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<document ID-DOI="http://dx.doi.org/10.3897/jhr.36.6830" ID-GBIF-Taxon="182224383" ID-Pensoft-Pub="1314-2607-36-131" ID-Pensoft-UUID="FFC442199F74FFDB022D1222FF875B6C" ID-Zenodo-Dep="574834" ModsDocID="1314-2607-36-131" checkinTime="1451252323662" checkinUser="pensoft" docAuthor="Nugroho, Hari, Kojima, Jun-ichi &amp; Ubaidillah, Rosichon" docDate="2014" docId="AB9F9996FE465C0A9ED3D57FB36ECB23" docLanguage="en" docName="JourHymenoptRes 36: 131-151" docOrigin="Journal of Hymenoptera Research 36" docPubDate="2014-02-14" docSource="http://dx.doi.org/10.3897/jhr.36.6830" docTitle="Apodynerus yayeyamensis" docType="treatment" docVersion="2" id="FFC442199F74FFDB022D1222FF875B6C" lastPageId="9" lastPageNumber="139" masterDocId="FFC442199F74FFDB022D1222FF875B6C" masterDocTitle="Synonymy of the potter wasp genus Philippodynerus Gusenleitner (Hymenoptera, Vespidae, Eumeninae) with Apodynerus Giordani Soika, with taxonomic notes on Apodynerus species" masterLastPageNumber="151" masterPageNumber="131" pageId="7" pageNumber="138" updateTime="1643517572533" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Synonymy of the potter wasp genus Philippodynerus Gusenleitner (Hymenoptera, Vespidae, Eumeninae) with Apodynerus Giordani Soika, with taxonomic notes on Apodynerus species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nugroho, Hari</mods:namePart>
<mods:affiliation>Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Sciences, Jl. Raya Jakarta-Bogor Km 46, Cibinong, Bogor, Indonesia</mods:affiliation>
<mods:nameIdentifier type="email">hntawon@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kojima, Jun-ichi</mods:namePart>
<mods:affiliation>Natural History Laboratory, Faculty of Science, Ibaraki University, Mito, 310 - 8512 Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ubaidillah, Rosichon</mods:namePart>
<mods:affiliation>Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Sciences, Jl. Raya Jakarta-Bogor Km 46, Cibinong, Bogor, Indonesia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-02-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>36</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>131</mods:start>
<mods:end>151</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.36.6830</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.36.6830</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-36-131</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFC442199F74FFDB022D1222FF875B6C</mods:identifier>
<mods:identifier type="Zenodo-Dep">574834</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182224383" LSID="urn:lsid:plazi:treatment:AB9F9996FE465C0A9ED3D57FB36ECB23" httpUri="http://treatment.plazi.org/id/AB9F9996FE465C0A9ED3D57FB36ECB23" lastPageId="9" lastPageNumber="139" pageId="7" pageNumber="138">
<subSubSection pageId="7" pageNumber="138" type="nomenclature">
<paragraph pageId="7" pageNumber="138">
<taxonomicName LSID="http://species-id.net/wiki/Apodynerus_yayeyamensis" authority="(Matsumura)" baseAuthorityName="Matsumura" class="Insecta" family="Eumenidae" genus="Apodynerus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Apodynerus yayeyamensis" order="Hymenoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="yayeyamensis">
<pageBreakToken pageId="7" pageNumber="138" start="start">Apodynerus</pageBreakToken>
yayeyamensis (Matsumura)
</taxonomicName>
<figureCitation captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Apodynerus species. 11 - 14 ♀ 15 - 18 ♂. 11, 15 Apodynerus yayeyamensis 12, 16 Apodynerus quadricolor 13, 17 Apodynerus flavospinosus, holotype and paratype - allotype, respectively 14, 18 Apodynerus icarioides 11 - 18 Clypeus, frontal view." httpUri="https://binary.pensoft.net/fig/11847" pageId="7" pageNumber="138">Figs 11</figureCitation>
