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<document id="2A1F09DDF20D96A1E113D818E6C784A8" ID-CLB-Dataset="54553" ID-DOI="10.11646/zootaxa.5221.1.1" ID-GBIF-Dataset="3c98b60d-917e-428e-af26-ae56accd2646" ID-ISSN="1175-5326" ID-Zenodo-Dep="7441371" ID-ZooBank="BECF280B-99E0-4DE3-874B-8585C1E4602E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1671100614055" checkinUser="plazi" docAuthor="Miller, Douglass R. &amp; Stocks, Ian C." docDate="2022" docId="AF31879188578115FF12FF5507C11A45" docLanguage="en" docName="zootaxa.5221.1.1.pdf" docOrigin="Zootaxa 5221 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Acanthococcus Signoret 1875" docType="treatment" docVersion="8" lastPageNumber="12" masterDocId="5308FFE9885C811EFF85FFC2025D1D10" masterDocTitle="New genera and species of felt scales (Hemiptera: Coccomorpha: Eriococcidae), with descriptions of new species and immature instars of described species" masterLastPageNumber="213" masterPageNumber="1" pageNumber="12" updateTime="1699413580947" updateUser="ExternalLinkService">
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<mods:title id="D42C3293E904CF1659E47F3A7FDA3373">New genera and species of felt scales (Hemiptera: Coccomorpha: Eriococcidae), with descriptions of new species and immature instars of described species</mods:title>
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<mods:namePart id="4FC87FD7964C1FC214F0106F514292FB">Stocks, Ian C.</mods:namePart>
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<treatment id="AF31879188578115FF12FF5507C11A45" ID-DOI="http://doi.org/10.5281/zenodo.7464838" ID-GBIF-Taxon="207926884" ID-Zenodo-Dep="7464838" LSID="urn:lsid:plazi:treatment:AF31879188578115FF12FF5507C11A45" httpUri="http://treatment.plazi.org/id/AF31879188578115FF12FF5507C11A45" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection id="6F82650C88578115FF12FF55005F1DA1" box="[151,514,151,178]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="2727368788578115FF12FF55005F1DA1" blockId="11.[151,514,151,178]" box="[151,514,151,178]" pageId="11" pageNumber="12">
<heading id="7C6F81EB88578115FF12FF55005F1DA1" bold="true" box="[151,514,151,178]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="4">
<taxonomicName id="E0984D0488578115FF12FF55005F1DA1" authority="Signoret 1875" authorityName="Signoret" authorityYear="1875" box="[151,514,151,178]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FF12FF55005F1DA1" bold="true" box="[151,514,151,178]" pageId="11" pageNumber="12">
<emphasis id="15ECEA9588578115FF12FF5503121DA2" bold="true" box="[151,335,151,178]" italics="true" pageId="11" pageNumber="12">Acanthococcus</emphasis>
<bibRefCitation id="43094B7688578115FED2FF55005F1DA1" author="Signoret, V." box="[343,514,151,177]" pageId="11" pageNumber="12" pagination="15 - 40" refId="ref122766" refString="Signoret, V. (1875) Essai sur les cochenilles ou gallinsectes (Homopteres - Coccides), 14 e partie. Annales de la Societe entomologique de France, Serie 5, 5, 15 - 40." type="journal article" year="1875">Signoret 1875</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="2727368788578115FF12FF1E06D01DE4" blockId="11.[151,1436,219,305]" box="[151,1165,219,244]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="078811A988578115FF12FF1E06D01DE4" box="[151,1165,219,244]" pageId="11" pageNumber="12">
<subSubSection id="6F82650C88578115FF12FF1E03AE1DE3" box="[151,499,219,244]" pageId="11" pageNumber="12" type="reference_group">
<treatmentCitation id="A639109688578115FF12FF1E03B21DE3" author="Signoret, V." box="[151,495,219,244]" page="16" pageId="11" pageNumber="12" year="1875">
