145 lines
18 KiB
XML
145 lines
18 KiB
XML
<document id="D04692C3E551677817603FD10645BD5B" ID-CLB-Dataset="35149" ID-DOI="http://dx.doi.org/10.3897/zookeys.536.5982" ID-GBIF-Dataset="f3546db1-d96a-40d8-ac8c-fe806547f201" ID-PMC="PMC4714036" ID-Pensoft-Pub="1313-2970-536-1" ID-PubMed="26798233" ID-ZooBank="A8B92CDD0B8C4384AAC559648BB45AA5" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-536-1" ModsDocOrigin="ZooKeys 536" ModsDocTitle="The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)" checkinTime="1451243741663" checkinUser="pensoft" docAuthor="Shi, Hongliang & Liang, Hongbin" docDate="2015" docId="AE1F585F11B2209DA93DDB633D427AC5" docLanguage="en" docName="ZooKeys 536: 1-92" docOrigin="ZooKeys 536" docSource="http://dx.doi.org/10.3897/zookeys.536.5982" docTitle="Pterostichus (Circinatus) zoiai Sciaky 1996" docType="treatment" docVersion="6" lastPageNumber="44" masterDocId="FFEAFF976442467ED413FFC17531FF81" masterDocTitle="The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="42" updateTime="1701379197761" updateUser="ExternalLinkService">
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<mods:title id="6FD19B771C516125A30EBEE18ED518E5">The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)</mods:title>
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<mods:namePart id="1732EF873687172AEADF7B9FF1121EA1">Shi, Hongliang</mods:namePart>
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<mods:namePart id="A8C45F9172F99D5B6E90C77963158097">Liang, Hongbin</mods:namePart>
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<mods:date id="D5EA1D3203172AC3F79A4BFB564A79D3">2015</mods:date>
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<mods:number id="658984FF0F0757104FBD8C59FFD4EBB6">536</mods:number>
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<mods:classification id="5FC08298AB3B83D4B966457D08E6A904">journal article</mods:classification>
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<mods:identifier id="7654AA54DCB9B29A0FB37F59CB445CEA" type="DOI">http://dx.doi.org/10.3897/zookeys.536.5982</mods:identifier>
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<treatment id="AE1F585F11B2209DA93DDB633D427AC5" ID-GBIF-Taxon="152063168" LSID="urn:lsid:plazi:treatment:AE1F585F11B2209DA93DDB633D427AC5" httpUri="http://treatment.plazi.org/id/AE1F585F11B2209DA93DDB633D427AC5" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
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<subSubSection id="7C9852015D5F6CAA3F6F7EB0F5228036" pageId="41" pageNumber="42" type="multiple">
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<paragraph id="98DD10D145AE360A1F6C9A907E0A1B52" pageId="41" pageNumber="42">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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</subSubSection>
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<subSubSection id="E760BF10DE3291277380BABE33B96AEF" pageId="41" pageNumber="42" type="nomenclature">
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<paragraph id="2B6808DDB9B7FE9011AA72AB30EC1E51" pageId="41" pageNumber="42">
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<taxonomicName id="251AAE38EF276F88B84C9C2485FA380E" ID-CoL="6WM49" LSID="Chinese common name: 佐亚通缘步甲 (Zuŏ Yà Tōng Yuán Bù Jiă)" authority="Sciaky, 1996" authorityName="Sciaky" authorityYear="1996" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Circinatus) zoiai" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="zoiai" subGenus="Circinatus">Pterostichus (Circinatus) zoiai Sciaky, 1996</taxonomicName>
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Figures 25, 49, 69, 88, 110
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</paragraph>
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</subSubSection>
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<subSubSection id="27D38C63315418C865D00DDA3F1FBFC1" pageId="42" pageNumber="43" type="reference_group">
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<paragraph id="94BDAA7E4E73FA9EF0BD6B329124F2AE" pageId="42" pageNumber="43">
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<taxonomicName id="856A48F9655AB374947A81678900592A" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Circinatus) zoiai" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="zoiai" subGenus="Circinatus">
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<pageBreakToken id="4D156A51399D963A204D5A4EB0FF7AF9" pageId="42" pageNumber="43" start="start">Pterostichus</pageBreakToken>
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(Circinatus) zoiai
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</taxonomicName>
