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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.123.1448" ID-GBIF-Dataset="f1dbd205-6f2f-4feb-867e-d52041258d12" ID-PMC="PMC3175121" ID-Pensoft-Pub="1313-2970-123-1" ID-PubMed="21998529" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-123-1" ModsDocOrigin="ZooKeys 123" ModsDocTitle="Australian Assassins, Part I: A review of the Assassin Spiders (Araneae, Archaeidae) of mid-eastern Australia" checkinTime="1451249962699" checkinUser="pensoft" docAuthor="Rix, Michael G. &amp; Harvey, Mark S." docDate="2011" docId="A68D6B03381F860CF7A777CF7B27A2F5" docLanguage="en" docName="ZooKeys 123: 1-100" docOrigin="ZooKeys 123" docSource="http://dx.doi.org/10.3897/zookeys.123.1448" docTitle="Austrarchaea helenae Illawarra Assassin Spider Rix &amp; Harvey, sp. n." docType="treatment" docVersion="5" lastPageNumber="37" masterDocId="A428FFEAFD3A7353FFACFFBCFFFA6D47" masterDocTitle="Australian Assassins, Part I: A review of the Assassin Spiders (Araneae, Archaeidae) of mid-eastern Australia" masterLastPageNumber="100" masterPageNumber="1" pageNumber="36" updateTime="1668152002706" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Australian Assassins, Part I: A review of the Assassin Spiders (Araneae, Archaeidae) of mid-eastern Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rix, Michael G.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harvey, Mark S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>123</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>100</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.123.1448</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.123.1448</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-123-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152030494" LSID="urn:lsid:zoobank.org:act:278EE43B-C8F7-4069-9F60-4B343BB6FF6E" httpUri="http://treatment.plazi.org/id/A68D6B03381F860CF7A777CF7B27A2F5" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<subSubSection pageId="35" pageNumber="36" type="nomenclature">
<paragraph pageId="35" pageNumber="36">
<taxonomicName LSID="urn:lsid:zoobank.org:act:278EE43B-C8F7-4069-9F60-4B343BB6FF6E" authority="Rix &amp; Harvey" class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea helenae" name="Illawarra Assassin Spider" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="vernacular name" species="helenae" vernacular="vernacular">Austrarchaea helenae Illawarra Assassin Spider Rix &amp; Harvey</taxonomicName>
<taxonomicNameLabel pageId="35" pageNumber="36">sp. n.</taxonomicNameLabel>
Figs 9I2644
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="type material">
<paragraph pageId="35" pageNumber="36">Type material.</paragraph>
<paragraph pageId="35" pageNumber="36">
Holotype male: Macquarie Pass National Park, Macquarie Pass, New South Wales, Australia,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-34.566666">34°34'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="150.65">150°39'E</geoCoordinate>
, pitfall trap, 12-26.IX.1999, M. Gray, G. Milledge, H. Smith (AMS KS62774).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="other material examined">
<paragraph pageId="35" pageNumber="36">Other material examined.</paragraph>
<paragraph pageId="35" pageNumber="36">
AUSTRALIA: New South Wales: Macquarie Pass National Park: Macquarie Pass, off Clover Hill Road,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-34.568054">34°34'05&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="150.65694">150°39'25&quot;E</geoCoordinate>
, sifting elevated leaf litter, subtropical rainforest, 828 m, 8.IV.2010, M. Rix, D. Harms, 3 juveniles (WAM T112561DNA: Ar30-124-J/Ar30-125-J/Ar30-126-J).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="additional material examined (of tentative identification)">
<paragraph pageId="35" pageNumber="36">Additional material examined (of tentative identification).</paragraph>
<paragraph pageId="35" pageNumber="36">AUSTRALIA: New South Wales: Morton National Park: Barrengarry Mountain, ANIC Berlesate, rainforest, 460 m, 20.XII.1967, R. Taylor, C. Brooks, 1 juvenile (ANIC).</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="etymology">
<paragraph pageId="35" pageNumber="36">Etymology.</paragraph>
<paragraph pageId="35" pageNumber="36">The specific epithet is a patronym in honour of Helen Rix, for her love of the Illawarra Escarpment, and for her hospitality to the senior author during field work in eastern Australia.</paragraph>
</subSubSection>
<subSubSection lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="diagnosis">
<paragraph pageId="35" pageNumber="36">Diagnosis.</paragraph>
<paragraph pageId="35" pageNumber="36">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea helenae" order="Araneae" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="helenae">Austrarchaea helenae</taxonomicName>
can be distinguished from all other
<taxonomicName family="Archaeidae" lsidName="" pageId="35" pageNumber="36" rank="family">Archaeidae</taxonomicName>
from mid-eastern Australia by the long, spiniform tegular sclerite 1 (TS 1) with a curled distal tip (Fig. 26D) combined with the bifurcate, plate-like TS 3 (Fig. 26D).
</paragraph>
<paragraph lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
This species can also be distinguished from other genotyped taxa from mid-eastern Australia (see Fig. 3B) by the following 14 unique nucleotide substitutions for COI
<pageBreakToken pageId="36" pageNumber="37" start="start">and</pageBreakToken>
COII (n = 3): G(243), A(291), T(555), C(654), C(843), G(849), C(901), T(903), T(990), A(1206), C(1209), C(1401), C(1500), C(1548).
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="description">
<paragraph pageId="36" pageNumber="37">Description.</paragraph>
<paragraph pageId="36" pageNumber="37">
Holotype male: Total length 3.13; leg I femur 2.69; F1/CL ratio 2.50. Cephalothorax dark reddish-brown; legs tan-brown with darker annulations; abdomen mottled grey-brown and beige, with darker reddish-brown dorsal scute and sclerites (Fig. 26A). Carapace tall (CH/CL ratio 2.01); 1.08 long, 2.17 high, 1.02 wide;
<normalizedToken originalValue="neck">'neck'</normalizedToken>
0.53 wide; bearing two pairs of rudimentary horns; highest point of pars cephalica (HPC) near middle of
<normalizedToken originalValue="head">'head'</normalizedToken>
(ratio of HPC to post-ocular length 0.57), carapace gently sloping and almost horizontal posterior to HPC;
<normalizedToken originalValue="head">'head'</normalizedToken>
moderately elevated postero-dorsally (post-ocular ratio 0.35) (Fig. 9I). Chelicerae with short brush of accessory setae on anterior face of paturon (Fig. 26B). Abdomen 1.79 long, 1.28 wide; with three pairs of dorsal hump-like tubercles (HT 1-6); dorsal scute fused anteriorly to epigastric sclerites, extending posteriorly to first pair of hump-like tubercles; HT 3-6 each covered by separate dorsal sclerites. Expanded pedipalp (Figs 26C-D) with broadly-triangular, pointed conductor; tegular sclerite 1 (TS 1) long, spiniform, with curled distal tip, visible in retro-ventral view; TS 2 spiniform, shorter than TS 1, largely obscured by embolus and TS 3; TS 2a sinuous, filiform, exposed distally; TS 3 exposed, plate-like, overlying TS 2, with bifurcate, triangular apex directed toward proximal conductor.
</paragraph>
<paragraph pageId="36" pageNumber="37">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="distribution">
<paragraph pageId="36" pageNumber="37">Distribution and habitat.</paragraph>
<paragraph pageId="36" pageNumber="37">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea helenae" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="helenae">Austrarchaea helenae</taxonomicName>
is known only from rainforest habitats in the Macquarie Pass National Park, on the Illawarra Escarpment of south-eastern New South Wales (Fig. 44). A juvenile specimen from Barrengarry Mountain (Morton National Park) may also belong to this species based on proximity.
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="conservation status">
<paragraph pageId="36" pageNumber="37">Conservation status.</paragraph>
<paragraph pageId="36" pageNumber="37">
This species appears to be a rare short-range endemic taxon (
<bibRefCitation author="Harvey, MS" journalOrPublisher="Invertebrate Systematics" pageId="39" pageNumber="40" pagination="555 - 570" title="Short-range endemism among the Australian fauna: some examples from non-marine environments." url="doi:10.1071/IS02009" volume="16" year="2002 b">Harvey 2002b</bibRefCitation>
), with populations on the Illawarra Escarpment potentially threatened by land-clearing, habitat fragmentation and fire. Much of the original rainforest of the Illawarra region has been cleared for agriculture and livestock, and only isolated fragments of forest remain.
</paragraph>
</subSubSection>
</treatment>
</document>