<taxonomicName id="E0984D0488578115FF12FF1E03B21DE3" authority="Signoret 1875: 16" authorityName="Signoret" authorityPageNumber="16" authorityYear="1875" box="[151,495,219,244]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FF12FF1E036D1DE4" box="[151,304,220,244]" italics="true" pageId="11" pageNumber="12">Acanthococcus</emphasis>
<bibRefCitation id="43094B7688578115FEB2FF1E03B21DE3" author="Signoret, V." box="[311,495,219,244]" pageId="11" pageNumber="12" pagination="15 - 40" refId="ref122766" refString="Signoret, V. (1875) Essai sur les cochenilles ou gallinsectes (Homopteres - Coccides), 14 e partie. Annales de la Societe entomologique de France, Serie 5, 5, 15 - 40." type="journal article" year="1875">Signoret 1875: 16</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</subSubSection>
<subSubSection id="6F82650C88578115FE7CFF1E06D01DE4" box="[505,1165,219,244]" pageId="11" pageNumber="12" type="type_taxon">
<typeStatus id="F823882588578115FE7CFF1E00761DE4" box="[505,555,220,244]" pageId="11" pageNumber="12">Type</typeStatus>
species:
<taxonomicName id="E0984D0488578115FD0DFF1E01A61DE3" authority="Signoret 1875" authorityName="Signoret" authorityYear="1875" box="[648,1019,219,244]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aceris">
<emphasis id="15ECEA9588578115FD0DFF1E01381DE3" box="[648,869,220,244]" italics="true" pageId="11" pageNumber="12">Acanthococcus aceris</emphasis>
<bibRefCitation id="43094B7688578115FCE9FF1E01A61DE3" author="Signoret, V." box="[876,1019,219,244]" pageId="11" pageNumber="12" pagination="15 - 40" refId="ref122766" refString="Signoret, V. (1875) Essai sur les cochenilles ou gallinsectes (Homopteres - Coccides), 14 e partie. Annales de la Societe entomologique de France, Serie 5, 5, 15 - 40." type="journal article" year="1875">Signoret 1875</bibRefCitation>
</taxonomicName>
by monotypy.
</subSubSection>
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</paragraph>
<subSubSection id="6F82650C88578115FF12FF3803691C21" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="2727368788578115FF12FF3803691C21" blockId="11.[151,1436,219,305]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="078811A988578115FF12FF3803691C21" pageId="11" pageNumber="12">
<treatmentCitation id="A639109688578115FF12FF3800321C02" author="Cockerell, T. D. A. &amp; Rohwer, S. A." box="[151,623,250,274]" page="169" pageId="11" pageNumber="12" year="1909">
<taxonomicName id="E0984D0488578115FF12FF3800321C02" ID-CoL="35CZ" authority="Cockerell and Rohwer 1909: 169" authorityName="Cockerell and Rohwer" authorityPageNumber="169" authorityYear="1909" box="[151,623,250,274]" class="Insecta" family="Eriococcidae" genus="Atriplicia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FF12FF3802A51C02" box="[151,248,250,274]" italics="true" pageId="11" pageNumber="12">Atriplicia</emphasis>
<bibRefCitation id="43094B7688578115FE80FF3800321C02" author="Cockerell, T. D. A. &amp; Rohwer, S. A." box="[261,623,250,274]" pageId="11" pageNumber="12" pagination="169 - 170" refId="ref119814" refString="Cockerell, T. D. A. &amp; Rohwer, S. A. (1909) A new gall-making coccid on Atriplex. Proceedings of the Entomological Society of Washington, 10, 169 - 170." type="journal article" year="1909">Cockerell and Rohwer 1909: 169</bibRefCitation>
</taxonomicName>
</treatmentCitation>
<emphasis id="15ECEA9588578115FDF9FF3800951C02" bold="true" box="[636,712,250,274]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="0EDF57EE88578115FDF9FF3800951C02" box="[636,712,250,274]" pageId="11" pageNumber="12">syn. n.</taxonomicNameLabel>
</emphasis>
<typeStatus id="F823882588578115FD50FF39015A1C03" box="[725,775,251,275]" pageId="11" pageNumber="12">Type</typeStatus>
species:
<taxonomicName id="E0984D0488578115FCF4FF38072B1C02" ID-CoL="JLT7" authority="Cockerell and Rohwer 1909" authorityName="Cockerell and Rohwer" authorityYear="1909" box="[881,1398,250,274]" class="Insecta" family="Eriococcidae" genus="Atriplicia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gallicola">
<emphasis id="15ECEA9588578115FCF4FF3806641C02" box="[881,1081,250,274]" italics="true" pageId="11" pageNumber="12">Atriplicia gallicola</emphasis>
<bibRefCitation id="43094B7688578115FBC3FF38072B1C02" author="Cockerell, T. D. A. &amp; Rohwer, S. A." box="[1094,1398,250,274]" pageId="11" pageNumber="12" pagination="169 - 170" refId="ref119814" refString="Cockerell, T. D. A. &amp; Rohwer, S. A. (1909) A new gall-making coccid on Atriplex. Proceedings of the Entomological Society of Washington, 10, 169 - 170." type="journal article" year="1909">Cockerell and Rohwer 1909</bibRefCitation>
</taxonomicName>
by monotypy.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6F82650C88578115FF12FE9D07371F66" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="2727368788578115FF12FE9D07371F66" blockId="11.[151,1437,351,1206]" pageId="11" pageNumber="12">
<emphasis id="15ECEA9588578115FF12FE9D00621C69" bold="true" box="[151,575,351,377]" pageId="11" pageNumber="12">Generic diagnosis of adult female:</emphasis>
Enlarged setae usually abundant on dorsum, but some species with setae restricted to all or part of body margin; anal lobes generally large and protruding, each with 1 or more enlarged setae, usually 3; macrotubular ducts present on adult female and second-instar male; microtubular ducts present, usually associated with enlarged setae; multilocular pores present at least on venter, usually each with 5 loculi; cruciform pores usually present; translucent pores present on hind legs of adult female and second-instar female, usually on femur and coxa; anal ring usually with 4 pairs of setae and 1 or 2 rows of pores; each femur usually with 5 setae; each tibia usually with 4‒6 setae; claw with denticle; digitules on tarsus and claw usually capitate; antennae 6- or 7-segmented, with sensory setae on distal 3 segments; labium 3-segmented, basal segment small, with 2 setae.
</paragraph>
</subSubSection>
<subSubSection id="6F82650C88578115FF42FDBD01921ECD" pageId="11" pageNumber="12" type="discussion">
<paragraph id="2727368788578115FF42FDBD00F51FAD" blockId="11.[151,1437,351,1206]" pageId="11" pageNumber="12">
<emphasis id="15ECEA9588578115FF42FDBD034E1F89" bold="true" box="[199,275,639,665]" pageId="11" pageNumber="12">Notes:</emphasis>
For an explanation and a list of the 20 Nearctic species transferred from
<taxonomicName id="E0984D0488578115FBD5FD4206931F89" authorityName="Targioni Tozzetti" authorityYear="1868" box="[1104,1230,640,665]" class="Insecta" family="Eriococcidae" genus="Eriococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FBD5FD4206931F89" box="[1104,1230,640,665]" italics="true" pageId="11" pageNumber="12">Eriococcus</emphasis>
</taxonomicName>
to
<taxonomicName id="E0984D0488578115FB76FDBD07C01F89" authorityName="Signoret" authorityYear="1875" box="[1267,1437,639,665]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FB76FDBD07C01F89" box="[1267,1437,639,665]" italics="true" pageId="11" pageNumber="12">Acanthococcus</emphasis>
</taxonomicName>
in this work, please see the Introduction above.
</paragraph>
<paragraph id="2727368788578115FF42FD0A01921ECD" blockId="11.[151,1437,351,1206]" pageId="11" pageNumber="12">
In North America, adult females of the species in
<taxonomicName id="E0984D0488578115FD6BFD0501C51FF1" authorityName="Signoret" authorityYear="1875" box="[750,920,711,737]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FD6BFD0501C51FF1" box="[750,920,711,737]" italics="true" pageId="11" pageNumber="12">Acanthococcus</emphasis>
</taxonomicName>
are most similar to adult females of species of
<taxonomicName id="E0984D0488578115FF12FD2E037C1E16" authorityName="Kloet" authorityYear="1944" box="[151,289,748,774]" class="Insecta" family="Eriococcidae" genus="Ovaticoccus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FF12FD2E037C1E16" box="[151,289,748,774]" italics="true" pageId="11" pageNumber="12">Ovaticoccus</emphasis>
</taxonomicName>
but differ as follows (character states of
<taxonomicName id="E0984D0488578115FD6EFD2901C81E15" authorityName="Signoret" authorityYear="1875" box="[747,917,747,773]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FD6EFD2901C81E15" box="[747,917,747,773]" italics="true" pageId="11" pageNumber="12">Acanthococcus</emphasis>
</taxonomicName>
are in brackets): anal ring normally with three or fewer pairs of setae (with four pairs of setae); anal lobes not protruding or protruding slightly, inconspicuous (protruding strongly, conspicuous); without frontal lobes (with or without frontal lobes); translucent pores usually on hind coxa only (on hind femur and coxa). Species of
<taxonomicName id="E0984D0488578115FCCCFC9501AE1E61" authorityName="Signoret" authorityYear="1875" box="[841,1011,855,881]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FCCCFC9501AE1E61" box="[841,1011,855,881]" italics="true" pageId="11" pageNumber="12">Acanthococcus</emphasis>
</taxonomicName>
also are similar to the Australasian and Neotropical genus
<taxonomicName id="E0984D0488578115FE22FCB900961E85" authority="Hoy 1962" authorityName="Hoy" authorityYear="1962" box="[423,715,891,918]" class="Insecta" family="Eriococcidae" genus="Madarococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FE22FCB900111E85" box="[423,588,891,917]" italics="true" pageId="11" pageNumber="12">Madarococcus</emphasis>
<bibRefCitation id="43094B7688578115FDDDFCBE00961E85" author="Hoy, J. M." box="[600,715,891,918]" pageId="11" pageNumber="12" pagination="1 - 219" refId="ref121371" refString="Hoy, J. M. (1962) Eriococcidae (Homoptera: Coccoidea) of New Zealand. New Zealand Department of Scientific and Industrial Research Bulletin, 146, 1 - 219." type="journal article" year="1962">Hoy 1962</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="43094B7688578115FD5BFCB901EE1E85" author="Hardy, N. B. &amp; Gullan, P. J. &amp; Henderson, R. C. &amp; Cook, L. G." box="[734,947,891,917]" pageId="11" pageNumber="12" pagination="365 - 405" refId="ref120977" refString="Hardy, N. B., Gullan, P. J., Henderson, R. C. &amp; Cook, L. G. (2008) Relationships among felt scale insects (Hemiptera: Coccoidea: Eriococcidae) of southern beech, Nothofagus (Nothofagaceae), with the first descriptions of Australian species of the Nothofagus - feeding genus Madarococcus Hoy. Invertebrate Systematics, 22 (3), 365 - 405. https: // doi. org / 10.1071 / IS 07032" type="journal article" year="2008">
Hardy
<emphasis id="15ECEA9588578115FCB4FCBE01331E85" box="[817,878,891,917]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2008
</bibRefCitation>
) but differ as follows (character states of
<taxonomicName id="E0984D0488578115FF12FC5D031C1EA9" authorityName="Signoret" authorityYear="1875" box="[151,321,927,953]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FF12FC5D031C1EA9" box="[151,321,927,953]" italics="true" pageId="11" pageNumber="12">Acanthococcus</emphasis>
</taxonomicName>
in brackets): suranal seta expanded (suranal seta flagellate); anal lobes large and almost plate like, heavily sclerotized (anal lobes not plate like, usually slightly sclerotized).
</paragraph>
</subSubSection>
<subSubSection id="6F82650C88578115FF42FC250198195A" pageId="11" pageNumber="12" type="etymology">
<paragraph id="2727368788578115FF42FC250198195A" blockId="11.[151,1437,351,1206]" pageId="11" pageNumber="12">
<emphasis id="15ECEA9588578115FF42FC25030F1911" bold="true" box="[199,338,999,1025]" pageId="11" pageNumber="12">Etymology:</emphasis>
The generic epithet “
<taxonomicName id="E0984D0488578115FDBBFC2500BA1911" authorityName="Signoret" authorityYear="1875" box="[574,743,999,1025]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="15ECEA9588578115FDBBFC2500BA1911" box="[574,743,999,1025]" italics="true" pageId="11" pageNumber="12">Acanthococcus</emphasis>
</taxonomicName>
” is a combination of the Greek word “
<emphasis id="15ECEA9588578115FB0AFC2506B41911" box="[1167,1257,999,1025]" italics="true" pageId="11" pageNumber="12">akantha</emphasis>
” meaning spine or thorn, and “
<emphasis id="15ECEA9588578115FEB9FBC903D51935" box="[316,392,1035,1061]" italics="true" pageId="11" pageNumber="12">kokkos</emphasis>
” meaning “seed” or “scale insect” and is a masculine noun. The generic epithet is formed to point out the enlarged setae on the dorsum of many species of the genus.
</paragraph>
</subSubSection>
<subSubSection id="6F82650C88578115FF42FB91060E19A5" pageId="11" pageNumber="12" type="biology_ecology">
<paragraph id="2727368788578115FF42FB91060E19A5" blockId="11.[151,1437,351,1206]" pageId="11" pageNumber="12">
<emphasis id="15ECEA9588578115FF42FB91032F197D" bold="true" box="[199,370,1107,1133]" pageId="11" pageNumber="12">Field features:</emphasis>
Body usually pear-shaped, with various arrangements and shapes of crystalline wax rods formed on the enlarged setae; forming white felted ovisac that covers body, with small posterior exit hole. Occurring on most parts of host including roots, crown, branches, and leaves, rarely inducing galls.
</paragraph>
</subSubSection>
<subSubSection id="6F82650C88578115FF12FAC507C11A45" pageId="11" pageNumber="12" type="key">
<paragraph id="2727368788578115FF12FAC506471831" blockId="11.[151,1050,1287,1314]" box="[151,1050,1287,1314]" pageId="11" pageNumber="12">
<emphasis id="15ECEA9588578115FF12FAC506471831" bold="true" box="[151,1050,1287,1314]" pageId="11" pageNumber="12">
Key to
<taxonomicName id="E0984D0488578115FF74FAC5005B1831" box="[241,518,1287,1314]" pageId="11" pageNumber="12">
<emphasis id="15ECEA9588578115FF74FAC503F41832" bold="true" box="[241,425,1287,1314]" italics="true" pageId="11" pageNumber="12">Acanthococcus</emphasis>
species
</taxonomicName>
treated here, based on first-instar nymphs
</emphasis>
</paragraph>
<key id="530961B688578115FF12FA8E07C11A45" pageId="11" pageNumber="12">
<keyStep id="9C6C2E2288578115FF12FA8E07C1186F" pageId="11" pageNumber="12">
<paragraph id="2727368788578115FF12FA8E07C11871" blockId="11.[151,1437,1356,1877]" box="[151,1436,1356,1378]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12FA8E07C11871" box="[151,1436,1356,1378]" pageId="11" pageNumber="12">1(0) Each hind tibia with 5 setae, including middle seta.......................................................... 4</keyLead>
</paragraph>
<paragraph id="2727368788578115FF12FAA807C1186F" blockId="11.[151,1437,1356,1877]" box="[151,1436,1385,1407]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12FAA807C1186F" box="[151,1436,1385,1407]" pageId="11" pageNumber="12">– Each hind tibia with 4 setae, without middle seta............................................................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6C2E2288578115FF12FA4507C118E4" pageId="11" pageNumber="12">
<paragraph id="2727368788578115FF12FA4507C118A9" blockId="11.[151,1437,1356,1877]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12FA4507C118A9" pageId="11" pageNumber="12">2(1) Each front tibia with 4 setae, without middle seta; medial enlarged setae on dorsum of abdomen as large as largest lateral setae............................................................................................... 3</keyLead>
</paragraph>
<paragraph id="2727368788578115FF12FA0007C118E4" blockId="11.[151,1437,1356,1877]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12FA0007C118E4" pageId="11" pageNumber="12">
– Each front tibia with 5 setae, with middle seta; medial enlarged setae on dorsum of abdomen smaller than largest lateral setae.....................................................................................
<taxonomicName id="E0984D0488578115FB68FA1D07C118E4" ID-CoL="8V8R" authority="(Parrott)" baseAuthorityName="Parrott" box="[1261,1436,1503,1524]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kemptoni">
<emphasis id="15ECEA9588578115FB68FA1D071818E4" box="[1261,1349,1503,1524]" italics="true" pageId="11" pageNumber="12">kemptoni</emphasis>
(Parrott)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6C2E2288578115FF12FA3E07C11B79" pageId="11" pageNumber="12">
<paragraph id="2727368788578115FF12FA3E07C11B3F" blockId="11.[151,1437,1356,1877]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12FA3E07C11B3F" pageId="11" pageNumber="12">
3(2) Multilocular pores near spiracles each often with 7 or 9 loculi; multilocular pores on abdomen predominantly each with 5 loculi; cruciform pores present on ventral margin of thorax.................................
<taxonomicName id="E0984D0488578115FB2BF9DB07C11B3F" ID-CoL="8V6H" authority="Miller and Miller" authorityName="Miller and Miller" box="[1198,1436,1561,1583]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="dennoi">
<emphasis id="15ECEA9588578115FB2BF9DB06B21B3E" box="[1198,1263,1561,1582]" italics="true" pageId="11" pageNumber="12">dennoi</emphasis>
Miller and Miller
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2727368788578115FF12F9F507C11B79" blockId="11.[151,1437,1356,1877]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12F9F507C11B79" pageId="11" pageNumber="12">
– Multilocular pores near spiracles each with 5 loculi; multilocular pores on abdomen predominantly each with 3 loculi; cruciform pores absent from ventral margin of thorax.......................................
<taxonomicName id="E0984D0488578115FBE5F997073D1B7A" authority="Miller and Stocks" authorityName="Miller and Stocks" authorityYear="2022" box="[1120,1376,1620,1642]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pennyae" status="sp. nov.">
<emphasis id="15ECEA9588578115FBE5F99706ED1B7A" box="[1120,1200,1621,1642]" italics="true" pageId="11" pageNumber="12">pennyae</emphasis>
Miller and Stocks
</taxonomicName>
<emphasis id="15ECEA9588578115FAE3F99607C11B79" bold="true" box="[1382,1436,1620,1641]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="0EDF57EE88578115FAE3F99607C11B79" box="[1382,1436,1620,1641]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6C2E2288578115FF12F9B007C11BB5" pageId="11" pageNumber="12">
<paragraph id="2727368788578115FF12F9B007C11B97" blockId="11.[151,1437,1356,1877]" box="[151,1436,1649,1671]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12F9B007C11B97" box="[151,1436,1649,1671]" pageId="11" pageNumber="12">4(1) Hind femora each with 5 setae.......................................................................... 5</keyLead>
</paragraph>
<paragraph id="2727368788578115FF12F94D07C11BB5" blockId="11.[151,1437,1356,1877]" box="[151,1436,1679,1701]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12F94D07C11BB5" box="[151,1436,1679,1701]" pageId="11" pageNumber="12">
– Hind femora each with 4 setae.........................................................
<taxonomicName id="E0984D0488578115FB4FF94D07C11BB5" authority="(Cockerell)" baseAuthorityName="Cockerell" baseAuthorityYear="1894" box="[1226,1436,1679,1701]" class="Insecta" family="Eriococcidae" genus="Rhizococcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="coccineus">
<emphasis id="15ECEA9588578115FB4FF94D07741BB4" box="[1226,1321,1679,1700]" italics="true" pageId="11" pageNumber="12">coccineus</emphasis>
(Cockerell)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6C2E2288578115FF12F96E07C11BCF" pageId="11" pageNumber="12">
<paragraph id="2727368788578115FF12F96E07C11BD2" blockId="11.[151,1437,1356,1877]" box="[151,1436,1708,1730]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12F96E07C11BD2" box="[151,1436,1708,1730]" pageId="11" pageNumber="12">5(4) Margin of each abdominal segment with 1 enlarged seta...................................................... 6</keyLead>
</paragraph>
<paragraph id="2727368788578115FF12F90807C11BCF" blockId="11.[151,1437,1356,1877]" box="[151,1436,1737,1759]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12F90807C11BCF" box="[151,1436,1737,1759]" pageId="11" pageNumber="12">
– Margin of each abdominal segment with 2 enlarged setae....................................
<taxonomicName id="E0984D0488578115FB43F90B07C11BCF" ID-CoL="64DQH" authority="Miller and Miller" authorityName="Miller and Miller" box="[1222,1436,1737,1759]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="hoyi">
<emphasis id="15ECEA9588578115FB43F90B06B21BCE" box="[1222,1263,1737,1758]" italics="true" pageId="11" pageNumber="12">hoyi</emphasis>
Miller and Miller
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6C2E2288578115FF12F92507C11A45" pageId="11" pageNumber="12">
<paragraph id="2727368788578115FF12F92507C11A0A" blockId="11.[151,1437,1356,1877]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12F92507C11A0A" pageId="11" pageNumber="12">
6(5) Medial and mediolateral enlarged setae on dorsum of abdomen about same size as lateral setae; cruciform pores absent..............................................................................
<taxonomicName id="E0984D0488578115FBE5F8C607C11A0A" authority="(Cockerell &amp; Rohwer)" authorityName="Miller &amp; Stocks" authorityYear="2022" baseAuthorityName="Cockerell and Rohwer" baseAuthorityYear="2022" box="[1120,1436,1796,1818]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gallicolus">
<emphasis id="15ECEA9588578115FBE5F8C606E21A09" box="[1120,1215,1796,1817]" italics="true" pageId="11" pageNumber="12">gallicolus</emphasis>
(Cockerell &amp; Rohwer)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="2727368788578115FF12F8E007C11A45" blockId="11.[151,1437,1356,1877]" pageId="11" pageNumber="12">
<keyLead id="9C6995B288578115FF12F8E007C11A45" pageId="11" pageNumber="12">
– Medial and mediolateral enlarged setae on dorsum of abdomen smaller than lateral setae; cruciform pores usually present near ventral margin of thorax...............................................................
<taxonomicName id="E0984D0488578115FB56F8FD07C11A45" ID-CoL="8V4B" authority="(Cockerell)" authorityName="Miller &amp; Stocks" authorityYear="2022" baseAuthorityName="Miller &amp; Stocks" baseAuthorityYear="2022" box="[1235,1436,1855,1877]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="arenosus">
<emphasis id="15ECEA9588578115FB56F8FD07741A44" box="[1235,1321,1855,1876]" italics="true" pageId="11" pageNumber="12">arenosus</emphasis>
(Cockerell)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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