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<bibRefCitation id="31A7852525194C5150432EBBA035FAB8" author="Sciaky, R" journalOrPublisher="Annuario del Museo Civico di Storia Naturale di Brescia" pageId="60" pageNumber="61" pagination="217 - 231" title="Circinatus new subgenus and three new species of Pterostichus from China (ColeopteraCarabidae). Natura Bresciana." volume="30" year="1996">Sciaky 1996</bibRefCitation>
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: 222 (Original:
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<taxonomicName id="CDFC864A5BDCCC1B0AB7A6DC609805F6" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Pterostichus</taxonomicName>
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(
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<taxonomicName id="B69EB924EDA8B350F4B9E2057009E041" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Circinatus)" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subGenus" subGenus="Circinatus">Circinatus</taxonomicName>
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); holotype deposited in NHMB);
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<bibRefCitation id="B7B92CE9DC25FDC580C9BFBCE897029F" author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="Apollo Books, Denmark" pageId="59" pageNumber="60" pagination="469 - 521" title="Carabidae: tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 1, Archostemata- Myxophaga- Adephaga" year="2003">Bousquet 2003</bibRefCitation>
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: 488;
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<bibRefCitation id="9A5D7B6ADE061A56A324CC0A33F992DB" author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing, Germany" pageId="59" pageNumber="60" title="Systematic list of extant ground beetles of the world (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="2005">Lorenz 2005</bibRefCitation>
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: 280;
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<bibRefCitation id="B08A8F84E229290420CA529C8BA1D2A9" author="Allegro, G" journalOrPublisher="Bollettino del Museo Civico di Storia Naturale di Verona (Botanica Zoologia)" pageId="59" pageNumber="60" pagination="3 - 18" title="Review of the Chinese subgenus Circinatus of Pterostichus, with description of six new species (Coleoptera, Carabidae)." volume="34" year="2010">Allegro and Sciaky 2010</bibRefCitation>
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: 4.
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</paragraph>
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</subSubSection>
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<subSubSection id="FCFFD649D9C651C90DE006234CB12D25" pageId="42" pageNumber="43" type="type locality">
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<paragraph id="95430A07498B001F15AA8645DACAD0CE" pageId="42" pageNumber="43">Type locality.</paragraph>
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<paragraph id="B0616F036988407569FF0C96B250F7EE" pageId="42" pageNumber="43">
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Sichuan: Emei Mt. (~
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<geoCoordinate id="C1269D33DD1D77FFA1F51CD16AA911A7" direction="north" orientation="latitude" precision="555" value="29.55">N29.55°</geoCoordinate>
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,
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<geoCoordinate id="16F9127EBAA5DCCE4A53828AE182606B" direction="east" orientation="longitude" precision="555" value="103.34">E103.34°</geoCoordinate>
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), 600-1050 m.
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</paragraph>
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</subSubSection>
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<subSubSection id="F715B7BDD9932AE6BA910B3EC0E158DD" pageId="42" pageNumber="43" type="type material">
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<paragraph id="A89C23F00B5AA8DEB5311430B565A595" pageId="42" pageNumber="43">Type material examined.</paragraph>
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<paragraph id="9A392F885756F0067D6DAFDBA300865E" pageId="42" pageNumber="43">
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Holotype of
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<taxonomicName id="98E8B06D64C4903416C4313BB2F7EE3C" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zoiai" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="zoiai">Pterostichus zoiai</taxonomicName>
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Sciaky (NHMB), male, body length = 10.7 mm, board mounted, genitalia dissected and glued on same board with specimen, "CHINA: Sichuan / Mt. EMEI, 600-1050 m / 5.-19.5.1989 / Lad.
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<normalizedToken id="B8FF4B89598E026AFEE35BE8A458C4B3" originalValue="Bocák">Bocak</normalizedToken>
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, lgt."; "HOLOTYPUS /
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<taxonomicName id="CAB1B53467277BB28264DF1D581A05F1" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Pterostichus</taxonomicName>
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/ (
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<taxonomicName id="00080D6726B1E031645765B213113439" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Circinatus)" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subGenus" subGenus="Circinatus">Circinatus</taxonomicName>
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)/
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<taxonomicName id="0E6A3A0730C0433DA3577D5239E159F1" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zoiai" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="zoiai">zoiai</taxonomicName>
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sp. n. / Det. Sciaky
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<normalizedToken id="065686FBF7016C22E3AEC1542F38A353" originalValue="‘94”">'94"</normalizedToken>
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[red label]. Paratype, 1 male, "CHINA, Sichuan, / Mt. Emei, 2640 m, 6.5.1989, Lad. Bocak, lgt."; "ex. coll. R. Sciaky, 2011".
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</paragraph>
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</subSubSection>
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<subSubSection id="5A7938712ADA1BB962A34DC8E2E87440" pageId="42" pageNumber="43" type="type material">
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<paragraph id="252306EC95F60EC8F6929D689DACC47B" pageId="42" pageNumber="43">Non-type material examined</paragraph>
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<paragraph id="551D450D413032C741795A624C0DA86E" pageId="42" pageNumber="43">
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(total 44 specimens). 9 males, 6 females (IZAS), "China, Sichuan, mount. Emei, Leidongping, broadleaf forest with dead log under forest,
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<geoCoordinate id="B37665E8D02AE1CC758845F6B77B6CB1" direction="north" orientation="latitude" precision="1" value="29.54669">N29.54669</geoCoordinate>
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,
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<geoCoordinate id="5E4A8BA12525B08A2A3BFCAFC17BC134" direction="east" orientation="longitude" precision="1" value="103.33552">E103.33552</geoCoordinate>
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; 2420 m, 2012.VII.02 N, on ground or tree trunk, SHI Hongliang, LIU Ye, leg."; 2 males, 2 females (IZAS), same data, but collecting date 2012.VII.03; 2 males, 7 females (IZAS), same data, but collecting date 2012.VII.05; 7 males, 8 females (IZAS), "China, Sichuan, mount. Emei, Leidongping, broadleaf forest with dead log under forest,
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<geoCoordinate id="C215A34B58CD5F0FDCE00539C6376751" direction="north" orientation="latitude" precision="1" value="29.54669">N29.54669</geoCoordinate>
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,
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<geoCoordinate id="D12C213B691872D867392290098C76F7" direction="east" orientation="longitude" precision="1" value="103.33552">E103.33552</geoCoordinate>
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; 2420 m, 2012.VII.06, by pitfall trap, SHI Hongliang, LIU Ye, leg."; 1 female (IZAS), "Mt. Emei, Sichuan Prov. 17-VII-2003, Hu & Tang leg.".
