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<document id="AE454E02AB2F53DA0630D5072CEABC61" ID-DOI="10.5281/zenodo.186268" ID-GBIF-Dataset="70978291-4fff-46ac-93c9-8c2f50900af1" ID-ISSN="1175-5326" ID-Zenodo-Dep="186268" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461049396867" checkinUser="plazi" docAuthor="Anlaş, Sinan" docDate="2009" docId="C1192B1A6414BE55FF32EB51B1445BB1" docLanguage="en" docName="zt02031p067.pdf" docOrigin="Zootaxa 2031" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tetartopeus tezcani Anlaş, 2009, sp. n." docType="treatment" docVersion="7" lastPageNumber="67" masterDocId="3D2053626416BE51FFA5EF6BB5555A56" masterDocTitle="On the genus Tetartopeus Czwalina, 1888 of Turkey. A new species and additional records (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="67" masterPageNumber="63" pageNumber="65" updateTime="1698578179513" updateUser="plazi">
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<mods:title id="C892E8C4D90C3016984707B6EA02DF3E">On the genus Tetartopeus Czwalina, 1888 of Turkey. A new species and additional records (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:namePart id="8B407B4B3C8B0BD6487A4B5A43F37CF9">Anlaş, Sinan</mods:namePart>
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<mods:date id="4DCC77C4DAC08C6B8B24E3BAF9487C5B">2009</mods:date>
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<treatment id="C1192B1A6414BE55FF32EB51B1445BB1" ID-DOI="http://doi.org/10.5281/zenodo.6225550" ID-GBIF-Taxon="119611506" ID-Zenodo-Dep="6225550" LSID="urn:lsid:plazi:treatment:C1192B1A6414BE55FF32EB51B1445BB1" httpUri="http://treatment.plazi.org/id/C1192B1A6414BE55FF32EB51B1445BB1" lastPageId="4" lastPageNumber="67" pageId="2" pageNumber="65">
<subSubSection id="01AAC9876414BE53FF32EB51B4735E20" pageId="2" pageNumber="65" type="nomenclature">
<paragraph id="490F9A0C6414BE53FF32EB51B49D5E05" blockId="2.[151,456,1081,1142]" box="[151,456,1081,1108]" pageId="2" pageNumber="65">
<heading id="12472D606414BE53FF32EB51B49D5E05" bold="true" box="[151,456,1081,1108]" fontSize="11" level="1" pageId="2" pageNumber="65" reason="1">
<emphasis id="7BC4461E6414BE53FF32EB51B49D5E05" bold="true" box="[151,456,1081,1108]" pageId="2" pageNumber="65">
<taxonomicName id="8EB0E18F6414BE53FF32EB51B42A5E02" ID-CoL="55K8M" box="[151,383,1082,1108]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="tezcani" status="sp. nov.">
<emphasis id="7BC4461E6414BE53FF32EB51B42A5E02" bold="true" box="[151,383,1082,1108]" italics="true" pageId="2" pageNumber="65">Tetartopeus tezcani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60F7FB656414BE53FE23EB52B49D5E05" box="[390,456,1081,1107]" pageId="2" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="490F9A0C6414BE53FF32EB37B4735E20" blockId="2.[151,456,1081,1142]" box="[151,294,1116,1142]" pageId="2" pageNumber="65">
(
<figureCitation id="D18B86896414BE53FF3AEB37B4485E20" box="[159,285,1116,1142]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="2" pageNumber="65">Figs. 112</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="01AAC9876414BE53FF32EBC2B0125F47" pageId="2" pageNumber="65" type="materials_examined">
<paragraph id="490F9A0C6414BE53FF32EBC2B0125F47" blockId="2.[151,1437,1192,2032]" pageId="2" pageNumber="65">
<emphasis id="7BC4461E6414BE53FF32EBC2B73A5E95" bold="true" box="[151,623,1193,1219]" pageId="2" pageNumber="65">
<typeStatus id="960B24AE6414BE53FF32EBC2B5825E95" box="[151,215,1193,1219]" pageId="2" pageNumber="65">Type</typeStatus>
material.
<typeStatus id="960B24AE6414BE53FEFBEBC2B48C5E95" box="[350,473,1193,1219]" pageId="2" pageNumber="65" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="31A7DA9C6414BE53FE42EBC2B73F5E95" box="[487,618,1193,1219]" name="Turkey" pageId="2" pageNumber="65">TURKEY</collectingCountry>
:
</emphasis>
3, “
<collectingCountry id="31A7DA9C6414BE53FD0FEBC1B7835E92" box="[682,726,1194,1220]" name="Chile" pageId="2" pageNumber="65">TR</collectingCountry>
. Adıyaman, Gölbaşı, Karakuyu village,
<quantity id="8E4837E96414BE53FB62EBC1B0505E92" box="[1223,1285,1194,1220]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="65" unit="km" value="1.0">1 km</quantity>
W,
<quantity id="8E4837E96414BE53FA9DEBC1B0C25E92" box="[1336,1431,1194,1220]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="2" pageNumber="65" unit="m" value="1210.0">1210 m</quantity>
, 37˚4152”N, 37˚3814”E,
<date id="3D0EBCCC6414BE53FE7DEBBAB7075EBD" box="[472,594,1233,1259]" pageId="2" pageNumber="65" value="2008-04-05">5.IV.2008</date>
, leg. Yaġmur /
<typeStatus id="960B24AE6414BE53FCABEBBAB6D95EBD" box="[782,908,1233,1259]" pageId="2" pageNumber="65" type="holotype">Holotypus</typeStatus>
3,
<taxonomicName id="8EB0E18F6414BE53FC19EBBAB1FC5EBC" box="[956,1193,1233,1258]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="tezcani" status="sp. nov.">
<emphasis id="7BC4461E6414BE53FC19EBBAB1FC5EBC" box="[956,1193,1233,1258]" italics="true" pageId="2" pageNumber="65">Tetartopeus tezcani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60F7FB656414BE53FB14EBBBB1A25EBC" box="[1201,1271,1232,1258]" pageId="2" pageNumber="65" rank="species">
<emphasis id="7BC4461E6414BE53FB14EBBBB1A25EBC" bold="true" box="[1201,1271,1232,1258]" pageId="2" pageNumber="65">sp. n.</emphasis>
</taxonomicNameLabel>
det. S. Anlaş 2008” (cAnl).
<emphasis id="7BC4461E6414BE53FEE4EB9CB7065F47" bold="true" box="[321,595,1271,1297]" pageId="2" pageNumber="65">
<typeStatus id="960B24AE6414BE53FEE4EB9CB4965F47" box="[321,451,1271,1297]" pageId="2" pageNumber="65" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="31A7DA9C6414BE53FE6AEB9CB71B5F47" box="[463,590,1271,1297]" name="Turkey" pageId="2" pageNumber="65">TURKEY</collectingCountry>
:
</emphasis>
833, 10ƤƤ, same data as
<typeStatus id="960B24AE6414BE53FC32EB9CB6AB5F47" box="[919,1022,1271,1297]" pageId="2" pageNumber="65" type="holotype">holotype</typeStatus>
(
<collectionCode id="2FA102C96414BE53FBA8EB9CB1355F47" LSID="urn:lsid:biocol.org:col:33701" box="[1037,1120,1271,1297]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33701" name="Ege University, Lodos Entomological Museum" pageId="2" pageNumber="65">LEMT</collectionCode>
, cAnl, cAss, cYag).
</paragraph>
</subSubSection>
<subSubSection id="01AAC9876414BE55FF63EA76B0C95B1A" lastPageId="4" lastPageNumber="67" pageId="2" pageNumber="65" type="description">
<paragraph id="490F9A0C6414BE53FF63EA76B07A5F85" blockId="2.[151,1437,1192,2032]" pageId="2" pageNumber="65">
<emphasis id="7BC4461E6414BE53FF63EA76B40B5F61" bold="true" box="[198,350,1309,1335]" pageId="2" pageNumber="65">Description.</emphasis>
Measurements (in mm) and ratios (range, arithmetic mean; n=19): AL: 1.772.15, 1.91; HL: 0.720.85, 0.78; HW: 0.680.75, 0.71; PW: 0.740.84, 0.78; PL: 0.861.00, 0.92; EL: 0.790.95, 0.87; EW: 0.841.00, 0.92; AW: 0.820.91, 0.86; TiL: 0.810.91, 0.87; TaL: 0.520.66, 0.60; ML: 0.911.02, 0.95 (n=9); TL: 6.37.4, 6.8; HL/HW: 1.061.13, 1.10; PW/HW: 1.091.12, 1.10; PW/PL: 0.840.86, 0.85; EL/ PL: 0.920.95, 0.94; EW/PW: 1.141.19, 1.17; AW/EW: 0.910.98, 0.94; TiL/TaL: 1.381.55, 1.45.
</paragraph>
<paragraph id="490F9A0C6414BE53FF63EAB4B6615C10" blockId="2.[151,1437,1192,2032]" pageId="2" pageNumber="65">
Habitus as in
<figureCitation id="D18B86896414BE53FED4EAB4B4EF5FAF" box="[369,442,1503,1529]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="2" pageNumber="65">Fig. 1</figureCitation>
. Coloration: head and pronotum black; elytra red, with scutellar region infuscate; abdomen black, with posterior margin of segment VII and apical segments (VIIIX) reddish; legs yellowish brown; antennae reddish brown, partly slightly infuscate.
</paragraph>
<paragraph id="490F9A0C6414BE53FF63E938B0735D5E" blockId="2.[151,1437,1192,2032]" pageId="2" pageNumber="65">
Head oblong (see measurements, ratio HL/HW and
<figureCitation id="D18B86896414BE53FC8CE938B6CC5C3B" box="[809,921,1619,1645]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="2" pageNumber="65">Figs. 1, 7</figureCitation>
); eyes moderately large (
<figureCitation id="D18B86896414BE53FB64E938B05F5C3B" box="[1217,1290,1619,1645]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="2" pageNumber="65">Fig. 7</figureCitation>
), punctation moderately coarse and dense, interstices in lateral dorsal region on average approximately as wide as punctures; punctation in central dorsal region slightly sparser; microsculpture in central dorsal region absent, in posterior and lateral region very shallow. Antenna slender; antennomere III longer than II; antennomeres IVXI slightly more than twice as long as wide, of subequal length, and about as long as II (
<figureCitation id="D18B86896414BE53FB75E985B0415D5E" box="[1232,1300,1774,1800]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="2" pageNumber="65">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="490F9A0C6414BE53FF63E87FB07A5D03" blockId="2.[151,1437,1192,2032]" pageId="2" pageNumber="65">
Pronotum slightly wider than head (see ratio PW/HW,
<figureCitation id="D18B86896414BE53FCEBE87FB6EB5D78" box="[846,958,1812,1838]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="2" pageNumber="65">Figs. 1, 7</figureCitation>
) and oblong (see ratio PW/PL and
<figureCitation id="D18B86896414BE53FAC7E87FB5905D03" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="2" pageNumber="65">Figs. 1, 7</figureCitation>
); punctation similar to that of head but much denser and slightly more defined than that of head.
</paragraph>
<paragraph id="490F9A0C6414BE53FF63E809B1385DF5" blockId="2.[151,1437,1192,2032]" pageId="2" pageNumber="65">
Elytra wider and at suture slightly shorter than pronotum; (see ratios EW/PW, EL/PL and
<figureCitation id="D18B86896414BE53FAB0E809B0D95D2A" box="[1301,1420,1890,1916]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="2" pageNumber="65">Figs. 1, 7</figureCitation>
); punctation finer and shallower than that of head and pronotum.
<collectingCountry id="31A7DA9C6414BE53FC24E8E2B6E95DF5" box="[897,956,1929,1955]" name="India" pageId="2" pageNumber="65">Hind</collectingCountry>
wings present.
</paragraph>
<paragraph id="490F9A0C6414BE53FF63E8C4B6A45DA6" blockId="2.[151,1437,1192,2032]" pageId="2" pageNumber="65">
Abdomen narrower than elytra (see ratio AW/EW and
<figureCitation id="D18B86896414BE53FCFCE8C4B6985D9F" box="[857,973,1967,1993]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="2" pageNumber="65">Figs. 1, 7</figureCitation>
); punctation very dense and very fine, pubescense blackish. Posterior margin of tergite VII with palisade fringe.
</paragraph>
<paragraph id="490F9A0C6415BE52FF63EFFCB7815B70" blockId="3.[151,1436,151,332]" pageId="3" pageNumber="66">
3: sternite VII with clusters of dark modified setae, posterior margin weakly concave (
<figureCitation id="D18B86896415BE52FB47EFF3B0795AE4" box="[1250,1324,152,178]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="3" pageNumber="66">Fig. 5</figureCitation>
); sternite VIII with two extensive and oblong clusters of black modified setae (
<figureCitation id="D18B86896415BE52FC48EFD5B16F5A8E" box="[1005,1082,190,216]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="3" pageNumber="66">Fig. 6</figureCitation>
); aedeagus with long ventral process of characteristic shape (
<figureCitation id="D18B86896415BE52FDB6EF8EB7C05AA9" box="[531,661,229,255]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="3" pageNumber="66">Figs. 810</figureCitation>
), particularly distinct of triangular shape, apically in lateral view (
<figureCitation id="D18B86896415BE52FF3AEE67B5B35B70" box="[159,230,268,294]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="3" pageNumber="66">Fig. 8</figureCitation>
). Internal sac structures as in
<figureCitation id="D18B86896415BE52FDE6EE67B7855B70" box="[579,720,268,294]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="3" pageNumber="66">Figs. 1112</figureCitation>
.
</paragraph>
<paragraph id="490F9A0C6415BE52FF63EE59B1035B1A" blockId="3.[151,1436,151,332]" box="[198,1110,306,332]" pageId="3" pageNumber="66">
Ƥ: tergite and sternite VIII as in
<figureCitation id="D18B86896415BE52FDE1EE59B7E65B1A" box="[580,691,306,332]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="3" pageNumber="66">Figs. 2, 3</figureCitation>
; female terga IX and X as in
<figureCitation id="D18B86896415BE52FBAEEE59B1075B1A" box="[1035,1106,306,332]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="3" pageNumber="66">Fig. 4</figureCitation>
.
</paragraph>
<caption id="1DCFCA846415BE52FF32E85DB1AB5D8E" httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="3" pageNumber="66" targetBox="[233,1360,393,1819]" targetPageId="3">
<paragraph id="490F9A0C6415BE52FF32E85DB1AB5D8E" blockId="3.[151,1437,1846,2008]" pageId="3" pageNumber="66">
<emphasis id="7BC4461E6415BE52FF32E85DB4015D18" bold="true" box="[151,340,1846,1870]" pageId="3" pageNumber="66">FIGURES 114.</emphasis>
Details of
<taxonomicName id="8EB0E18F6415BE52FE75E85DB7F35D1B" box="[464,678,1846,1869]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="tezcani" status="sp. nov.">
<emphasis id="7BC4461E6415BE52FE75E85DB7F35D1B" box="[464,678,1846,1869]" italics="true" pageId="3" pageNumber="66">Tetartopeus tezcani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60F7FB656415BE52FD08E85DB7BE5D18" box="[685,747,1846,1870]" pageId="3" pageNumber="66" rank="species">
<emphasis id="7BC4461E6415BE52FD08E85DB7BE5D18" bold="true" box="[685,747,1846,1870]" pageId="3" pageNumber="66">sp. n.</emphasis>
</taxonomicNameLabel>
(112),
<taxonomicName id="8EB0E18F6415BE52FCEEE85DB1725D1B" authority="Assing" authorityName="Assing" box="[843,1063,1846,1869]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="adanensis">
<emphasis id="7BC4461E6415BE52FCEEE85DB6865D1B" box="[843,979,1846,1869]" italics="true" pageId="3" pageNumber="66">T. adanensis</emphasis>
Assing
</taxonomicName>
(13) and
<taxonomicName id="8EB0E18F6415BE52FB36E85DB0355D1B" authority="(Reitter)" authorityName="Reitter" baseAuthorityName="Reitter" box="[1171,1376,1846,1869]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="stylifer">
<emphasis id="7BC4461E6415BE52FB36E85DB1AE5D1B" box="[1171,1275,1846,1869]" italics="true" pageId="3" pageNumber="66">T. stylifer</emphasis>
(Reitter)
</taxonomicName>
(14). 1—habitus; 2—female tergite VIII; 3—female sternite VIII; 4—female terga IX and X; 5—male sternite VII; 6—male sternite VIII; 7—forebody; 8—apex of aedeagus, lateral view; 9, 13, 14—aedeagus, lateral view; 10—aedeagus, ventral view; 11—retracted internal sac with sclerites, lateral view; 12—retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1.0 mm (1, 7); 0.3 mm (26); 0,04 mm (8); 0.5 mm (910); 0,1 mm (1112); 0,2 mm (1314).
</paragraph>
</caption>
<paragraph id="490F9A0C6412BE55FF63EFFCB0C95B1A" blockId="4.[151,1436,151,487]" pageId="4" pageNumber="67">
<emphasis id="7BC4461E6412BE55FF63EFFCB4945AE7" bold="true" box="[198,449,151,177]" pageId="4" pageNumber="67">Comparative notes.</emphasis>
The species is distinguished from all its congeners by the male primary and secondary sexual characters, especially by the distinctive shape of the apex of the ventral process of the aedeagus. From other species occurring in central-eastern and central-southeastern
<collectingCountry id="31A7DA9C6412BE55FBD2EF8EB1985AA9" box="[1143,1229,229,255]" name="Turkey" pageId="4" pageNumber="67">Turkey</collectingCountry>
, it is additionally separated as follows: from
<taxonomicName id="8EB0E18F6412BE55FE70EE67B7325B73" box="[469,615,268,293]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="adanensis">
<emphasis id="7BC4461E6412BE55FE70EE67B7325B73" box="[469,615,268,293]" italics="true" pageId="4" pageNumber="67">T. adanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="8EB0E18F6412BE55FD05EE67B65B5B73" box="[672,782,268,293]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="stylifer">
<emphasis id="7BC4461E6412BE55FD05EE67B65B5B73" box="[672,782,268,293]" italics="true" pageId="4" pageNumber="67">T. stylifer</emphasis>
</taxonomicName>
by distincly smaller size; from
<taxonomicName id="8EB0E18F6412BE55FB26EE67B1AA5B73" box="[1155,1279,268,293]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="7BC4461E6412BE55FB26EE67B1AA5B73" box="[1155,1279,268,293]" italics="true" pageId="4" pageNumber="67">T. persicus</emphasis>
</taxonomicName>
by the shape of head. For illustrations of the habitus and genitalia of these species see
<figureCitation id="D18B86896412BE55FC40EE59B13B5B1A" box="[997,1134,306,332]" captionStart="FIGURES 1 14" captionStartId="3.[151,272,1846,1870]" captionTargetBox="[233,1360,393,1819]" captionTargetId="figure@3.[227,1361,390,1820]" captionTargetPageId="3" captionText="FIGURES 1 14. Details of Tetartopeus tezcani sp. n. (1 12), T. adanensis Assing (13) and T. stylifer (Reitter) (14). 1 — habitus; 2 — female tergite VIII; 3 — female sternite VIII; 4 — female terga IX and X; 5 — male sternite VII; 6 — male sternite VIII; 7 — forebody; 8 — apex of aedeagus, lateral view; 9, 13, 14 — aedeagus, lateral view; 10 — aedeagus, ventral view; 11 — retracted internal sac with sclerites, lateral view; 12 — retracted internal sac with sclerites, dorso-lateral view. Scale bars: 1. 0 mm (1, 7); 0. 3 mm (2 6); 0,04 mm (8); 0. 5 mm (9 10); 0,1 mm (11 12); 0,2 mm (13 14)." httpUri="https://zenodo.org/record/186269/files/figure.png" pageId="4" pageNumber="67">Figs. 13, 14</figureCitation>
and
<bibRefCitation id="2D21E7FD6412BE55FB03EE59B0105B1A" author="Assing" box="[1190,1349,306,332]" pageId="4" pageNumber="67" refString="Assing, V. (2004) New species and records of Staphylinidae from Turkey III (Insecta: Coleoptera). Linzer biologische Beitrage, 36, 669 - 733." type="journal article" year="2004">Assing (2004</bibRefCitation>
,
<bibRefCitation id="2D21E7FD6412BE55FAF5EE59B0DA5B1A" author="Assing" box="[1360,1423,306,332]" pageId="4" pageNumber="67" refString="Assing, V. (2008) On the taxonomy and zoogeography of some Palaearctic Paederinae and Xantholinini (Coleoptera: Staphylinidae). Linzer biologische Beitrage, 40, 1237 - 1294." type="journal article" year="2008">2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="01AAC9876412BE55FF63EE33B6065BCC" pageId="4" pageNumber="67" type="etymology">
<paragraph id="490F9A0C6412BE55FF63EE33B6065BCC" blockId="4.[151,1436,151,487]" pageId="4" pageNumber="67">
<emphasis id="7BC4461E6412BE55FF63EE33B4065B24" bold="true" box="[198,339,344,370]" pageId="4" pageNumber="67">Etymology.</emphasis>
The species is dedicated to Prof. Dr. Serdar Tezcan, Izmir, a specialist on
<taxonomicName id="8EB0E18F6412BE55FB68EE32B0095B25" box="[1229,1372,345,371]" class="Insecta" family="Buprestidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="family">Buprestidae</taxonomicName>
, who has carried out important entomological research in
<collectingCountry id="31A7DA9C6412BE55FD5FEEEBB6055BCC" box="[762,848,384,410]" name="Turkey" pageId="4" pageNumber="67">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="01AAC9876412BE55FF63EECDB1445BB1" pageId="4" pageNumber="67" type="distribution">
<paragraph id="490F9A0C6412BE55FF63EECDB1445BB1" blockId="4.[151,1436,151,487]" pageId="4" pageNumber="67">
<emphasis id="7BC4461E6412BE55FF63EECDB77C5B96" bold="true" box="[198,553,422,448]" pageId="4" pageNumber="67">Distribution and bionomics.</emphasis>
The species was collected in only one locality in Adıyaman (Map 1), in moist grassland near edges of small standing water at an altitude of
<quantity id="8E4837E96412BE55FC14EEA6B1585BB1" box="[945,1037,461,487]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="4" pageNumber="67" unit="m" value="1210.0">1210 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>