<figureCitation captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Apodynerus species. 11 - 14 ♀ 15 - 18 ♂. 11, 15 Apodynerus yayeyamensis 12, 16 Apodynerus quadricolor 13, 17 Apodynerus flavospinosus, holotype and paratype - allotype, respectively 14, 18 Apodynerus icarioides 11 - 18 Clypeus, frontal view." httpUri="https://binary.pensoft.net/fig/11847" pageId="7" pageNumber="138">, 15</figureCitation>
<figureCitation captionStart="Figures 19–29" captionStartId="F3" captionText="Figures 19 - 29. Apodynerus species. 19 - 27 ♀ 28 - 29 ♂ 19, 24, 29 Apodynerus icarioides 20, 23 Apodynerus flavospinosus, holotype 25 Apodynerus troglodytes 21, 26 Apodynerus yayeyamensis 22, 27, 28 Apodynerus quadricolor 19 - 22 Head, lateral view 23 - 27 Metasomal segments I-II, lateral view 28, 29 Apical part of antenna. Scale 1 mm." httpUri="https://binary.pensoft.net/fig/11848" pageId="7" pageNumber="138">, 21</figureCitation>
<figureCitation captionStart="Figures 19–29" captionStartId="F3" captionText="Figures 19 - 29. Apodynerus species. 19 - 27 ♀ 28 - 29 ♂ 19, 24, 29 Apodynerus icarioides 20, 23 Apodynerus flavospinosus, holotype 25 Apodynerus troglodytes 21, 26 Apodynerus yayeyamensis 22, 27, 28 Apodynerus quadricolor 19 - 22 Head, lateral view 23 - 27 Metasomal segments I-II, lateral view 28, 29 Apical part of antenna. Scale 1 mm." httpUri="https://binary.pensoft.net/fig/11848" pageId="7" pageNumber="138">, 26</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="138" type="reference_group">
<paragraph pageId="7" pageNumber="138">
<taxonomicName class="Hexapoda" family="Vespidae" genus="Odynerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="yayeyamensis">Odynerus yayeyamensis</taxonomicName>
Matsumura, 1926,
<emphasis italics="true" pageId="7" pageNumber="138">in</emphasis>
<bibRefCitation author="Matsumura, S" journalOrPublisher="Insecta Matsumurana" pageId="17" pageNumber="148" pagination="32 - 52" refId="B22" refString="Matsumura, S, Uchida, T, 1926. Die Hymenopteren-Fauna von den Riukiu-Inseln. Insecta Matsumurana 1 (1): 32 - 52" title="Die Hymenopteren-Fauna von den Riukiu-Inseln." volume="1" year="1926">Matsumura and Uchida 1926</bibRefCitation>
: 36, ♂, &quot;Okinawa (Okinawa-honto)&quot; [possibly erroneously listed locality], lectotype from Yaeyama Islands (SEIHU).
</paragraph>
<paragraph pageId="7" pageNumber="138">
<taxonomicName authorityName="Sonan" authorityYear="1929" class="Hexapoda" family="Vespidae" genus="Odynerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="hokotoensis">Odynerus hokotoensis</taxonomicName>
<bibRefCitation author="Sonan, J" journalOrPublisher="Transactions of the Natural History Society of Formosa" pageId="18" pageNumber="149" pagination="533 - 540" refId="B28" refString="Sonan, J, 1929. Some wasps and bees of Hokoto (Pescadores Islands). Transactions of the Natural History Society of Formosa 19: 533 - 540" title="Some wasps and bees of Hokoto (Pescadores Islands)." volume="19" year="1929">Sonan 1929</bibRefCitation>
: 534, ♀♂, [Taiwan:]
<normalizedToken originalValue="“Hôkotô">&quot;Hokoto</normalizedToken>
... Takao&quot;. Holotype ♀ originally in &quot;Entomological Laboratory, Taihoku Imperial University&quot;, but current depository unknown.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138" type="description of female and lectotype male">
<paragraph pageId="7" pageNumber="138">Description of female and lectotype male.</paragraph>
<paragraph pageId="7" pageNumber="138">
FEMALE. Head in frontal view about as wide as high. Clypeus in frontal view wider than high, 1.2
<normalizedToken originalValue="×">x</normalizedToken>
as wide as high (
<figureCitation captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Apodynerus species. 11 - 14 ♀ 15 - 18 ♂. 11, 15 Apodynerus yayeyamensis 12, 16 Apodynerus quadricolor 13, 17 Apodynerus flavospinosus, holotype and paratype - allotype, respectively 14, 18 Apodynerus icarioides 11 - 18 Clypeus, frontal view." httpUri="https://binary.pensoft.net/fig/11847" pageId="7" pageNumber="138">Fig. 11</figureCitation>
). Vertical anterior face of pronotum with series of conspicuously produced striae. Metanotum compressed, obtusely dentiform; posterior margin oblique in lateral view. Metasomal tergum II in lateral view distinctly swollen anterodorsally and barely convex in posterior two-thirds (
<figureCitation captionStart="Figures 19–29" captionStartId="F3" captionText="Figures 19 - 29. Apodynerus species. 19 - 27 ♀ 28 - 29 ♂ 19, 24, 29 Apodynerus icarioides 20, 23 Apodynerus flavospinosus, holotype 25 Apodynerus troglodytes 21, 26 Apodynerus yayeyamensis 22, 27, 28 Apodynerus quadricolor 19 - 22 Head, lateral view 23 - 27 Metasomal segments I-II, lateral view 28, 29 Apical part of antenna. Scale 1 mm." httpUri="https://binary.pensoft.net/fig/11848" pageId="7" pageNumber="138">Fig. 26</figureCitation>
).
</paragraph>
<paragraph pageId="7" pageNumber="138">Head and mesosoma black, with following yellow markings: interantennal spot, ocular sinus, scape ventrally, mandible basally, band on gena, large dorsal spot and ventral scrobal spot on clypeus; anterior half of dorsal face of pronotum interrupted medially; spot on upper part of mesepisternum, tegula except median ferruginous spot, parategula, scutellum nearly entirely, large spots on dorsal face of propodeum; antenna dark brown. Legs black, but apical part of fore and mid femora and outer face of tibiae yellow. Metasoma black, with following yellow markings: tergum I: paired sublateral bands running obliquely towards posterolateral corner of tergum and apical band; segment II: basal lateral spots on tergum, and apical band.</paragraph>
<paragraph pageId="7" pageNumber="138">
Lectotype male. Clypeus in frontal view about as wide as high; ventral margin proportionally narrow (
<figureCitation captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Apodynerus species. 11 - 14 ♀ 15 - 18 ♂. 11, 15 Apodynerus yayeyamensis 12, 16 Apodynerus quadricolor 13, 17 Apodynerus flavospinosus, holotype and paratype - allotype, respectively 14, 18 Apodynerus icarioides 11 - 18 Clypeus, frontal view." httpUri="https://binary.pensoft.net/fig/11847" pageId="7" pageNumber="138">Fig. 15</figureCitation>
), about 0.2
<normalizedToken originalValue="×">x</normalizedToken>
the maximum width of clypeus. Scape 3.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as its maximum width; flagellomere X small; flagellomere XI minute, curved backward, with apex reaching about half length of flagellomere IX. Mandible with five teeth; apicalmost tooth long and sharp.
</paragraph>
<paragraph pageId="7" pageNumber="138">
Mesosoma in lateral view about 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as high; in dorsal view about 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. Metanotum dorsally with closely pair of obtuse teeth. Propodeum with somewhat deep median concavity on posterior face, in lateral view strongly convex.
</paragraph>
<paragraph pageId="7" pageNumber="138">Lateral face of propodeum with dense, strong punctures, in most specimens forming ridges between punctures. Metasomal segment I with dense, conspicuous deep punctures; segments II-VII with dense, deep punctures.</paragraph>
<paragraph lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138">
Black, with following yellow markings: interantennal spot, spot filling ocular sinus extending ventrally to antennal socket, band on gena, clypeus except apical ferruginous band and black transverse band at level of lateral lobes, scape except black dorsal face, mandible except ferruginous tip and narrow black band along outer margin; anterodorsal part of pronotum, scrobal spot on mesepisternum, tegula except ferruginous mid spot and outer margin, parategula, scutellum nearly entirely, spot on metanotal
<pageBreakToken pageId="8" pageNumber="139" start="start">teeth</pageBreakToken>
, paired large dorsal spots on propodeum; markings on metasoma as in female, but terga IV-V with posteromedian yellow spot.
</paragraph>
<paragraph pageId="8" pageNumber="139">Color variation in males from Taiwan. Sublateral yellow bands on tergum I in some specimens reduced to small spots or lacking, terga IV-V and occasionally also VI with posteromedian yellow spot, and sternum III sometimes with apical yellow band.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="139" type="material examined">
<paragraph pageId="8" pageNumber="139">Material examined.</paragraph>
<paragraph pageId="8" pageNumber="139">
JAPAN: 1 ♂ (SEIHU; lectotype of
<taxonomicName class="Insecta" family="Eumenidae" genus="Apodynerus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="yayeyamensis">
<emphasis italics="true" pageId="8" pageNumber="139">Apodynerus yayeyamensis</emphasis>
</taxonomicName>
), labeled &quot;SK, Japan Matsumra, Yaeyama, VIII 07&quot;, &quot;
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Vespidae" genus="Odynerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="8" pageNumber="139">Odynerus</emphasis>
</taxonomicName>
n. sp.
<emphasis italics="true" pageId="8" pageNumber="139">yayeyamensis</emphasis>
, det. Matsumura&quot;, &quot;Type Matsumura&quot;, and &quot;
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1943" class="Hexapoda" family="Vespidae" genus="Pachymenes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis italics="true" pageId="8" pageNumber="139">Pachymenes fragilis</emphasis>
</taxonomicName>
(Smith) det. K. Yasumatsu, 1938&quot;; 1 ♀, Motobu, Sesoko I., 9 Sep.1982, J. Kojima (IUNH). TAIWAN: 2 ♂, Tojuko, Prov. Taichu [=Taichung], 1-8 Aug.1931, N. Owaki (SEIHU).
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="140" pageId="8" pageNumber="139" type="remarks">
<paragraph pageId="8" pageNumber="139">Remarks.</paragraph>
<paragraph lastPageId="9" lastPageNumber="140" pageId="8" pageNumber="139">
<bibRefCitation author="Matsumura, S" journalOrPublisher="Insecta Matsumurana" pageId="17" pageNumber="148" pagination="32 - 52" refId="B22" refString="Matsumura, S, Uchida, T, 1926. Die Hymenopteren-Fauna von den Riukiu-Inseln. Insecta Matsumurana 1 (1): 32 - 52" title="Die Hymenopteren-Fauna von den Riukiu-Inseln." volume="1" year="1926">Matsumura and Uchida (1926)</bibRefCitation>
mentioned the two males collected on
<normalizedToken originalValue="“Okinawa-honto”">&quot;Okinawa-honto&quot;</normalizedToken>
[=Okinawa Island] by S. Kiyamu and S. Sakaguchi in the description of
<taxonomicName class="Hexapoda" family="Vespidae" genus="Odynerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="9" pageNumber="140" phylum="Arthropoda" rank="species" species="yayeyamensis">
<emphasis italics="true" pageId="9" pageNumber="140">
<pageBreakToken pageId="9" pageNumber="140" start="start">Odynerus</pageBreakToken>
yayeyamensis
</emphasis>
</taxonomicName>
[=
<taxonomicName class="Insecta" family="Eumenidae" genus="Apodynerus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="9" pageNumber="140" phylum="Arthropoda" rank="species" species="yayeyamensis">
<emphasis italics="true" pageId="9" pageNumber="140">Apodynerus yayeyamensis</emphasis>
</taxonomicName>
]. Although, as
<bibRefCitation author="Yamane, S" journalOrPublisher="Insecta matsumurana (new series)" pageId="18" pageNumber="149" pagination="1 - 189" refId="B30" refString="Yamane, S, 1990. A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea). Insecta matsumurana (new series) 43: 1 - 189" title="A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea)." volume="43" year="1990">Yamane (1990)</bibRefCitation>
pointed out, the locality
<normalizedToken originalValue="“Yaeyama”">&quot;Yaeyama&quot;</normalizedToken>
written on the label was not mentioned in the original description, the two male specimens referred to by
<bibRefCitation author="Yamane, S" journalOrPublisher="Insecta matsumurana (new series)" pageId="18" pageNumber="149" pagination="1 - 189" refId="B30" refString="Yamane, S, 1990. A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea). Insecta matsumurana (new series) 43: 1 - 189" title="A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea)." volume="43" year="1990">Yamane (1990</bibRefCitation>
: 137) as &quot;The type specimen&quot; and that &quot;believed to be the one in the type-series&quot; are certainly the syntypes.
<bibRefCitation author="Yamane, S" journalOrPublisher="Insecta matsumurana (new series)" pageId="18" pageNumber="149" pagination="1 - 189" refId="B30" refString="Yamane, S, 1990. A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea). Insecta matsumurana (new series) 43: 1 - 189" title="A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea)." volume="43" year="1990">Yamane (1990</bibRefCitation>
: 137) inadvertently designated the lectotype by specifically referring to one of the two syntypes as &quot;The type specimen&quot; (Article 74.6.1 in the Code, ICZN 1999). The lectotype we located in the SEIHU bears the following labels: (1) on upper side
<normalizedToken originalValue="“SK”">&quot;SK&quot;</normalizedToken>
[=?S. Kiyamu] in handwriting and &quot;Japan Matsumra&quot; in print; on underside,
<normalizedToken originalValue="“Yaeyama”">&quot;Yaeyama&quot;</normalizedToken>
[Chinese characters] and &quot;VIII 07&quot; [=August 1907;
<bibRefCitation author="Yamane, S" journalOrPublisher="Insecta matsumurana (new series)" pageId="18" pageNumber="149" pagination="1 - 189" refId="B30" refString="Yamane, S, 1990. A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea). Insecta matsumurana (new series) 43: 1 - 189" title="A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea)." volume="43" year="1990">Yamane (1990)</bibRefCitation>
read &quot;vii[?] 0
<normalizedToken originalValue="’7”">'7&quot;</normalizedToken>
], both in handwriting, (2) &quot;
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Hexapoda" family="Vespidae" genus="Odynerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="9" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="140">Odynerus</emphasis>
</taxonomicName>
n.sp.
<emphasis italics="true" pageId="9" pageNumber="140">yayeyamensis</emphasis>
&quot; in handwriting, and &quot;det. Matsumura&quot; in print, (3) red label &quot;Type Matsumura&quot; in print, (4)
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1943" class="Hexapoda" family="Vespidae" genus="Pachymenes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="9" pageNumber="140" phylum="Arthropoda" rank="species" species="fragilis">
<emphasis italics="true" pageId="9" pageNumber="140">Pachymenes fragilis</emphasis>
</taxonomicName>
(Smith) det. K. Yasumatsu, 1938&quot; in handwriting. We have added a label &quot;LECTOTYPE [in red]/
<taxonomicName class="Hexapoda" family="Vespidae" genus="Odynerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="9" pageNumber="140" phylum="Arthropoda" rank="species" species="yayeyamensis">
<emphasis italics="true" pageId="9" pageNumber="140">Odynerus yayeyamensis</emphasis>
</taxonomicName>
/ Matsumura, 1926/by designation of/
<bibRefCitation author="Yamane, S" journalOrPublisher="Insecta matsumurana (new series)" pageId="18" pageNumber="149" pagination="1 - 189" refId="B30" refString="Yamane, S, 1990. A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea). Insecta matsumurana (new series) 43: 1 - 189" title="A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea)." volume="43" year="1990">Yamane (1990)</bibRefCitation>
&quot; [a slash denotes the start of a new line]. We searched in vain in the SEIHU for the other specimen, the paralectotype, that according to
<bibRefCitation author="Yamane, S" journalOrPublisher="Insecta matsumurana (new series)" pageId="18" pageNumber="149" pagination="1 - 189" refId="B30" refString="Yamane, S, 1990. A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea). Insecta matsumurana (new series) 43: 1 - 189" title="A revision of the Japanese Eumenidae (Hymenoptera, Vespoidea)." volume="43" year="1990">Yamane (1990)</bibRefCitation>
is labeled (1)
<normalizedToken originalValue="“56”">&quot;56&quot;</normalizedToken>
in handwriting and (2) &quot;Okinawa S. Sakaguchi&quot; in print.
</paragraph>
</subSubSection>
</treatment>
</document>
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