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</paragraph>
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</subSubSection>
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<subSubSection id="C401BE5AE6CAEFEF7088836C075F9C2F" pageId="42" pageNumber="43" type="diagnosis">
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<paragraph id="BADAF59B78128210E5FBA663F309C17C" pageId="42" pageNumber="43">Diagnosis.</paragraph>
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<paragraph id="5D190DC47118ADC04027E816DFC09666" pageId="42" pageNumber="43">
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Pronotum with single mid-lateral seta; posterior seta distant from hind angle, hind angle completely rounded; basal foveal outer groove absent, outer region of inner groove completely flat; basal fovea area finely punctate between two inner grooves; elytron with humeral tooth almost invisible; elytral microsculpture faint, linear; fifth tarsomeres glabrous beneath; terminal sternum of male not modified. This species is very similar to
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<taxonomicName id="B8398718DD03D1F6527EBAE19A61E317" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus beneshi" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="beneshi">Pterostichus beneshi</taxonomicName>
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. Their comparison has been provided in the diagnosis under
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<taxonomicName id="BDF01B1547F8561D38ADC17FDF5B85CA" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus beneshi" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="beneshi">Pterostichus beneshi</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="28ED418E63CDD46D9664A015D418113C" pageId="42" pageNumber="43" type="supplemental description">
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<paragraph id="843692F6FFEC10327651E9B212AD0ED8" pageId="42" pageNumber="43">Supplemental description.</paragraph>
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<paragraph id="917D6AD7D341C5EF7710EC87369A116E" pageId="42" pageNumber="43">Male sternum. Penultimate and terminal sterna without special structures. Endophallus (Fig. 49D, E, G) extended in dorsal-basal direction of aedeagus, major portion of endophallus located on dorsal side of aedeagus; gonopore (gp, gonopore lobe folded in Fig. 49) located at level near middle of aedeagus, pointing to right. Five distinct lobes recognized: left lobe (ll) largest, located on left face near gp, its apex branched, forming two rounded sub-lobes, upper half of ll decorated with scales, scales near apex of ll very coarse; ventral lobe I (vl-I) tubiform, pointing to dorsal side of aedeagus, apex a little enlarged; ventral lobe II (vl-II) much smaller than vl-I; right lobe (rl) longer and thicker than vl-II, tubiform, straight; pre-apical lobe (pa) located on right face, close to gp, apex enlarged and branched, with very fine scales; basal-dorsal region on endophallus heavily pigmented; endophallus without chitinized piece. Female genitalia. Spermatheca with seminal canal approx three times as long as receptaculum; receptaculum capitate (Fig. 69), club approx half length of receptaculum; seminal canal inserted at base of common oviduct, base of seminal canal with sclerotized region short and thick. Stylomere II with two ensiform setae at basal half of outer margin, and a small one near middle of inner margin; two short nematiform setae located in a furrow near apex. Female sternum VIII (Fig. 88B) with fine setae on posterior margin, setae on middle region a little thicker; posterior margin almost straight, notched in middle; posterior region well chitinized; anterior region semi-chitinized, deeply notched in middle; middle transparent region small, roughly diamond-shaped, with two short oblique extensions; middle transparent region adjacent to anterior and posterior notches in middle. Female tergum VIII (Fig. 88A) short, posterior region semi-chitinized, without denser pigmentation; anterior region evenly chitinized; anterior margin notched in middle, posterior margin evenly arcuate.</paragraph>
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</subSubSection>
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<subSubSection id="2FE336AEC2CB06295FD4124606757439" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="distribution">
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<paragraph id="1A563A4B3BD4FA5077EB1C2C29C05B72" pageId="42" pageNumber="43">Distribution.</paragraph>
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<paragraph id="7B7B76DB7855BDD0EDE7F0D9FA2AD6C7" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
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Known only from Emei mountain in central Sichuan province (Map 2). The label of the holotype indicates that this species is from low altitude (600-1050 m) on Emei Mt. As a result of our expedition to Emei Mt. in 2012, we
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<pageBreakToken id="9552AECBDE48CD2994F5CD7E5A39C863" pageId="43" pageNumber="44" start="start">found</pageBreakToken>
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that this species is very common at relatively high altitudes (more than 2000 m) of Emei Mountain.
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</paragraph>
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</subSubSection>
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<subSubSection id="D18563781EF3B822ED6D5758D66FDD88" pageId="43" pageNumber="44" type="affinities">
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<paragraph id="CF75E9BEFB270412AD924A8295E225C1" pageId="43" pageNumber="44">Affinities.</paragraph>
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<paragraph id="44F4F252D1706B343F8EC321DF4815A5" pageId="43" pageNumber="44">
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This species is close to
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<taxonomicName id="C6B722F15B37E61AE9965DDB3774022C" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus beneshi" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="beneshi">Pterostichus beneshi</taxonomicName>
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in similarities of external appearance and male genitalia. Their endophalli are also similar, all lobes being recognizable as homologous.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |