cirri smooth, tapered, surpassing chaetal lobe.
</paragraph>
<paragraph id="496036ABFFF8FFD2FC97FE5BD9B3FD7B" blockId="35.[813,1456,217,1040]" pageId="35" pageNumber="260">
Neuraciculae blackish, 1-2 markedly thicker and 1 very thin, very close to each other, making their detection difficult. Acicular lobe double, tapered, blunt, upper lobe twice as long as lower one, half as long as chaetal fascicle width (
<figureCitation id="D1E42A2EFFF8FFD2FCE0FDDBD9D7FD7B" box="[820,917,632,658]" captionStart="FIG" captionStartId="34.[133,144,1229,1247]" captionTargetBox="[172,1417,215,1184]" captionTargetId="figure@34.[172,1417,214,1184]" captionTargetPageId="34" captionText="FIG. 18. — Hesione hartmanae n. sp., holotype, LACM 10158: A, anterior region, dorsal view; B, prostomium, dorsal view (arrows point to antennae); C, chaetiger 8, right parapodium, anterior view (inset: acicular lobe); D, same, neurochaetal blades (insets: tips of blades); E, posterior region, dorsal view. Scale bars: A, 2.2 mm; B, 0.3 mm; C, 0.6 mm; D, 65 µm; E, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3741586" httpUri="https://zenodo.org/record/3741586/files/figure.png" pageId="35" pageNumber="260">Fig. 18C</figureCitation>
[inset]).
</paragraph>
<paragraph id="496036ABFFF8FFD2FC97FD3BD9BEFCDB" blockId="35.[813,1456,217,1040]" pageId="35" pageNumber="260">
Neurochaetae about 30 per bundle, blades bidentate (sometimes guard and teeth eroded), subdistal tooth smaller, blades at a certain angle from the handle, 3-6 times as long as wide, decreasing in size ventrally, guard approaching distal tooth (
<figureCitation id="D1E42A2EFFF8FFD2FCE0FCB4D9D5FCD8" box="[820,919,791,817]" captionStart="FIG" captionStartId="34.[133,144,1229,1247]" captionTargetBox="[172,1417,215,1184]" captionTargetId="figure@34.[172,1417,214,1184]" captionTargetPageId="34" captionText="FIG. 18. — Hesione hartmanae n. sp., holotype, LACM 10158: A, anterior region, dorsal view; B, prostomium, dorsal view (arrows point to antennae); C, chaetiger 8, right parapodium, anterior view (inset: acicular lobe); D, same, neurochaetal blades (insets: tips of blades); E, posterior region, dorsal view. Scale bars: A, 2.2 mm; B, 0.3 mm; C, 0.6 mm; D, 65 µm; E, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3741586" httpUri="https://zenodo.org/record/3741586/files/figure.png" pageId="35" pageNumber="260">Fig. 18D</figureCitation>
[insets]).
</paragraph>
<paragraph id="496036ABFFF8FFD2FC97FC94DFF2FC98" blockId="35.[813,1456,217,1040]" pageId="35" pageNumber="260">
Posterior region tapered into a blunt cone (
<figureCitation id="D1E42A2EFFF8FFD2FAC3FC94DF35FCB8" box="[1303,1399,823,849]" captionStart="FIG" captionStartId="34.[133,144,1229,1247]" captionTargetBox="[172,1417,215,1184]" captionTargetId="figure@34.[172,1417,214,1184]" captionTargetPageId="34" captionText="FIG. 18. — Hesione hartmanae n. sp., holotype, LACM 10158: A, anterior region, dorsal view; B, prostomium, dorsal view (arrows point to antennae); C, chaetiger 8, right parapodium, anterior view (inset: acicular lobe); D, same, neurochaetal blades (insets: tips of blades); E, posterior region, dorsal view. Scale bars: A, 2.2 mm; B, 0.3 mm; C, 0.6 mm; D, 65 µm; E, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3741586" httpUri="https://zenodo.org/record/3741586/files/figure.png" pageId="35" pageNumber="260">Fig. 18E</figureCitation>
); pygidium swollen, slightly depressed, anus with 6 digitate papillae.
</paragraph>
<paragraph id="496036ABFFF8FFD2FC97FCD4D9D6FBF9" blockId="35.[813,1456,217,1040]" pageId="35" pageNumber="260">
Pharynx not everted in
<typeStatus id="96648809FFF8FFD2FB99FCD4DEECFC78" box="[1101,1198,887,913]" pageId="35" pageNumber="260" type="holotype">holotype</typeStatus>
(smaller
<typeStatus id="96648809FFF8FFD2FAC3FCD4DF37FC78" box="[1303,1397,887,913]" pageId="35" pageNumber="260" type="paratype">paratype</typeStatus>
with pharynx almost fully everted, margin slightly eroded, made up by three rings, basal one slightly corrugated, dorsal papillae rounded, as long as wide). Oocytes about
<quantity id="8E279B4EFFF8FFD2FAEFFC74DFD0FC18" box="[1339,1426,983,1009]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="35" pageNumber="260" unit="mm" value="100.0">100 µm</quantity>
in diameter.
</paragraph>
<paragraph id="496036ABFFF8FFD2FCF9FB94D9CDFBB9" blockId="35.[813,1456,1079,1392]" box="[813,911,1079,1104]" pageId="35" pageNumber="260">
<emphasis id="7BABEAB9FFF8FFD2FCF9FB94D9CDFBB9" box="[813,911,1079,1104]" italics="true" pageId="35" pageNumber="260">Variation</emphasis>
</paragraph>
<paragraph id="496036ABFFF8FFD2FCF9FBF5DE46FA86" blockId="35.[813,1456,1079,1392]" pageId="35" pageNumber="260">
A rather weak dorsal darkening in one of the smaller
<typeStatus id="96648809FFF8FFD2FAACFBF4D925FB78" pageId="35" pageNumber="260" type="paratype">paratypes</typeStatus>
, even less defined in largest
<typeStatus id="96648809FFF8FFD2FB5BFBD4DEADFB78" box="[1167,1263,1143,1169]" pageId="35" pageNumber="260" type="paratype">paratype</typeStatus>
. Prostomial shape varies depending on pharynx exposure from having a squarrish outlook, with lateral margins slightly divergent in smallest
<typeStatus id="96648809FFF8FFD2FCF9FB75D9CBFB19" box="[813,905,1238,1264]" pageId="35" pageNumber="260" type="paratype">paratype</typeStatus>
(
<figureCitation id="D1E42A2EFFF8FFD2FC43FB75DE57FB19" box="[919,1045,1238,1264]" captionStart="FIG" captionStartId="36.[132,143,1151,1169]" captionTargetBox="[172,1416,216,1105]" captionTargetId="figure@36.[172,1416,216,1105]" captionTargetPageId="36" captionText="FIG. 19. — Hesione hartmanae n. sp., paratypes, LACM 10159: A, smallest paratype, anterior region, dorsal view; B, same, prostomium and everted pharynx (arrow points to dorsal papilla); C, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe); D, largest paratype, anterior region, slightly oblique dorsal view; E, same,prostomium slightly oblique dorsal view; F, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe). Scale bars: A, 1.3 mm; B, 0.7 mm; C, 0.4 mm; D, 1.8 mm; E, 0.3 mm; F, 0.6 mm." figureDoi="http://doi.org/10.5281/zenodo.3741588" httpUri="https://zenodo.org/record/3741588/files/figure.png" pageId="35" pageNumber="260">Fig. 19A, B</figureCitation>
), to one with rounded lateral margins, markedly divergent if the pharynx is not exposed (
<figureCitation id="D1E42A2EFFF8FFD2FA92FB56D903FAD9" captionStart="FIG" captionStartId="36.[132,143,1151,1169]" captionTargetBox="[172,1416,216,1105]" captionTargetId="figure@36.[172,1416,216,1105]" captionTargetPageId="36" captionText="FIG. 19. — Hesione hartmanae n. sp., paratypes, LACM 10159: A, smallest paratype, anterior region, dorsal view; B, same, prostomium and everted pharynx (arrow points to dorsal papilla); C, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe); D, largest paratype, anterior region, slightly oblique dorsal view; E, same,prostomium slightly oblique dorsal view; F, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe). Scale bars: A, 1.3 mm; B, 0.7 mm; C, 0.4 mm; D, 1.8 mm; E, 0.3 mm; F, 0.6 mm." figureDoi="http://doi.org/10.5281/zenodo.3741588" httpUri="https://zenodo.org/record/3741588/files/figure.png" pageId="35" pageNumber="260">Fig. 19D, E</figureCitation>
). In smaller
<typeStatus id="96648809FFF8FFD2FC17FAB5DE69FAD9" box="[963,1067,1302,1328]" pageId="35" pageNumber="260" type="paratype">paratypes</typeStatus>
, acicular lobes with upper tine twice as long as lower ones (
<figureCitation id="D1E42A2EFFF8FFD2FBC9FA96DEC2FAA6" box="[1053,1152,1333,1359]" captionStart="FIG" captionStartId="36.[132,143,1151,1169]" captionTargetBox="[172,1416,216,1105]" captionTargetId="figure@36.[172,1416,216,1105]" captionTargetPageId="36" captionText="FIG. 19. — Hesione hartmanae n. sp., paratypes, LACM 10159: A, smallest paratype, anterior region, dorsal view; B, same, prostomium and everted pharynx (arrow points to dorsal papilla); C, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe); D, largest paratype, anterior region, slightly oblique dorsal view; E, same,prostomium slightly oblique dorsal view; F, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe). Scale bars: A, 1.3 mm; B, 0.7 mm; C, 0.4 mm; D, 1.8 mm; E, 0.3 mm; F, 0.6 mm." figureDoi="http://doi.org/10.5281/zenodo.3741588" httpUri="https://zenodo.org/record/3741588/files/figure.png" pageId="35" pageNumber="260">Fig. 19C</figureCitation>
), or of similar size in largest
<typeStatus id="96648809FFF8FFD2FCF9FAF5D9CBFA99" box="[813,905,1366,1392]" pageId="35" pageNumber="260" type="paratype">paratype</typeStatus>
(
<figureCitation id="D1E42A2EFFF8FFD2FC4CFAF6D9B5FA86" box="[920,1015,1365,1391]" captionStart="FIG" captionStartId="36.[132,143,1151,1169]" captionTargetBox="[172,1416,216,1105]" captionTargetId="figure@36.[172,1416,216,1105]" captionTargetPageId="36" captionText="FIG. 19. — Hesione hartmanae n. sp., paratypes, LACM 10159: A, smallest paratype, anterior region, dorsal view; B, same, prostomium and everted pharynx (arrow points to dorsal papilla); C, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe); D, largest paratype, anterior region, slightly oblique dorsal view; E, same,prostomium slightly oblique dorsal view; F, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe). Scale bars: A, 1.3 mm; B, 0.7 mm; C, 0.4 mm; D, 1.8 mm; E, 0.3 mm; F, 0.6 mm." figureDoi="http://doi.org/10.5281/zenodo.3741588" httpUri="https://zenodo.org/record/3741588/files/figure.png" pageId="35" pageNumber="260">Fig. 19F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="01C56520FFF8FFD5FCF9FA37DB32F804" lastPageId="36" lastPageNumber="261" pageId="35" pageNumber="260" type="discussion">
<paragraph id="496036ABFFF8FFD2FCF9FA37D9D3FA44" blockId="35.[813,1456,1428,2030]" box="[813,913,1428,1454]" pageId="35" pageNumber="260">REMARKS</paragraph>
<paragraph id="496036ABFFF8FFD2FCF9FA15DE0DF864" blockId="35.[813,1456,1428,2030]" pageId="35" pageNumber="260">
<taxonomicName id="8EDF4D28FFF8FFD2FCF9FA15D9BBFA26" authority="Salazar-Vallejo, 2018" authorityName="Salazar-Vallejo" authorityYear="2018" box="[813,1017,1461,1487]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="35" pageNumber="260" phylum="Annelida" rank="species" species="hartmanae" status="sp. nov.">
<emphasis id="7BABEAB9FFF8FFD2FCF9FA15D9BBFA26" box="[813,1017,1461,1487]" italics="true" pageId="35" pageNumber="260">Hesione hartmanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="609857C2FFF8FFD2FBD5FA16DE7DFA26" box="[1025,1087,1461,1487]" pageId="35" pageNumber="260" rank="species">n. sp.</taxonomicNameLabel>
was likely recorded before under three different names.
<bibRefCitation id="2D4E4B5AFFF8FFD2FBF2FA76DEB1FA19" author="Monro" box="[1062,1267,1493,1520]" firstAuthor="Monro" pageId="35" pageNumber="260" pagination="1 - 96" refId="ref96564" refString="MONRO C. C. A. 1933. - The Polychaeta Errantia collected by Dr C. Crossland at Colon, in the Panama Region, and the Galapagos Islands during the Expedition of the S. Y. &quot; St. George &quot;. Proceedings of the Zoological Society, London 1933 (1): 1 - 96. http: // onlinelibrary. wiley. com / doi / 10.1111 / j. 1096 - 3642.1933. tb 01578. x / full" type="journal article" year="1933">Monro (1933: 26)</bibRefCitation>
cited what Cyril Crossland noticed about the pigmentation of some Panamian
<taxonomicName id="8EDF4D28FFF8FFD2FCBBF9B6D981F9C7" authorityName="Savigny in Lamarck" authorityYear="1818" box="[879,963,1557,1582]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="35" pageNumber="260" phylum="Annelida" rank="genus">
<emphasis id="7BABEAB9FFF8FFD2FCBBF9B6D981F9C7" box="[879,963,1557,1582]" italics="true" pageId="35" pageNumber="260">Hesione</emphasis>
</taxonomicName>
: “There are five transverse brown bands of irregular outline anteriorly, the first on the anterior border of the peristomium, the others occupying the anterior third of each segment, the middle third being white, while the posterior third is a light brown but darker than the general body color behind the fifth segment.” Monro referred that specimen to
<taxonomicName id="8EDF4D28FFF8FFD2FC62F976DF23F907" authority="Savigny in Lamarck, 1818" authorityName="Savigny in Lamarck" authorityYear="1818" box="[950,1377,1748,1774]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="35" pageNumber="260" phylum="Annelida" rank="species" species="splendida">
<emphasis id="7BABEAB9FFF8FFD2FC62F976DE79F907" box="[950,1083,1748,1774]" italics="true" pageId="35" pageNumber="260">H. splendida</emphasis>
Savigny
<emphasis id="7BABEAB9FFF8FFD2FB49F976DEF1F907" box="[1181,1203,1749,1774]" italics="true" pageId="35" pageNumber="260">in</emphasis>
<bibRefCitation id="2D4E4B5AFFF8FFD2FB6EF977DF23F907" author="Lamarck" box="[1210,1377,1748,1774]" firstAuthor="Lamarck" pageId="35" pageNumber="260" refId="ref95809" refString="LAMARCK J. B. 1818. - Histoire Naturelle des Animaux sans Vertebres, presentant les Caracteres Generaux et Particuliers de ces Animaux, leur Distribution, leurs Classes, leurs Familles, leurs Genres, et la Citation des principales Especes qui s'y rapportent; Precedes d'une Introduction offrant la Determination des Caracteres Essentiels de l'Animal, sa distinction du Vegetal et des autres Corps Naturels, en fin, l'Exposition des Principes fondamentaux de la Zoologie, Deterville &amp; Verdiere, Paris, volume 5, 612 p. http: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 64280058" type="book" year="1818">Lamarck, 1818</bibRefCitation>
</taxonomicName>
, which had been observed alive and described with a pearly iridescent body, but without pigmentation. It must be noted that this Panamian specimen had been less than 10 years in ethanol before Monro studied it, and he indicated there was no pigmentation left on its body.
</paragraph>
<paragraph id="496036ABFFF8FFD5FC97F837DB85F8E7" blockId="35.[813,1456,1428,2030]" lastBlockId="36.[132,776,1301,2030]" lastPageId="36" lastPageNumber="261" pageId="35" pageNumber="260">
<bibRefCitation id="2D4E4B5AFFF8FFD2FC97F837DE6EF844" author="Hartman" box="[835,1068,1939,1966]" firstAuthor="Hartman" pageId="35" pageNumber="260" pagination="173 - 287" refId="ref94489" refString="HARTMAN O. 1940. - Polychaetous annelids, 2. Chrysopetalidae to Goniadidae. Allan Hancock Pacific Expeditions 7: 173 - 287, pls 31 - 34. http: // cdm 15799. contentdm. oclc. org / cdm / ref / collection / p 15799 coll 82 / id / 5767" type="journal article" year="1940">Hartman (1940: 212)</bibRefCitation>
preferred to use
<taxonomicName id="8EDF4D28FFF8FFD2FB0EF837D92AF824" authority="Grube, 1878" authorityName="Grube" authorityYear="1878" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="35" pageNumber="260" phylum="Annelida" rank="species" species="intertexta">
<emphasis id="7BABEAB9FFF8FFD2FB0EF837DF1FF844" box="[1242,1373,1940,1965]" italics="true" pageId="35" pageNumber="260">H. intertexta</emphasis>
<bibRefCitation id="2D4E4B5AFFF8FFD2FAB7F830D92AF824" author="Grube" firstAuthor="Grube" pageId="35" pageNumber="260" pagination="1 - 300" refId="ref94269" refString="GRUBE A. E. 1878. - Annulata Semperiana. Beitrage zur Kenntnis der Annelidenfauna der Philippinen nach den von Herrn Prof. Semper mitgebrachten Sammlungen. Memoires de l'Academie Imperiale des Sciences de St-Petersbourg (serie 7) 25 (8): 1 - 300, pls 1 - 15." type="journal article" year="1878">Grube, 1878</bibRefCitation>
</taxonomicName>
, and, as indicated below in the remarks for
<taxonomicName id="8EDF4D28FFF8FFD2FA99F817DE79F804" authority="Chamberlin, 1919" authorityName="Chamberlin" authorityYear="1919" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="35" pageNumber="260" phylum="Annelida" rank="species" species="panamena">
<emphasis id="7BABEAB9FFF8FFD2FA99F817D924F804" italics="true" pageId="35" pageNumber="260">H. panamena</emphasis>
<bibRefCitation id="2D4E4B5AFFF8FFD2FCB9F870DE79F804" author="Chamberlin" box="[877,1083,2003,2029]" firstAuthor="Chamberlin" pageId="35" pageNumber="260" pagination="1 - 514" refId="ref91326" refString="CHAMBERLIN R. V. 1919. - The Annelida Polychaeta of the Albatross Tropical Pacific Expedition, 1891 - 1905. Memoirs of the Museum of Comparative Zoology of Harvard College 31: 1 - 514. https: // doi. org / 10.5962 / bhl. title. 49195" type="journal article" year="1919">Chamberlin, 1919</bibRefCitation>
</taxonomicName>
, her illustrations show an acicular lobe double with tines of similar size, which are not present in
<taxonomicName id="8EDF4D28FFFFFFD5FF76FA95DB69FAA6" authorityName="Grube" authorityYear="1878" box="[162,299,1334,1359]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="261" phylum="Annelida" rank="species" species="intertexta">
<emphasis id="7BABEAB9FFFFFFD5FF76FA95DB69FAA6" box="[162,299,1334,1359]" italics="true" pageId="36" pageNumber="261">H. intertexta</emphasis>
</taxonomicName>
, meaning she had two species in her material. It seems she was unaware of the previous indications by
<bibRefCitation id="2D4E4B5AFFFFFFD5FF50FAD5DB0FFA66" author="Horst" box="[132,333,1397,1424]" firstAuthor="Horst" pageId="36" pageNumber="261" pagination="145 - 198" refId="ref95128" refString="HORST R. 1924. - Polychaeta Errantia of the Siboga - Expedition, 3. Nereidae and Hesionidae. Siboga-Expeditie Monographs 24 (1 c): 145 - 198." type="journal article" year="1924">Horst (1924: 193)</bibRefCitation>
, who emphasised that acicular lobes were of very different size and shape in
<taxonomicName id="8EDF4D28FFFFFFD5FDDEFA35D8D4FA46" authorityName="Grube" authorityYear="1878" box="[522,662,1430,1455]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="261" phylum="Annelida" rank="species" species="intertexta">
<emphasis id="7BABEAB9FFFFFFD5FDDEFA35D8D4FA46" box="[522,662,1430,1455]" italics="true" pageId="36" pageNumber="261">H. intertexta</emphasis>
</taxonomicName>
: a longer, digitate and the other one shorter, rounded. Different sized tines are also present in
<taxonomicName id="8EDF4D28FFFFFFD5FE46FA76D867FA07" authorityName="Chamberlin" authorityYear="1919" box="[402,549,1493,1518]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="261" phylum="Annelida" rank="species" species="panamena">
<emphasis id="7BABEAB9FFFFFFD5FE46FA76D867FA07" box="[402,549,1493,1518]" italics="true" pageId="36" pageNumber="261">H. panamena</emphasis>
</taxonomicName>
, as indicated above. Hartman made no indication of transverse bands but stated that in her material: “most individuals retain the reticulated, fulvous pattern on the dorsum of the anterior segments, but in some there are almost none.” This fading is typical for those species having longitudinal thin lines along the body. In some of the specimens regarded as belonging to
<taxonomicName id="8EDF4D28FFFFFFD5FD7AF936DA88F927" authority="Salazar-Vallejo, 2018" authorityName="Salazar-Vallejo" authorityYear="2018" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="261" phylum="Annelida" rank="species" species="hartmanae" status="sp. nov.">
<emphasis id="7BABEAB9FFFFFFD5FD7AF936DA88F927" italics="true" pageId="36" pageNumber="261">H. hartmanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="609857C2FFFFFFD5FF06F916DB53F926" box="[210,273,1717,1743]" pageId="36" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
, there is a fine indication for darker tranverse areas in anterior segments, despite the fact they have been in ethanol for about 70 years.
</paragraph>
<caption id="1DA06623FFFFFFD5FF50FBDCD81CFB3C" ID-DOI="http://doi.org/10.5281/zenodo.3741588" ID-Zenodo-Dep="3741588" httpUri="https://zenodo.org/record/3741588/files/figure.png" pageId="36" pageNumber="261" startId="36.[132,143,1151,1169]" targetBox="[172,1416,216,1105]" targetPageId="36">
<paragraph id="496036ABFFFFFFD5FF50FBDCD81CFB3C" blockId="36.[132,1456,1150,1237]" pageId="36" pageNumber="261">
FIG. 19. —
<taxonomicName id="8EDF4D28FFFFFFD5FF35FBDCDBC1FB78" authority="Salazar-Vallejo, 2018" authorityName="Salazar-Vallejo" authorityYear="2018" box="[225,387,1151,1169]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="261" phylum="Annelida" rank="species" species="hartmanae" status="sp. nov.">
<emphasis id="7BABEAB9FFFFFFD5FF35FBDCDBC1FB78" box="[225,387,1151,1169]" italics="true" pageId="36" pageNumber="261">Hesione hartmanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="609857C2FFFFFFD5FE5DFBDCDBFFFB78" box="[393,445,1151,1169]" pageId="36" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
, paratypes, LACM 10159:
<emphasis id="7BABEAB9FFFFFFD5FD49FBDCD8E8FB78" bold="true" box="[669,682,1151,1169]" pageId="36" pageNumber="261">A</emphasis>
, smallest paratype, anterior region, dorsal view;
<emphasis id="7BABEAB9FFFFFFD5FB91FBDCDE10FB78" bold="true" box="[1093,1106,1151,1169]" pageId="36" pageNumber="261">B</emphasis>
, same, prostomium and everted pharynx (arrow points to dorsal papilla);
<emphasis id="7BABEAB9FFFFFFD5FE59FB36DBD9FB4E" bold="true" box="[397,411,1173,1191]" pageId="36" pageNumber="261">C</emphasis>
, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe);
<emphasis id="7BABEAB9FFFFFFD5FBD7FB35DE53FB41" bold="true" box="[1027,1041,1174,1192]" pageId="36" pageNumber="261">D</emphasis>
, largest paratype, anterior region, slightly oblique dorsal view;
<emphasis id="7BABEAB9FFFFFFD5FF38FB0FDABAFB57" bold="true" box="[236,248,1196,1214]" pageId="36" pageNumber="261">E</emphasis>
, same,prostomium slightly oblique dorsal view;
<emphasis id="7BABEAB9FFFFFFD5FD5FFB0FD8D4FB57" bold="true" box="[651,662,1196,1214]" pageId="36" pageNumber="261">F</emphasis>
, same, chaetiger 9, right parapodium, anterior view (inset: acicular lobe). Scale bars: A, 1.3 mm; B, 0.7 mm; C, 0.4 mm; D, 1.8 mm; E, 0.3 mm; F, 0.6 mm.
