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<document id="7A93B79BDCC3EEC0C37352A95954917A" ID-DOI="10.5281/zenodo.4618877" ID-ISSN="2345-7600" ID-RefBankID="0758FFA7FFE3FFD7F61CFFD21A74FFD5" ID-Zenodo-Dep="4618877" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616092816892" checkinUser="carolina" docAuthor="Ng, Heok Hee" docDate="2004" docId="B80187B9FF979E49FC4611D6185AFAD7" docLanguage="en" docName="RafflesBZool.52.1.209-213.pdf" docOrigin="Raffles Bulletin of Zoology 52 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Oreoglanis macronemus Ng 2004, new species" docType="treatment" docVersion="3" lastPageNumber="211" masterDocId="4438FFC1FF979E4BFFF617231961FFEE" masterDocTitle="Oreoglanis Macronemus, A New Species Of Glyptosternine Catfish (Teleostei: Siluriformes: Sisoridae) From Northern Laos" masterLastPageNumber="213" masterPageNumber="209" pageNumber="209" updateTime="1698890571937" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="85D66BAF3EABD289F21C3ADA79A6BE11">Oreoglanis Macronemus, A New Species Of Glyptosternine Catfish (Teleostei: Siluriformes: Sisoridae) From Northern Laos</mods:title>
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<mods:namePart id="D2A8437B4C2279645F25F3DC5359A0AE">Ng, Heok Hee</mods:namePart>
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<mods:title id="5B13113FC29EB69508624A175B4DB369">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="8E55CD345A4EDF8DD5B2FC444232A900">2004</mods:date>
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<subSubSection id="78B26524FF979E4BFC4611D61C34F8E3" box="[944,1365,1781,1805]" pageId="0" pageNumber="209" type="nomenclature">
<paragraph id="301736AFFF979E4BFC4611D61C34F8E3" blockId="0.[944,1365,1781,1836]" box="[944,1365,1781,1805]" pageId="0" pageNumber="209">
<heading id="6B5F81C3FF979E4BFC4611D61C34F8E3" bold="true" box="[944,1365,1781,1805]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="209" reason="2">
<emphasis id="02DCEABDFF979E4BFC4611D61C34F8E3" bold="true" box="[944,1365,1781,1805]" pageId="0" pageNumber="209">
<emphasis id="02DCEABDFF979E4BFC4611D61DA8F8E3" bold="true" box="[944,1225,1781,1805]" italics="true" pageId="0" pageNumber="209">
<taxonomicName id="F7A84D2CFF979E4BFC4611D61DA2F8E3" ID-CoL="6T4H2" authority="Ng, 2004" authorityName="Ng" authorityYear="2004" box="[944,1219,1781,1805]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="0" pageNumber="209" phylum="Chordata" rank="species" species="macronemus" status="sp. nov.">Oreoglanis macronemus</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="19EF57C6FF979E4BFB2611D61C34F8E3" box="[1232,1365,1781,1805]" pageId="0" pageNumber="209" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="78B26524FF979E4BFBAC10361DCDF8C2" box="[1114,1196,1813,1836]" pageId="0" pageNumber="209" type="description">
<paragraph id="301736AFFF979E4BFBAC10361DCDF8C2" blockId="0.[944,1365,1781,1836]" box="[1114,1196,1813,1836]" pageId="0" pageNumber="209">
(
<figureCitation id="A8932A2AFF979E4BFB9410361DC0F8C2" box="[1122,1185,1813,1836]" captionStart="Fig" captionStartId="2.[136,170,2066,2087]" captionTargetBox="[136,776,1441,2032]" captionTargetId="figure-815@2.[136,776,1441,2032]" captionTargetPageId="2" captionText="Fig. 1. Oreoglanis macronemus, new species, BMNH 1933.8.19.52, paratype male, 61.5 mm SL; Laos: Xieng Khouang." figureDoi="http://doi.org/10.5281/zenodo.4618881" httpUri="https://zenodo.org/record/4618881/files/figure.png" pageId="0" pageNumber="209">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="78B26524FF979E4BFCB510711C5AF834" pageId="0" pageNumber="209" type="materials_examined">
<paragraph id="301736AFFF979E4BFCB510711CA2F86A" blockId="0.[835,1475,1874,1924]" pageId="0" pageNumber="209">
<emphasis id="02DCEABDFF979E4BFCB510711D74F889" bold="true" box="[835,1045,1874,1895]" italics="true" pageId="0" pageNumber="209">Material examined. </emphasis>
<materialsCitation id="80C03CF2FF979E4BFBEC10711CA2F86A" collectionCode="BMNH" collectorName="J. Delacour &amp; W. Lowe" country="Laos" location="Xieng Khouang" pageId="0" pageNumber="209" specimenCode="BMNH 1933.8" specimenCount="1" stateProvince="Xiangkhouang" typeStatus="holotype">
<typeStatus id="EF13880DFF979E4BFBEC10711D12F889" box="[1050,1139,1874,1895]" pageId="0" pageNumber="209" type="holotype">Holotype</typeStatus>
<specimenCode id="600E9ED4FF979E4BFB7F10711C7DF889" box="[1161,1308,1874,1895]" collectionCode="BMNH" pageId="0" pageNumber="209">BMNH 1933.8</specimenCode>
.19.51, male,
<quantity id="F7509B4AFF979E4BFA6E10711A09F86A" metricMagnitude="-2" metricUnit="m" metricValue="6.47" pageId="0" pageNumber="209" unit="mm" value="64.7">64.7 mm</quantity>
SL,
<collectingCountry id="48BF763FFF979E4BFC62104C1AA7F86A" box="[916,966,1903,1924]" name="Laos" pageId="0" pageNumber="209">Laos</collectingCountry>
:
<collectingRegion id="F26CF84DFF979E4BFC39104C1D08F86A" box="[975,1129,1903,1924]" country="Laos" name="Xiangkhouang" pageId="0" pageNumber="209">Xieng Khouang</collectingRegion>
, coll.
<collectorName id="9D5D5379FF979E4BFB56104C1C6DF86A" box="[1184,1292,1903,1924]" pageId="0" pageNumber="209">J. Delacour</collectorName>
&amp;
<collectorName id="9D5D5379FF979E4BFADF104C1CE2F86A" box="[1321,1411,1903,1924]" pageId="0" pageNumber="209">W. Lowe</collectorName>
, 1926.
</materialsCitation>
</paragraph>
<paragraph id="301736AFFF979E4BFCB5108B1C5AF834" blockId="0.[835,1474,1960,2010]" pageId="0" pageNumber="209">
<materialsCitation id="80C03CF2FF979E4BFCB5108B1A03F834" collectionCode="BMNH" pageId="0" pageNumber="209" specimenCount="2" typeStatus="paratype">
<typeStatus id="EF13880DFF979E4BFCB5108B1AC1F853" box="[835,928,1960,1981]" pageId="0" pageNumber="209" type="paratype">Paratypes</typeStatus>
<collectionCode id="56B9AE6AFF979E4BFC43108B1A9CF853" box="[949,1021,1960,1981]" pageId="0" pageNumber="209">BMNH</collectionCode>
1933.8.19.5254,
<specimenCount id="26AEFD26FF979E4BFB5B108B1DBDF853" box="[1197,1244,1960,1981]" pageId="0" pageNumber="209" type="generic">2 ex.</specimenCount>
, females,
<quantity id="F7509B4AFF979E4BFACF108B1CA3F853" box="[1337,1474,1960,1981]" metricMagnitude="-2" metricUnit="m" metricValue="5.36" metricValueMax="5.97" metricValueMin="4.75" pageId="0" pageNumber="209" unit="mm" value="53.6" valueMax="59.7" valueMin="47.5">47.559.7 mm</quantity>
SL
</materialsCitation>
,
<materialsCitation id="80C03CF2FF979E4BFC9B10E61C56F834" box="[877,1335,1989,2010]" collectionCode="BMNH" collectorName="J. Delacour &amp; W. Lowe" country="Laos" location="Xieng Khouang" pageId="0" pageNumber="209" specimenCount="1" stateProvince="Xiangkhouang" typeStatus="paratype">
<specimenCount id="26AEFD26FF979E4BFC9B10E61AC1F834" box="[877,928,1989,2010]" pageId="0" pageNumber="209" type="generic">1 ex.</specimenCount>
, male,
<quantity id="F7509B4AFF979E4BFC1110E61D5EF834" box="[999,1087,1989,2010]" metricMagnitude="-2" metricUnit="m" metricValue="6.18" pageId="0" pageNumber="209" unit="mm" value="61.8">61.8 mm</quantity>
SL, data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78B26524FF979E4AFCB510DC198AF963" lastPageId="1" lastPageNumber="210" pageId="0" pageNumber="209" type="diagnosis">
<paragraph id="301736AFFF979E4BFCB510DC1CA3F7D8" blockId="0.[835,1475,2047,2102]" pageId="0" pageNumber="209">
<emphasis id="02DCEABDFF979E4BFCB510DC1DBBF7F8" box="[835,1242,2047,2071]" italics="true" pageId="0" pageNumber="209">
<emphasis id="02DCEABDFF979E4BFCB510DC1AADF7F9" bold="true" box="[835,972,2047,2071]" italics="true" pageId="0" pageNumber="209">Diagnosis. </emphasis>
<taxonomicName id="F7A84D2CFF979E4BFC2410DC1DBBF7F8" authorityName="Ng" authorityYear="2004" box="[978,1242,2047,2070]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="0" pageNumber="209" phylum="Chordata" rank="species" species="macronemus">Oreoglanis macronemus</taxonomicName>
</emphasis>
shares with members of the
<taxonomicName id="F7A84D2CFF979E4BFC731F3C1D6BF7D8" authorityName=": MNHN" authorityYear="1936" box="[901,1034,2079,2102]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="0" pageNumber="209" phylum="Chordata" rank="species" species="delacouri">
<emphasis id="02DCEABDFF979E4BFC731F3C1D6BF7D8" box="[901,1034,2079,2102]" italics="true" pageId="0" pageNumber="209">O. delacouri</emphasis>
</taxonomicName>
group a lower lip lacking a median notch,
</paragraph>
<paragraph id="301736AFFF969E4AFF8713F5198AF963" blockId="1.[113,753,1238,1677]" pageId="1" pageNumber="210">
with prominent projections along the posterior margin and a lunate caudal fin with extended principal rays. This distinguishes it from congeners of the
<taxonomicName id="F7A84D2CFF969E4AFDE712341BFAFAC0" box="[529,667,1303,1326]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="02DCEABDFF969E4AFDE712341BFAFAC0" box="[529,667,1303,1326]" italics="true" pageId="1" pageNumber="210">O. siamensis</emphasis>
</taxonomicName>
species group (
<taxonomicName id="F7A84D2CFF969E4AFF3E12141826FAA0" box="[200,327,1335,1358]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="insignis">
<emphasis id="02DCEABDFF969E4AFF3E12141826FAA0" box="[200,327,1335,1358]" italics="true" pageId="1" pageNumber="210">O. insignis</emphasis>
</taxonomicName>
,
<taxonomicName id="F7A84D2CFF969E4AFEAC12141B71FAA0" box="[346,528,1335,1358]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="macropterus">
<emphasis id="02DCEABDFF969E4AFEAC12141B71FAA0" box="[346,528,1335,1358]" italics="true" pageId="1" pageNumber="210">O. macropterus</emphasis>
</taxonomicName>
,
<taxonomicName id="F7A84D2CFF969E4AFDD512141BF7FAA0" box="[547,662,1335,1358]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="setiger">
<emphasis id="02DCEABDFF969E4AFDD512141BF7FAA0" box="[547,662,1335,1358]" italics="true" pageId="1" pageNumber="210">O. setiger</emphasis>
</taxonomicName>
and
<taxonomicName id="F7A84D2CFF969E4AFD23121419B8FA80" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="02DCEABDFF969E4AFD23121419B8FA80" italics="true" pageId="1" pageNumber="210">O. siamensis</emphasis>
</taxonomicName>
), which have a median notch on the lower lip, with an entire or weakly laciniate posterior margin, and emarginate caudal fins. It can be distinguished from congeners of the
<taxonomicName id="F7A84D2CFF969E4AFD2012B419B9FA20" authorityName=": MNHN" authorityYear="1936" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="delacouri">
<emphasis id="02DCEABDFF969E4AFD2012B419B9FA20" italics="true" pageId="1" pageNumber="210">O. delacouri</emphasis>
</taxonomicName>
species group in having a shorter caudal peduncle (15.617.3% SL vs. 18.727.1) and longer nasal barbels (35.441.7% HL vs. 11.232.0). It is further distinguished from
<taxonomicName id="F7A84D2CFF969E4AFF5C1134184EF9C0" authorityName=": MNHN" authorityYear="1936" box="[170,303,1559,1582]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="delacouri">
<emphasis id="02DCEABDFF969E4AFF5C1134184EF9C0" box="[170,303,1559,1582]" italics="true" pageId="1" pageNumber="210">O. delacouri</emphasis>
</taxonomicName>
,
<taxonomicName id="F7A84D2CFF969E4AFECD113418D6F9C0" box="[315,439,1559,1582]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="infulatus">
<emphasis id="02DCEABDFF969E4AFECD113418D6F9C0" box="[315,439,1559,1582]" italics="true" pageId="1" pageNumber="210">O. infulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="F7A84D2CFF969E4AFE3511341B28F9C0" box="[451,585,1559,1582]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="hypsiurus">
<emphasis id="02DCEABDFF969E4AFE3511341B28F9C0" box="[451,585,1559,1582]" italics="true" pageId="1" pageNumber="210">O. hypsiurus</emphasis>
</taxonomicName>
and
<taxonomicName id="F7A84D2CFF969E4AFD8C11341B91F9C0" box="[634,752,1559,1582]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="lepturus">
<emphasis id="02DCEABDFF969E4AFD8C11341B91F9C0" box="[634,752,1559,1582]" italics="true" pageId="1" pageNumber="210">O. lepturus</emphasis>
</taxonomicName>
in having a deeper caudal peduncle (4.24.9% SL vs. 2.1 3.6), and from
<taxonomicName id="F7A84D2CFF969E4AFEE7117418EBF980" box="[273,394,1623,1646]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="02DCEABDFF969E4AFEE7117418EBF980" box="[273,394,1623,1646]" italics="true" pageId="1" pageNumber="210">O. frenatus</emphasis>
</taxonomicName>
in having more vertebrae (3940 vs. 3638).
</paragraph>
</subSubSection>
<subSubSection id="78B26524FF969E49FF87119518D6FBD7" lastPageId="2" lastPageNumber="211" pageId="1" pageNumber="210" type="description">
<paragraph id="301736AFFF969E4AFF87119519A0F863" blockId="1.[113,753,1718,1933]" pageId="1" pageNumber="210">
<emphasis id="02DCEABDFF969E4AFF8711951875F920" bold="true" box="[113,276,1718,1742]" italics="true" pageId="1" pageNumber="210">Description. </emphasis>
Morphometric data as in Table 1. Head moderately broad and extremely depressed; dorsal profile slightly convex and ventral profile almost straight. Eye ovoid, horizontal axis longest, subcutaneous; located dorsally on head. Gill openings extremely narrow, extending from middle of pectoral-fin base to level just medial to base of last pectoralfin ray.
</paragraph>
<paragraph id="301736AFFF969E4AFF8710951D5DFA83" blockId="1.[113,753,1974,2093]" lastBlockId="1.[813,1452,1238,1389]" pageId="1" pageNumber="210">Mouth strongly inferior, with broad, thin and papillate lips. Lower lip lacking median notch, posterior margin with lobulate projections. Postlabial groove on lower jaw present and uninterrupted. Jaw teeth pointed and in large, broad band with small median indentation and rounded ends on both sides in upper jaws. Two kinds of teeth present on lower jaw in two separate, roughly triangular patches: anterior teeth short and spatulate, with curved inner face; posterior teeth pointed as in those of upper jaw.</paragraph>
<paragraph id="301736AFFF969E4AFCDB12B51DD7F923" blockId="1.[813,1452,1430,1741]" pageId="1" pageNumber="210">Barbels in four pairs. Maxillary barbels flattened, with membranous flap of skin on dorsoposterior edge and rounded tip. Ventral surface with numerous plicae, with crenulate ventroposterior edge. Nasal barbel extending to point midway between barbel base and anterior orbital margin. Inner mandibular-barbel origin close to midline immediately under lower lip and extending to imaginary line through bases of the first pectoral-fin elements. Outer mandibular barbel originates posterolateral of inner mandibular barbel, extending to base of first pectoral-fin element.</paragraph>
<paragraph id="301736AFFF969E4AFCDB11D51C12F843" blockId="1.[813,1452,1782,1965]" pageId="1" pageNumber="210">Body extremely depressed, becoming compressed towards caudal peduncle. Dorsal profile rising very gently from tip of snout to origin of dorsal fin, then almost horizontal to end of adipose fin base, sloping gently ventrally thereafter to end of caudal peduncle. Ventral profile horizontal to anus and thereafter sloping dorsally to end of caudal peduncle.</paragraph>
<paragraph id="301736AFFF969E4AFCDB10F51D28F7C3" blockId="1.[813,1453,2006,2093]" pageId="1" pageNumber="210">Skin smooth. Lateral line complete and midlateral in position. Vertebrae 24+15=39 (n=1), 25+14=39 (n=1), 26+13=39 (n=1) or 26+14=40 (n=1).</paragraph>
<paragraph id="301736AFFF959E49FF7E17E11806FDB7" blockId="2.[136,776,194,601]" pageId="2" pageNumber="211">Dorsal fin with first element not ossified and i,6 (n=4) rays; fin margin convex. Pectoral fin greatly enlarged, with first element thickened but not ossified, and i,15,i (n=2) or i,17 (n=2) rays. Ventral surface of thickened first pectoral-fin element with numerous striae. Pectoral-fin margin strongly convex. Pelvic-fin origin at midpoint of body. Pelvic fin enlarged, with i,5 (n=4) rays and convex margin. Ventral surface of first pelvic-fin element with numerous striae. Anus and urogenital openings located at vertical through line through posterior quarter of adpressed pelvic fin. Anal fin with short base and ii,4 (n=4) rays. Caudal peduncle short. Caudal fin lunate, with i,5,5,i (n=3), i,6,5,i (n=4) or i,6,6,i (n=1) principal rays.</paragraph>
<paragraph id="301736AFFF959E49FF7E15A1199EFD17" blockId="2.[136,775,642,761]" pageId="2" pageNumber="211">Males with small conical genital papilla immediately posterior to anus in depression terminating posteriorly with straight transverse wall. Females with tube-shaped papilla similarly positioned.</paragraph>
<paragraph id="301736AFFF959E49FF7E140118D6FBD7" blockId="2.[136,776,802,1081]" pageId="2" pageNumber="211">
<emphasis id="02DCEABDFF959E49FF7E14011996FCD4" bold="true" box="[136,247,802,826]" italics="true" pageId="2" pageNumber="211">Colour. </emphasis>
In 70% ethanol: brown on dorsal and lateral surfaces of the head and body, dark yellow on ventral region. A paler, dark yellow ovoid patch on the dorsum at the base of the first dorsal-fin element present. Dorsal and caudal fins brown; dorsal surfaces of pectoral and pelvic fins brown, with light yellow ventral surfaces. Anal fin yellow. Dorsal surface of barbels brown, ventral surface light yellow. Caudal-fin base with a vertical darker band that extends along outermost lower principal caudal rays.
</paragraph>
</subSubSection>
<subSubSection id="78B26524FF959E49FF7E13411BE1FB57" pageId="2" pageNumber="211" type="etymology">
<paragraph id="301736AFFF959E49FF7E13411BE1FB57" blockId="2.[136,775,1122,1209]" pageId="2" pageNumber="211">
<emphasis id="02DCEABDFF959E49FF7E1341187AFB94" bold="true" box="[136,283,1122,1146]" italics="true" pageId="2" pageNumber="211">Etymology. </emphasis>
From the Greek makros, meaning long, and nema, meaning thread. In reference to the long nasal barbels of this species. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="78B26524FF959E49FF7E13C1185AFAD7" pageId="2" pageNumber="211" type="distribution">
<paragraph id="301736AFFF959E49FF7E13C1185AFAD7" blockId="2.[136,775,1250,1337]" pageId="2" pageNumber="211">
<emphasis id="02DCEABDFF959E49FF7E13C11845FB14" bold="true" box="[136,292,1250,1274]" italics="true" pageId="2" pageNumber="211">Distribution. </emphasis>
Known from the
<collectingRegion id="F26CF84DFF959E49FE2C13C11BAAFB17" box="[474,715,1250,1273]" country="Laos" name="Xiangkhouang" pageId="2" pageNumber="211">Xieng Khouang region</collectingRegion>
, most likely from either the Nam Ngiap or the Nam Sen drainage (see discussion).
</paragraph>
</subSubSection>
</treatment>
</document>