308 lines
40 KiB
XML
308 lines
40 KiB
XML
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<mods:title id="40C158B3CFF10F19A95BA05BE4532E4B">New species of Cirratulidae (Annelida, Polychaeta) from abyssal depths of the Clarion-Clipperton Fracture Zone, North Equatorial Pacific Ocean</mods:title>
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<mods:namePart id="EFD556DF53A4B20B0DC5C8F98CF0A18F">Blake, James A.</mods:namePart>
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<mods:title id="F56493651480C583D336C2B707AE0999">Zootaxa</mods:title>
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<mods:part id="8AB8964F3C42F896B2EC3D203C62A6B2">
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<mods:date id="32AE7D560DFBF330EB959D290BF097C4">2019</mods:date>
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<mods:identifier id="ED0D39FAA0FB6DD1FBF44004E4BE66EF" type="DOI">10.11646/zootaxa.4629.2.1</mods:identifier>
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<treatment id="AB0E185AC466FFAF4FE91DF8AF3726D0" ID-DOI="http://doi.org/10.5281/zenodo.5619243" ID-GBIF-Taxon="158525132" ID-Zenodo-Dep="5619243" LSID="urn:lsid:zoobank.org:act:7472BB3A-8A19-42D9-B2EF-114C06DA01B7" httpUri="http://treatment.plazi.org/id/AB0E185AC466FFAF4FE91DF8AF3726D0" lastPageId="31" lastPageNumber="182" pageId="30" pageNumber="181">
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<paragraph id="2318A94CC466FFAE4FE91DF8ADCF2188" blockId="30.[151,937,153,244]" box="[151,491,153,180]" pageId="30" pageNumber="181">
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<heading id="78501E20C466FFAE4FE91DF8ADCF2188" bold="true" box="[151,491,153,180]" fontSize="11" level="1" pageId="30" pageNumber="181" reason="1">
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<emphasis id="11D3755EC466FFAE4FE91DF8ADCF2188" bold="true" box="[151,491,153,180]" pageId="30" pageNumber="181">
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<taxonomicName id="E4A7D2CFC466FFAE4FE91DF8AD70218F" ID-CoL="85S5H" authority="Blake, 2019" authorityName="Blake" authorityYear="2019" box="[151,340,153,180]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="30" pageNumber="181" phylum="Annelida" rank="species" species="hessleri" status="sp. nov.">
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<emphasis id="11D3755EC466FFAE4FE91DF8AD70218F" bold="true" box="[151,340,153,180]" italics="true" pageId="30" pageNumber="181">Tharyx hessleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="0AE0C825C466FFAE4E251DFBADCF2188" box="[347,491,153,180]" pageId="30" pageNumber="181" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6BBDFAC7C466FFAE4FE91DDCAF6D2015" pageId="30" pageNumber="181" type="description">
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<paragraph id="2318A94CC466FFAE4FE91DDCAD1C21EC" blockId="30.[151,937,153,244]" box="[151,312,189,215]" pageId="30" pageNumber="181">
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<figureCitation id="BB9CB5C9C466FFAE4FE91DDCAD1C21EC" box="[151,312,189,215]" captionStart-0="FIGURE 19" captionStart-1="FIGURE 20" captionStartId-0="30.[151,250,1973,1998]" captionStartId-1="31.[151,250,957,982]" captionTargetBox-0="[159,1401,734,1944]" captionTargetBox-1="[155,1429,189,930]" captionTargetId-0="figure@30.[151,1409,720,1954]" captionTargetId-1="figure@31.[151,1436,181,938]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="FIGURE 19. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary." captionText-1="FIGURE 20. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287):A, anterior end through setiger 14, right lateral view; B, far posterior segments, dorsal view; C, posterior 3 setigers and pygidium, dorsal view. Stained with Shirlastain A." httpUri-0="https://zenodo.org/record/3269029/files/figure.png" httpUri-1="https://zenodo.org/record/3269031/files/figure.png" pageId="30" pageNumber="181">Figures 19–20</figureCitation>
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</paragraph>
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<paragraph id="2318A94CC466FFAE4FE91DBDAF8D21CF" blockId="30.[151,937,153,244]" box="[151,937,220,244]" pageId="30" pageNumber="181">
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<uri id="5736A54EC466FFAE4FE91DBDAF8D21CF" box="[151,937,220,244]" pageId="30" pageNumber="181">
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urn:lsid:zoobank.org:act:
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<uuid id="57019399C466FFAE4EDE1DBDAF8D21CF" box="[416,937,220,244]" pageId="30" pageNumber="181">7472BB3A-8A19-42D9-B2EF-114C06DA01B7</uuid>
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</uri>
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</paragraph>
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<paragraph id="2318A94CC466FFAE4FE91C77AF6D2015" blockId="30.[151,841,277,302]" box="[151,841,277,302]" pageId="30" pageNumber="181">
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<treatmentCitationGroup id="03B78E62C466FFAE4FE91C77AF6D2015" box="[151,841,277,302]" pageId="30" pageNumber="181">
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<taxonomicName id="E4A7D2CFC466FFAE4FE91C77AD2B2015" ID-CoL="84M8M" authorityName="Malmgren" authorityYear="1867" box="[151,271,278,302]" class="Polychaeta" family="Cirratulidae" genus="Chaetozone" kingdom="Animalia" order="Terebellida" pageId="30" pageNumber="181" phylum="Annelida" rank="genus">
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<emphasis id="11D3755EC466FFAE4FE91C77AD2B2015" box="[151,271,278,302]" italics="true" pageId="30" pageNumber="181">Chaetozone</emphasis>
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</taxonomicName>
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||
sp. A:
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<treatmentCitation id="A2068F5DC466FFAE4E281C74AE4E2015" author="Wilson, G. D. F. & Hessler, R. R." box="[342,618,277,302]" page="66" pageId="30" pageNumber="181" year="1987">
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<bibRefCitation id="4736D4BDC466FFAE4E281C74AE4E2015" author="Wilson, G. D. F. & Hessler, R. R." box="[342,618,277,302]" pageId="30" pageNumber="181" pagination="24 - 86" refId="ref21781" refString="Wilson, G. D. F. & Hessler, R. R. (1987) The effects of manganese nodule test mining on the benthic fauna in the North Equatorial Pacific. In: Spiess, F. N., Hessler, R., Wilson, G. & Weydert, M. (Eds.), Environmental effects of deep sea dredging. Final Report prepared for the National Oceanographic and Atmospheric Administration under Contract NO. 83 - SAC- 00659. Scripps Institution of Oceanography, La Jolla, SIO Reference 87 - 5, pp. 24 - 86, appendices A-H. https: // doi. org / 10.13140 / RG. 2.1.1024.2080" type="journal article" year="1987">Wilson & Hessler 1987: 66</bibRefCitation>
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</treatmentCitation>
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||
Appendix E (in part).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="6BBDFAC7C466FFAE4FE91C3AAF7F20D9" pageId="30" pageNumber="181" type="materials_examined">
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<paragraph id="2318A94CC466FFAE4FE91C3AAF7F20D9" blockId="30.[151,1437,347,699]" pageId="30" pageNumber="181">
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<emphasis id="11D3755EC466FFAE4FE91C3AAD53204D" bold="true" box="[151,375,347,374]" pageId="30" pageNumber="181">Material examined</emphasis>
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||
.
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||
<emphasis id="11D3755EC466FFAE4EFD1C3DAC8D20A1" bold="true" pageId="30" pageNumber="181">
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||
North Equatorial Pacific Ocean, abyssal plain, Clarion-Clipperton Fracture Zone,
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||
<taxonomicName id="E4A7D2CFC466FFAE4A341C3AA9B8204D" box="[1354,1436,347,374]" class="Insecta" family="Calopterygidae" genus="Echo" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="186" phylum="Arthropoda" rank="genus">ECHO</taxonomicName>
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||
I,
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||
</emphasis>
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||
<materialsCitation id="93CFA311C466FFAE4FCF1C1EAF6D2085" ID-GBIF-Occurrence="2412500457" collectingDate="1983-06-04" collectingDateMax="1983-06-17" collectingDateMin="1983-06-04" collectionCode="DOMES, LACM-AHF" collectorName="Site C & V Melville & R. Hessler & Sta. H" elevation="4506" latitude="14.679978" longLatPrecision="1" longitude="-125.3673" pageId="30" pageNumber="181" specimenCount="2" typeStatus="holotype">
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<emphasis id="11D3755EC466FFAE4FCF1C1EAD4320A1" bold="true" box="[177,359,383,410]" pageId="30" pageNumber="181">
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<collectionCode id="45B63189C466FFAE4FCF1C1EAD3C20A1" box="[177,280,383,410]" pageId="30" pageNumber="181">DOMES</collectionCode>
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<collectorName id="8E52CC9AC466FFAE4E611C1EAD4320A1" box="[287,359,383,410]" pageId="30" pageNumber="181">Site C</collectorName>
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||
</emphasis>
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||
, R/
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||
<collectorName id="8E52CC9AC466FFAE4EEF1CE1AE2C20A1" box="[401,520,384,410]" pageId="30" pageNumber="181">
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||
V
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||
<emphasis id="11D3755EC466FFAE4ED21CE1AE2C20A1" box="[428,520,384,410]" italics="true" pageId="30" pageNumber="181">Melville</emphasis>
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</collectorName>
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cruise, coll.