</paragraph>
</caption>
<paragraph id="496036ABFFFFFFD5FF4FF8B7DB32F804" blockId="36.[132,776,1301,2030]" pageId="36" pageNumber="261">
As indicated in the key below,
<taxonomicName id="8EDF4D28FFFFFFD5FE37F8B7D83AF8C7" authority="Salazar-Vallejo, 2018" authorityName="Salazar-Vallejo" authorityYear="2018" box="[483,632,1812,1838]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="261" phylum="Annelida" rank="species" species="hartmanae" status="sp. nov.">
<emphasis id="7BABEAB9FFFFFFD5FE37F8B7D83AF8C7" box="[483,632,1812,1838]" italics="true" pageId="36" pageNumber="261">H. hartmanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="609857C2FFFFFFD5FDAAF8B7D8FBF8C7" box="[638,697,1812,1838]" pageId="36" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="8EDF4D28FFFFFFD5FF67F897DBDAF8A7" authority="Muller, 1858" authorityName="Muller" authorityYear="1858" box="[179,408,1843,1870]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="261" phylum="Annelida" rank="species" species="picta">
<emphasis id="7BABEAB9FFFFFFD5FF67F897DB44F8A4" box="[179,262,1844,1869]" italics="true" pageId="36" pageNumber="261">H. picta</emphasis>
<bibRefCitation id="2D4E4B5AFFFFFFD5FED8F890DBDAF8A7" author="Muller" box="[268,408,1843,1870]" firstAuthor="Muller" pageId="36" pageNumber="261" pagination="211 - 220" refId="ref96748" refString="MULLER F. 1858. - Einiges uber die Annelidenfauna der Insel Santa Catharina an der brasilianischen Kuste. Archiv fur Naturgeschichte 24: 211 - 220, pls 6 - 7. https: // biodiversitylibrary. org / page / 7460059" type="journal article" year="1858">Müller, 1858</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="8EDF4D28FFFFFFD5FE1FF897DA82F887" authority="von Marenzeller, 1879" authorityName="von Marenzeller" authorityYear="1879" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="261" phylum="Annelida" rank="species" species="reticulata">
<emphasis id="7BABEAB9FFFFFFD5FE1FF897D80DF8A4" box="[459,591,1843,1869]" italics="true" pageId="36" pageNumber="261">H. reticulata</emphasis>
<bibRefCitation id="2D4E4B5AFFFFFFD5FD80F897DA82F887" author="von Marenzeller" firstAuthor="von Marenzeller" pageId="36" pageNumber="261" pagination="109 - 154" refId="ref98946" refString="VON MARENZELLER E. 1879. - Sudjapanische Anneliden, 1. Amphinomea, Aphroditea, Lycoridea, Phyllodocea, Hesionea, Syllidea, Eunicea, Glycerea, Sternaspidea, Chaetopterea, Cirratulea, Amphictenea. Denskschriften der Mathematisch-Naturwissenschaftlichen classe der Kaiserlichen Akademie der Wissenschaften 41 (2): 109 - 154. https: // biodiversitylibrary. org / page / 7215498" type="journal article" year="1879">von Marenzeller, 1879</bibRefCitation>
</taxonomicName>
. Their main difference is the length of the acicular lobe in comparison to the width of the corresponding chaetal lobe. In
<taxonomicName id="8EDF4D28FFFFFFD5FF35F837DB35F844" authority="Salazar-Vallejo, 2018" authorityName="Salazar-Vallejo" authorityYear="2018" box="[225,375,1939,1965]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="261" phylum="Annelida" rank="species" species="hartmanae" status="sp. nov.">
<emphasis id="7BABEAB9FFFFFFD5FF35F837DB35F844" box="[225,375,1939,1965]" italics="true" pageId="36" pageNumber="261">H. hartmanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="609857C2FFFFFFD5FEAAF837DBF8F847" box="[382,442,1940,1966]" pageId="36" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
it is longer, about half as long as chaetal fascicle width, whereas it is shorter (1/3 as long) in the two other species.
</paragraph>
</subSubSection>
</treatment>
</document>