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<collectorName id="8E52CC9AC466FFAE4DE71CE0AF3420A1" box="[665,784,384,411]" pageId="30" pageNumber="181">R. Hessler</collectorName>
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,
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||
<collectorName id="8E52CC9AC466FFAE4C621CE1AF4720A1" box="[796,867,383,410]" pageId="30" pageNumber="181">Sta. H</collectorName>
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350C,
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||
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fraction,
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<collectingDate id="475D7664C466FFAE4B101CE1A8C920A1" box="[1134,1261,384,411]" pageId="30" pageNumber="181" value="1983-06-04">
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<date id="57198F8CC466FFAE4B101CE1A8C920A1" box="[1134,1261,384,411]" pageId="30" pageNumber="181" value="1983-06-04">4 Jun 1983</date>
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</collectingDate>
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,
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<geoCoordinate id="4693CF8BC466FFAE4B871C1EA9BC20A1" box="[1273,1432,383,410]" degrees="14" direction="north" minutes="38.1226" orientation="latitude" pageId="30" pageNumber="181" precision="1" value="14.635377">14°38.1226ʹN</geoCoordinate>
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,
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<geoCoordinate id="4693CF8BC466FFAE4FE91CC2AD6D2085" box="[151,329,419,446]" degrees="125" direction="west" minutes="26.8208" orientation="longitude" pageId="30" pageNumber="181" precision="1" value="-125.447014">125°26.8208ʹW</geoCoordinate>
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,
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<elevation id="A88A4E7FC466FFAE4E2C1CC2AD832084" box="[338,423,419,447]" metricMagnitude="3" metricUnit="m" metricValue="4.506" pageId="30" pageNumber="181" unit="m" value="4506.0">
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</elevation>
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,
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<typeStatus id="FC1C17EEC466FFAE4ECC1CC5AE3D2085" box="[434,537,420,446]" pageId="30" pageNumber="181" type="holotype">
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<emphasis id="11D3755EC466FFAE4ECC1CC5AE3D2085" bold="true" box="[434,537,420,446]" pageId="30" pageNumber="181">holotype</emphasis>
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</typeStatus>
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(
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<collectionCode id="45B63189C466FFAE4D5B1CC5AE9D2085" box="[549,697,420,446]" pageId="30" pageNumber="181">LACM-AHF</collectionCode>
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Poly 11287)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="93CFA311C466FFAE4C2D1CC5AF7320D9" ID-GBIF-Occurrence="2412500458" collectingDate="1983-06-04" collectingDateMax="1983-06-17" collectingDateMin="1983-06-04" collectionCode="DOMES, LACM-AHF" collectorName="Site C & V Melville & R. Hessler & Sta. H" elevation="4506" latitude="14.679978" longLatPrecision="1" longitude="-125.3673" pageId="30" pageNumber="181" specimenCount="2" typeStatus="paratype">
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<collectorName id="8E52CC9AC466FFAE4C2D1CC5AFBD2085" box="[851,921,420,446]" pageId="30" pageNumber="181">Sta. H</collectorName>
|
||
360, nodule wash,
|
||
<collectingDate id="475D7664C466FFAE4B181CC5A8CB2085" box="[1126,1263,420,447]" pageId="30" pageNumber="181" value="1983-06-17">
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||
<date id="57198F8CC466FFAE4B181CC5A8CB2085" box="[1126,1263,420,447]" pageId="30" pageNumber="181" value="1983-06-17">17 Jun 1983</date>
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||
</collectingDate>
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||
,
|
||
<geoCoordinate id="4693CF8BC466FFAE4B871CC2A9BC2085" box="[1273,1432,419,446]" degrees="14" direction="north" minutes="40.7987" orientation="latitude" pageId="30" pageNumber="181" precision="1" value="14.679978">14°40.7987ʹN</geoCoordinate>
|
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,
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||
<geoCoordinate id="4693CF8BC466FFAE4FE91CA6AD6D20D9" box="[151,329,455,482]" degrees="125" direction="west" minutes="22.0379" orientation="longitude" pageId="30" pageNumber="181" precision="1" value="-125.3673">125°22.0379ʹW</geoCoordinate>
|
||
,
|
||
<elevation id="A88A4E7FC466FFAE4E2D1CA6AD8E20D8" box="[339,426,455,483]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="30" pageNumber="181" unit="m" value="4500.0">
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<quantity id="E45F04A9C466FFAE4E2D1CA6AD8E20D8" box="[339,426,455,483]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="30" pageNumber="181" unit="m" value="4500.0">4500 m</quantity>
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||
</elevation>
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||
,
|
||
<typeStatus id="FC1C17EEC466FFAE4EC81CA9AE0520D9" box="[438,545,456,482]" pageId="30" pageNumber="181" type="paratype">
|
||
<emphasis id="11D3755EC466FFAE4EC81CA9AE0520D9" bold="true" box="[438,545,456,482]" pageId="30" pageNumber="181">paratype</emphasis>
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||
</typeStatus>
|
||
(
|
||
<collectionCode id="45B63189C466FFAE4D4E1CA9AEE020D9" box="[560,708,456,482]" pageId="30" pageNumber="181">LACM-AHF</collectionCode>
|
||
Poly 11288)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6BBDFAC7C466FFAF4FB91C8AAF232765" lastPageId="31" lastPageNumber="182" pageId="30" pageNumber="181" type="description">
|
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<paragraph id="2318A94CC466FFAE4FB91C8AAFEC2381" blockId="30.[151,1437,347,699]" pageId="30" pageNumber="181">
|
||
<emphasis id="11D3755EC466FFAE4FB91C8AAD75233D" bold="true" box="[199,337,491,518]" pageId="30" pageNumber="181">Description</emphasis>
|
||
. A small, threadlike species;
|
||
<typeStatus id="FC1C17EEC466FFAE4DEB1C8DAED3233D" box="[661,759,492,518]" pageId="30" pageNumber="181" type="holotype">holotype</typeStatus>
|
||
complete,
|
||
<quantity id="E45F04A9C466FFAE4C0F1C8DAFF0233C" box="[881,980,492,519]" metricMagnitude="-3" metricUnit="m" metricValue="3.72" pageId="30" pageNumber="181" unit="mm" value="3.72">3.72 mm</quantity>
|
||
long, peristomium
|
||
<quantity id="E45F04A9C466FFAE4BD31C8DA8D1233C" box="[1197,1269,492,519]" metricMagnitude="-2" metricUnit="m" metricValue="9.8" pageId="30" pageNumber="181" unit="mm" value="98.0">98 µm</quantity>
|
||
wide; anterior, middle, and posterior setigers of same width, about
|
||
<quantity id="E45F04A9C466FFAE4D991F71AF102310" box="[743,820,528,555]" metricMagnitude="-2" metricUnit="m" metricValue="9.2" pageId="30" pageNumber="181" unit="mm" value="92.0">92 µm</quantity>
|
||
, last few setigers narrowing slightly; with 39 setigers. Anterior setigers about 1.5 times as wide as long (
|
||
<figureCitation id="BB9CB5C9C466FFAE4DAC1F55AF1F2375" box="[722,827,564,590]" captionStart="FIGURE 19" captionStartId="30.[151,250,1973,1998]" captionTargetBox="[159,1401,734,1944]" captionTargetId="figure@30.[151,1409,720,1954]" captionTargetPageId="30" captionText="FIGURE 19. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary." httpUri="https://zenodo.org/record/3269029/files/figure.png" pageId="30" pageNumber="181">Fig. 19A</figureCitation>
|
||
); middle and posterior segments becoming somewhat rounded, as long as wide, with far posterior setigers weakly moniliform (
|
||
<figureCitation id="BB9CB5C9C466FFAE4CC41F39A80D2349" box="[954,1065,600,626]" captionStart="FIGURE 19" captionStartId="30.[151,250,1973,1998]" captionTargetBox="[159,1401,734,1944]" captionTargetId="figure@30.[151,1409,720,1954]" captionTargetPageId="30" captionText="FIGURE 19. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary." httpUri="https://zenodo.org/record/3269029/files/figure.png" pageId="30" pageNumber="181">Figs. 19B</figureCitation>
|
||
,
|
||
<figureCitation id="BB9CB5C9C466FFAE4B4D1F39A8742349" box="[1075,1104,600,626]" captionStart="FIGURE 20" captionStartId="31.[151,250,957,982]" captionTargetBox="[155,1429,189,930]" captionTargetId="figure@31.[151,1436,181,938]" captionTargetPageId="31" captionText="FIGURE 20. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287):A, anterior end through setiger 14, right lateral view; B, far posterior segments, dorsal view; C, posterior 3 setigers and pygidium, dorsal view. Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269031/files/figure.png" pageId="30" pageNumber="181">20</figureCitation>
|
||
B–C).
|
||
<typeStatus id="FC1C17EEC466FFAE4BE71F38A8DF2348" box="[1177,1275,601,627]" pageId="30" pageNumber="181" type="paratype">Paratype</typeStatus>
|
||
(LACM-AHF Poly H360) smaller, broken in two parts, only 2.0 mm long with 20 setigers. Body cylindrical in cross section, with no dorsal or ventral grooves. Color in alcohol opaque white; no pigment.
|
||
</paragraph>
|
||
<caption id="77D8F9C4C466FFAE4FE91AD4AF5826C9" httpUri="https://zenodo.org/record/3269029/files/figure.png" pageId="30" pageNumber="181" startId="30.[151,250,1973,1998]" targetBox="[159,1401,734,1944]" targetPageId="30">
|
||
<paragraph id="2318A94CC466FFAE4FE91AD4AF5826C9" blockId="30.[151,1436,1973,2034]" pageId="30" pageNumber="181">
|
||
<emphasis id="11D3755EC466FFAE4FE91AD4AE2E26F5" bold="true" box="[151,522,1973,1998]" pageId="30" pageNumber="181">
|
||
FIGURE 19.
|
||
<taxonomicName id="E4A7D2CFC466FFAE4E591AD4ADED26F5" authority="Blake, 2019" authorityName="Blake" authorityYear="2019" box="[295,457,1973,1998]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="30" pageNumber="181" phylum="Annelida" rank="species" species="hessleri" status="sp. nov.">
|
||
<emphasis id="11D3755EC466FFAE4E591AD4ADED26F5" bold="true" box="[295,457,1973,1998]" italics="true" pageId="30" pageNumber="181">Tharyx hessleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0AE0C825C466FFAE4EAE1AD7AE2E26F5" box="[464,522,1974,1998]" pageId="30" pageNumber="181" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="77D8F9C4C467FFAF4FE91EDCA91722C1" httpUri="https://zenodo.org/record/3269031/files/figure.png" pageId="31" pageNumber="182" startId="31.[151,250,957,982]" targetBox="[155,1429,189,930]" targetPageId="31">
|
||
<paragraph id="2318A94CC467FFAF4FE91EDCA91722C1" blockId="31.[151,1436,957,1018]" pageId="31" pageNumber="182">
|
||
<emphasis id="11D3755EC467FFAF4FE91EDCAE2222ED" bold="true" box="[151,518,957,982]" pageId="31" pageNumber="182">
|
||
FIGURE 20.
|
||
<taxonomicName id="E4A7D2CFC467FFAF4E5B1EDCADE322ED" authority="Blake, 2019" authorityName="Blake" authorityYear="2019" box="[293,455,957,982]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="182" phylum="Annelida" rank="species" species="hessleri" status="sp. nov.">
|
||
<emphasis id="11D3755EC467FFAF4E5B1EDCADE322ED" bold="true" box="[293,455,957,982]" italics="true" pageId="31" pageNumber="182">Tharyx hessleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0AE0C825C467FFAF4EB31EDFAE2222ED" box="[461,518,958,982]" pageId="31" pageNumber="182" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Holotype (LACM-AHF Poly 11287): A, anterior end through setiger 14, right lateral view; B, far posterior segments, dorsal view; C, posterior 3 setigers and pygidium, dorsal view. Stained with Shirlastain A.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2318A94CC467FFAF4FB91949A89225CC" blockId="31.[151,1437,1064,2027]" pageId="31" pageNumber="182">
|
||
Pre-setiger region approximately twice as long as wide, slightly longer than first three setigers (
|
||
<figureCitation id="BB9CB5C9C467FFAF4A5C1949A9BC2579" box="[1314,1432,1064,1090]" captionStart="FIGURE 19" captionStartId="30.[151,250,1973,1998]" captionTargetBox="[159,1401,734,1944]" captionTargetId="figure@30.[151,1409,720,1954]" captionTargetPageId="30" captionText="FIGURE 19. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary." httpUri="https://zenodo.org/record/3269029/files/figure.png" pageId="31" pageNumber="182">Figs. 19A</figureCitation>
|
||
,
|
||
<figureCitation id="BB9CB5C9C467FFAF4FE9192DACE8255D" box="[151,204,1100,1126]" captionStart="FIGURE 20" captionStartId="31.[151,250,957,982]" captionTargetBox="[155,1429,189,930]" captionTargetId="figure@31.[151,1436,181,938]" captionTargetPageId="31" captionText="FIGURE 20. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287):A, anterior end through setiger 14, right lateral view; B, far posterior segments, dorsal view; C, posterior 3 setigers and pygidium, dorsal view. Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269031/files/figure.png" pageId="31" pageNumber="182">20A</figureCitation>
|
||
). Prostomium triangular, tapering to narrow apex (
|
||
<figureCitation id="BB9CB5C9C467FFAF4C6D192DAFA3255D" box="[787,903,1100,1126]" captionStart="FIGURE 19" captionStartId="30.[151,250,1973,1998]" captionTargetBox="[159,1401,734,1944]" captionTargetId="figure@30.[151,1409,720,1954]" captionTargetPageId="30" captionText="FIGURE 19. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary." httpUri="https://zenodo.org/record/3269029/files/figure.png" pageId="31" pageNumber="182">Figs. 19A</figureCitation>
|
||
,
|
||
<figureCitation id="BB9CB5C9C467FFAF4CEB192DAFED255D" box="[917,969,1100,1126]" captionStart="FIGURE 20" captionStartId="31.[151,250,957,982]" captionTargetBox="[155,1429,189,930]" captionTargetId="figure@31.[151,1436,181,938]" captionTargetPageId="31" captionText="FIGURE 20. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287):A, anterior end through setiger 14, right lateral view; B, far posterior segments, dorsal view; C, posterior 3 setigers and pygidium, dorsal view. Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269031/files/figure.png" pageId="31" pageNumber="182">20A</figureCitation>
|
||
); eyespots absent; nuchal organs not observed. Peristomium entire, smooth, without annular rings (
|
||
<figureCitation id="BB9CB5C9C467FFAF4C491911AFBB25B1" box="[823,927,1136,1162]" captionStart="FIGURE 19" captionStartId="30.[151,250,1973,1998]" captionTargetBox="[159,1401,734,1944]" captionTargetId="figure@30.[151,1409,720,1954]" captionTargetPageId="30" captionText="FIGURE 19. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary." httpUri="https://zenodo.org/record/3269029/files/figure.png" pageId="31" pageNumber="182">Fig. 19A</figureCitation>
|
||
); dorsal tentacles arising from posterior margin; first pair of branchiae arising immediately posterior and slightly lateral to dorsal tentacles (
|
||
<figureCitation id="BB9CB5C9C467FFAF4BB319F5A9112595" box="[1229,1333,1172,1198]" captionStart="FIGURE 19" captionStartId="30.[151,250,1973,1998]" captionTargetBox="[159,1401,734,1944]" captionTargetId="figure@30.[151,1409,720,1954]" captionTargetPageId="30" captionText="FIGURE 19. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary." httpUri="https://zenodo.org/record/3269029/files/figure.png" pageId="31" pageNumber="182">Fig. 19A</figureCitation>
|
||
). Second pair of branchiae arising from setiger 1, dorsal to notosetae; subsequent branchiae from similar location. Branchiae present as stubs over anterior and some middle segments, not observed on posterior segments.
|
||
</paragraph>
|
||
<paragraph id="2318A94CC467FFAF4FB91861A972272D" blockId="31.[151,1437,1064,2027]" pageId="31" pageNumber="182">
|
||
Parapodia reduced to low mounds from which setae arise. Setae of anterior segments all capillaries with 4–5 notosetae, including some long, natatory-like, continuing through middle body (
|
||
<figureCitation id="BB9CB5C9C467FFAF4B491845A8892405" box="[1079,1197,1316,1342]" captionStart="FIGURE 19" captionStartId="30.[151,250,1973,1998]" captionTargetBox="[159,1401,734,1944]" captionTargetId="figure@30.[151,1409,720,1954]" captionTargetPageId="30" captionText="FIGURE 19. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary." httpUri="https://zenodo.org/record/3269029/files/figure.png" pageId="31" pageNumber="182">Figs. 19A</figureCitation>
|
||
,
|
||
<figureCitation id="BB9CB5C9C467FFAF4BC21845A8D42405" box="[1212,1264,1316,1342]" captionStart="FIGURE 20" captionStartId="31.[151,250,957,982]" captionTargetBox="[155,1429,189,930]" captionTargetId="figure@31.[151,1436,181,938]" captionTargetPageId="31" captionText="FIGURE 20. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287):A, anterior end through setiger 14, right lateral view; B, far posterior segments, dorsal view; C, posterior 3 setigers and pygidium, dorsal view. Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269031/files/figure.png" pageId="31" pageNumber="182">20A</figureCitation>
|
||
); capillaries of middle and posterior segments shorter, 4–5 per fascicle.
|
||
<typeStatus id="FC1C17EEC467FFAF4C6C1829AF5E2459" box="[786,890,1352,1378]" pageId="31" pageNumber="182" type="holotype">Holotype</typeStatus>
|
||
with neuro-acicular spines first present from setiger
|
||
<quantity id="E45F04A9C467FFAF4FAE180DAD2C24BD" box="[208,264,1388,1414]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="31" pageNumber="182" unit="in" value="18.0">18 in</quantity>
|
||
neuropodia and far posterior setiger
|
||
<quantity id="E45F04A9C467FFAF4DE1180DAEF324BD" box="[671,727,1388,1414]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="31" pageNumber="182" unit="in" value="32.0">32 in</quantity>
|
||
notopodia. Neuropodial spines typically numbering 1–2 at first, accompanied by 2–3 capillaries; in posterior segments notopodia with 1–2 spines and 1–2 capillaries, neuropodia with 2–3 spines and capillaries present or absent. Notopodial spines of far posterior segments longer than those in neuropodia, but both sets of spines emerging prominently from posteriormost setigers (
|
||
<figureCitation id="BB9CB5C9C467FFAF4B1318B9A8FA24C9" box="[1133,1246,1496,1522]" captionStart="FIGURE 19" captionStartId="30.[151,250,1973,1998]" captionTargetBox="[159,1401,734,1944]" captionTargetId="figure@30.[151,1409,720,1954]" captionTargetPageId="30" captionText="FIGURE 19. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary." httpUri="https://zenodo.org/record/3269029/files/figure.png" pageId="31" pageNumber="182">Figs. 19B</figureCitation>
|
||
,
|
||
<figureCitation id="BB9CB5C9C467FFAF4B9418B9A92324C9" box="[1258,1287,1496,1522]" captionStart="FIGURE 20" captionStartId="31.[151,250,957,982]" captionTargetBox="[155,1429,189,930]" captionTargetId="figure@31.[151,1436,181,938]" captionTargetPageId="31" captionText="FIGURE 20. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287):A, anterior end through setiger 14, right lateral view; B, far posterior segments, dorsal view; C, posterior 3 setigers and pygidium, dorsal view. Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269031/files/figure.png" pageId="31" pageNumber="182">20</figureCitation>
|
||
B–C). Spines curved, geniculate, terminating in stubby knob-tipped apex (
|
||
<figureCitation id="BB9CB5C9C467FFAF4C42189DAFAF272D" box="[828,907,1532,1558]" captionStart="FIGURE 19" captionStartId="30.[151,250,1973,1998]" captionTargetBox="[159,1401,734,1944]" captionTargetId="figure@30.[151,1409,720,1954]" captionTargetPageId="30" captionText="FIGURE 19. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary." httpUri="https://zenodo.org/record/3269029/files/figure.png" pageId="31" pageNumber="182">Fig. 19</figureCitation>
|
||
C–D); no denticles observed along shaft.
|
||
</paragraph>
|
||
<paragraph id="2318A94CC467FFAF4FB91B41A9542701" blockId="31.[151,1437,1064,2027]" box="[199,1392,1568,1595]" pageId="31" pageNumber="182">
|
||
Pygidium a thick rounded, ventrally directed lobe, surface with a coarse granular texture (
|
||
<figureCitation id="BB9CB5C9C467FFAF4BCB1B41A9022701" box="[1205,1318,1568,1594]" captionStart="FIGURE 19" captionStartId="30.[151,250,1973,1998]" captionTargetBox="[159,1401,734,1944]" captionTargetId="figure@30.[151,1409,720,1954]" captionTargetPageId="30" captionText="FIGURE 19. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287): A, anterior end, right lateral view; B, posterior end, dorsal view; C, neuroacicular spines; D, notoacicular spine and capillary." httpUri="https://zenodo.org/record/3269029/files/figure.png" pageId="31" pageNumber="182">Figs. 19B</figureCitation>
|
||
,
|
||
<figureCitation id="BB9CB5C9C467FFAF4A4F1B41A9402701" box="[1329,1380,1568,1594]" captionStart="FIGURE 20" captionStartId="31.[151,250,957,982]" captionTargetBox="[155,1429,189,930]" captionTargetId="figure@31.[151,1436,181,938]" captionTargetPageId="31" captionText="FIGURE 20. Tharyx hessleri n. sp. Holotype (LACM-AHF Poly 11287):A, anterior end through setiger 14, right lateral view; B, far posterior segments, dorsal view; C, posterior 3 setigers and pygidium, dorsal view. Stained with Shirlastain A." httpUri="https://zenodo.org/record/3269031/files/figure.png" pageId="31" pageNumber="182">20C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2318A94CC467FFAF4FB91B24AF232765" blockId="31.[151,1437,1064,2027]" box="[199,775,1603,1631]" pageId="31" pageNumber="182">
|
||
<emphasis id="11D3755EC467FFAF4FB91B24AD8B2765" bold="true" box="[199,431,1603,1631]" pageId="31" pageNumber="182">Methyl Green stain</emphasis>
|
||
. No pattern, de-stains entirely.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6BBDFAC7C467FFAF4FB91B08AD4527F1" pageId="31" pageNumber="182" type="etymology">
|
||
<paragraph id="2318A94CC467FFAF4FB91B08AD4527F1" blockId="31.[151,1437,1064,2027]" pageId="31" pageNumber="182">
|
||
<emphasis id="11D3755EC467FFAF4FB91B08AD6A27B8" bold="true" box="[199,334,1641,1667]" pageId="31" pageNumber="182">Etymology.</emphasis>
|
||
This species is named for Dr. Robert E. Hessler, prominent deep-sea ecologist whose pioneering work on deep-sea benthos, including collection of the
|
||
<taxonomicName id="E4A7D2CFC467FFAF4D871BEDAF61279D" box="[761,837,1676,1702]" class="Insecta" family="Calopterygidae" genus="Echo" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="186" phylum="Arthropoda" rank="genus">ECHO</taxonomicName>
|
||
I samples reported in this study, has inspired generations of scientists.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6BBDFAC7C467FFAF4FB91BB4ACDA26FD" pageId="31" pageNumber="182" type="discussion">
|
||
<paragraph id="2318A94CC467FFAF4FB91BB4ACDA26FD" blockId="31.[151,1437,1064,2027]" pageId="31" pageNumber="182">
|
||
<emphasis id="11D3755EC467FFAF4FB91BB4AD1027D4" bold="true" box="[199,308,1749,1775]" pageId="31" pageNumber="182">Remarks</emphasis>
|
||
. Among the 15 described species of
|
||
<taxonomicName id="E4A7D2CFC467FFAF4D9F1BB5AF1427D5" authorityName="Webster & Benedict" authorityYear="1887" box="[737,816,1748,1774]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="182" phylum="Annelida" rank="genus">
|
||
<emphasis id="11D3755EC467FFAF4D9F1BB5AF1427D5" box="[737,816,1748,1774]" italics="true" pageId="31" pageNumber="182">Tharyx</emphasis>
|
||
</taxonomicName>
|
||
, only two species have been recorded from depths of
|
||
<quantity id="E45F04A9C467FFAF4FE91B99ACC92628" box="[151,237,1784,1811]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="182" unit="m" value="2000.0">2000 m</quantity>
|
||
or greater:
|
||
<taxonomicName id="E4A7D2CFC467FFAF4E0E1B98AEB82629" authority="Blake, 1991" authorityName="Blake" authorityYear="1991" box="[368,668,1784,1810]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="182" phylum="Annelida" rank="species" species="kirkegaardi">
|
||
<emphasis id="11D3755EC467FFAF4E0E1B98AE292629" box="[368,525,1784,1810]" italics="true" pageId="31" pageNumber="182">T. kirkegaardi</emphasis>
|
||
<bibRefCitation id="4736D4BDC467FFAF4D6A1B99AEB82629" author="Blake, J. A." box="[532,668,1784,1810]" pageId="31" pageNumber="182" pagination="17 - 30" refId="ref19317" refString="Blake, J. A. (1991) Revision of some genera and species of Cirratulidae from the Western North Atlantic. Ophelia, Supplement No. 5, 17 - 30." type="book chapter" year="1991">Blake, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
from off the
|
||
<collectingCountry id="5BB0E9DCC467FFAF4C4D1B99AF402629" box="[819,868,1784,1810]" name="United States of America" pageId="31" pageNumber="182">U.S.</collectingCountry>
|
||
Atlantic coast in slope and abyssal depths to
|
||
<quantity id="E45F04A9C467FFAF4A1A1B99AC89260C" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="182" unit="m" value="3000.0">3000 m</quantity>
|
||
and
|
||
<taxonomicName id="E4A7D2CFC467FFAF4F9D1A7CADA9260D" authorityName="Blake" authorityYear="2018" box="[227,397,1820,1846]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="182" phylum="Annelida" rank="species" species="moniliformis">
|
||
<emphasis id="11D3755EC467FFAF4F9D1A7CADA9260D" box="[227,397,1820,1846]" italics="true" pageId="31" pageNumber="182">T. moniliformis</emphasis>
|
||
</taxonomicName>
|
||
from the Weddell Sea in
|
||
<quantity id="E45F04A9C467FFAF4DD91A7DAEDB260C" box="[679,767,1820,1847]" metricMagnitude="3" metricUnit="m" metricValue="2.086" pageId="31" pageNumber="182" unit="m" value="2086.0">2086 m</quantity>
|
||
.
|
||
<taxonomicName id="E4A7D2CFC467FFAF4C741A7DAF92260D" authority="Blake, 2019" authorityName="Blake" authorityYear="2019" box="[778,950,1820,1846]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="182" phylum="Annelida" rank="species" species="hessleri" status="sp. nov.">
|
||
<emphasis id="11D3755EC467FFAF4C741A7DAF92260D" box="[778,950,1820,1846]" italics="true" pageId="31" pageNumber="182">Tharyx hessleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0AE0C825C467FFAF4CC31A7CAFDF260C" box="[957,1019,1821,1847]" pageId="31" pageNumber="182" rank="species">
|
||
<emphasis id="11D3755EC467FFAF4CC31A7CAFDF260C" bold="true" box="[957,1019,1821,1847]" pageId="31" pageNumber="182">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from
|
||
<quantity id="E45F04A9C467FFAF4B411A7AA8B0260C" box="[1087,1172,1819,1847]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="182" unit="m" value="4500.0">4500 m</quantity>
|
||
is therefore the deepest recorded for any species of the genus. Both
|
||
<taxonomicName id="E4A7D2CFC467FFAF4DF41A20AF0D2661" authorityName="Blake" authorityYear="1991" box="[650,809,1856,1882]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="182" phylum="Annelida" rank="species" species="kirkegaardi">
|
||
<emphasis id="11D3755EC467FFAF4DF41A20AF0D2661" box="[650,809,1856,1882]" italics="true" pageId="31" pageNumber="182">T. kirkegaardi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E4A7D2CFC467FFAF4C1E1A20A8282661" authorityName="Blake" authorityYear="2018" box="[864,1036,1856,1882]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="182" phylum="Annelida" rank="species" species="moniliformis">
|
||
<emphasis id="11D3755EC467FFAF4C1E1A20A8282661" box="[864,1036,1856,1882]" italics="true" pageId="31" pageNumber="182">T. moniliformis</emphasis>
|
||
</taxonomicName>
|
||
have serrations along the shaft and prominent knobs on the tips of the acicular spines thus providing a sub-bidentate appearance; in contrast, the knobs on the spines of
|
||
<taxonomicName id="E4A7D2CFC467FFAF4E2A1AE8ADEC2699" authority="Blake, 2019" authorityName="Blake" authorityYear="2019" box="[340,456,1928,1954]" class="Polychaeta" family="Cirratulidae" genus="Tharyx" kingdom="Animalia" order="Terebellida" pageId="31" pageNumber="182" phylum="Annelida" rank="species" species="hessleri" status="sp. nov.">
|
||
<emphasis id="11D3755EC467FFAF4E2A1AE8ADEC2699" box="[340,456,1928,1954]" italics="true" pageId="31" pageNumber="182">T. hessleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0AE0C825C467FFAF4EAE1AE8AE342698" box="[464,528,1929,1955]" pageId="31" pageNumber="182" rank="species">
|
||
<emphasis id="11D3755EC467FFAF4EAE1AE8AE342698" bold="true" box="[464,528,1929,1955]" pageId="31" pageNumber="182">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are rounded off, providing a worn appearance and there are no serrations along the shaft.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6BBDFAC7C467FFAF4FB91AAEAF3726D0" box="[199,787,1999,2027]" pageId="31" pageNumber="182" type="distribution">
|
||
<paragraph id="2318A94CC467FFAF4FB91AAEAF3726D0" blockId="31.[151,1437,1064,2027]" box="[199,787,1999,2027]" pageId="31" pageNumber="182">
|
||
<emphasis id="11D3755EC467FFAF4FB91AAEAD7D26D1" bold="true" box="[199,345,1999,2026]" pageId="31" pageNumber="182">Distribution</emphasis>
|
||
. Abyssal Pacific Ocean,
|
||
<quantity id="E45F04A9C467FFAF4D0F1AAEAF2B26D0" box="[625,783,1999,2027]" metricMagnitude="3" metricUnit="m" metricValue="4.503" metricValueMax="4.506" metricValueMin="4.5" pageId="31" pageNumber="182" unit="m" value="4503.0" valueMax="4506.0" valueMin="4500.0">4500–4506 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |