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<mods:title id="11FAA55E082CD2356AB1E9A01AEAC340">Review of the brevicollis group of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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(
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, 
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: 
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, 136 [= 1872: 109, 110]; 
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, Jakobson 1910: 461
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<emphasis id="1EF4EAD6FFEE8B00FF60FEBCF181FED1" box="[151,445,317,341]" italics="true" pageId="12" pageNumber="593">Neogeodromicus transversus</emphasis>
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: 
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: 
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; 
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Khachikov 
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2021: 250
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<emphasis id="1EF4EAD6FFEE8B00FF60FE18F081FE35" box="[151,189,409,433]" italics="true" pageId="12" pageNumber="593">&lt;?&gt;</emphasis>
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aff. 
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<taxonomicName id="EB804D47FFEE8B00FF60FE39F264FE4B" authorityName="Motschulsky" authorityYear="1860" box="[151,600,440,464]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus" subGenus="Brachydromicus">
<emphasis id="1EF4EAD6FFEE8B00FF60FE39F123FE54" box="[151,287,440,464]" italics="true" pageId="12" pageNumber="593">Geodromicus</emphasis>
(
<emphasis id="1EF4EAD6FFEE8B00FED9FE39F1E9FE54" box="[302,469,440,464]" italics="true" pageId="12" pageNumber="593">Brachydromicus</emphasis>
) 
<emphasis id="1EF4EAD6FFEE8B00FE14FE39F264FE4B" box="[483,600,440,463]" italics="true" pageId="12" pageNumber="593">transversus</emphasis>
</taxonomicName>
: 
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<bibRefCitation id="48114B35FFEE8B00FD92FE39F314FE54" author="Khachikov, E. A." box="[613,808,440,464]" pageId="12" pageNumber="593" refId="ref16971" refString="Khachikov, E. A. (1998) Materialy k faune zhukov (Coleoptera) Nizhnego Dona i Severnogo Kavkaza. Zhuki-stafiliny. (Staphylinidae). Chast' II. Triba Staphylinini [Materials to the fauna of beetles (Coleoptera) of Lower Don and the North Caucasus. Rove beetles (Staphylinidae). Part II. The tribe Staphylinini]. Rostov Institute of Advanced Training and Retraining of Educators, Rostov-on-Don, 49 pp. [in Russian]" type="book" year="1998">Khachikov 1998: 8</bibRefCitation>
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</paragraph>
<paragraph id="2C3F36C4FFEE8B00FF60FE57F294FE6A" blockId="12.[151,1436,255,648]" box="[151,680,470,494]" pageId="12" pageNumber="593">
<treatmentCitationGroup id="0C9011EAFFEE8B00FF60FE57F294FE6A" box="[151,680,470,494]" pageId="12" pageNumber="593">
<treatmentCitation id="AD2110D5FFEE8B00FF60FE57F260FE6A" author="Motschulsky, V. de" box="[151,604,470,494]" page="551" pageId="12" pageNumber="593" year="1860">
<taxonomicName id="EB804D47FFEE8B00FF60FE57F260FE6A" authority="Motschulsky, 1860: 551" authorityName="Motschulsky" authorityPageNumber="551" authorityYear="1860" box="[151,604,470,494]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes" status="syn. nov.">
<emphasis id="1EF4EAD6FFEE8B00FF60FE57F15EFE6A" box="[151,354,470,494]" italics="true" pageId="12" pageNumber="593">Geodromus flavipes</emphasis>
<bibRefCitation id="48114B35FFEE8B00FE9FFE57F260FE6A" author="Motschulsky, V. de" box="[360,604,470,494]" pageId="12" pageNumber="593" pagination="539 - 588" refId="ref17233" refString="Motschulsky, V. de. (1860) Enumeration des nouvelles especes de coleopteres rapportees de ses voyages. 3 - ieme article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 33 (2), 539 - 588." type="journal article" year="1860">Motschulsky, 1860: 551</bibRefCitation>
</taxonomicName>
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<emphasis id="1EF4EAD6FFEE8B00FD95FE57F294FE6A" bold="true" box="[610,680,470,494]" pageId="12" pageNumber="593">
<taxonomicNameLabel id="05C757ADFFEE8B00FD95FE57F294FE6A" box="[610,680,470,494]" pageId="12" pageNumber="593">syn. n.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="2C3F36C4FFEE8B00FF60FE74F4ADFD89" blockId="12.[151,1436,255,648]" box="[151,1169,501,525]" pageId="12" pageNumber="593">
<treatmentCitationGroup id="0C9011EAFFEE8B00FF60FE74F4ADFD89" box="[151,1169,501,525]" pageId="12" pageNumber="593">
<taxonomicName id="EB804D47FFEE8B00FF60FE74F19EFD89" authorityName="Mannerheim" authorityYear="1830" box="[151,418,501,525]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="globulicollis">
<emphasis id="1EF4EAD6FFEE8B00FF60FE74F19EFD89" box="[151,418,501,525]" italics="true" pageId="12" pageNumber="593">Anthophagus globulicollis</emphasis>
</taxonomicName>
: 
<treatmentCitation id="AD2110D5FFEE8B00FE58FE74F2C2FD89" author="Gemminger, M. &amp; Harold, B. de" box="[431,766,501,525]" page="658" pageId="12" pageNumber="593" year="1868">
<bibRefCitation id="48114B35FFEE8B00FE58FE74F2C2FD89" author="Gemminger, M. &amp; Harold, B. de" box="[431,766,501,525]" pageId="12" pageNumber="593" pagination="425 - 752" refId="ref16410" refString="Gemminger, M. &amp; Harold, B. de. (1868) n. k. In: Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom II. Dytiscidae, Gyrinidae, Hydrophilidae, Staphylinidae, Pselaphidae, Gnostidae, Paussidae, Scydmaenidae, Silphidae, Trichopterygidae, Scaphidiidae. E. H. Gummi, Monachii, pp. 425 - 752 + (6). https: // doi. org / 10.5962 / bhl. title. 9089" type="journal article" year="1868">Gemminger &amp; Harold 1868: 658</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="AD2110D5FFEE8B00FCFEFE74F380FD89" author="Fauvel, A." box="[777,956,501,525]" page="493" pageId="12" pageNumber="593" year="1869">
<bibRefCitation id="48114B35FFEE8B00FCFEFE74F380FD89" author="Fauvel, A." box="[777,956,501,525]" pageId="12" pageNumber="593" pagination="479 - 494" refId="ref16207" refString="Fauvel, A. (1869) Remarques synonymiques sur les staphylinides du Catalogus Coleopterorum de MM v. Harold et Gemminger. L'Abeille. Memoires d'Entomologie, 5, 479 - 494." type="journal article" year="1869">Fauvel 1869: 493</bibRefCitation>
</treatmentCitation>
, Jakobson 1910: 461
</treatmentCitationGroup>
</paragraph>
<paragraph id="2C3F36C4FFEE8B00FF60FD95F258FDAF" blockId="12.[151,1436,255,648]" box="[151,612,531,556]" pageId="12" pageNumber="593">
<treatmentCitationGroup id="0C9011EAFFEE8B00FF60FD95F258FDAF" box="[151,612,531,556]" pageId="12" pageNumber="593">
<taxonomicName id="EB804D47FFEE8B00FF60FD95F1ABFDA8" box="[151,407,532,556]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFEE8B00FF60FD95F1ABFDA8" box="[151,407,532,556]" italics="true" pageId="12" pageNumber="593">Neogeodromicus flavipes</emphasis>
</taxonomicName>
: 
<treatmentCitation id="AD2110D5FFEE8B00FE53FD95F258FDAF" author="Bordoni, A." box="[420,612,531,556]" page="158" pageId="12" pageNumber="593" year="1993">
<bibRefCitation id="48114B35FFEE8B00FE53FD95F258FDAF" author="Bordoni, A." box="[420,612,531,556]" pageId="12" pageNumber="593" pagination="157 - 160" refId="ref15928" refString="Bordoni, A. (1993) Note intorno ad alcuni Omaliinae. Atti Museo Civico di Storia Naturale, Grosseto, 15, 157 - 160." type="journal article" year="1993">Bordoni 1993: 158</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="2C3F36C4FFEE8B00FF60FDB3F15AFDED" blockId="12.[151,1436,255,648]" pageId="12" pageNumber="593">
<treatmentCitationGroup id="0C9011EAFFEE8B00FF60FDB3F15AFDED" pageId="12" pageNumber="593">
<taxonomicName id="EB804D47FFEE8B00FF60FDB3F1B9FDCE" ID-CoL="58CBJ" baseAuthorityName="Motschulsky" baseAuthorityYear="1860" box="[151,389,562,586]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFEE8B00FF60FDB3F1B9FDCE" box="[151,389,562,586]" italics="true" pageId="12" pageNumber="593">Trichodromeus flavipes</emphasis>
</taxonomicName>
: 
<treatmentCitation id="AD2110D5FFEE8B00FE65FDB3F27BFDCE" author="Zerche, L." box="[402,583,562,586]" page="107" pageId="12" pageNumber="593" year="1992">
<bibRefCitation id="48114B35FFEE8B00FE65FDB3F27BFDCE" author="Zerche, L." box="[402,583,562,586]" pageId="12" pageNumber="593" pagination="105 - 142" refId="ref17790" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche 1992: 107</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="AD2110D5FFEE8B00FDA6FDB2F32DFDCE" author="Herman, L. H." box="[593,785,562,586]" page="373" pageId="12" pageNumber="593" year="2001">
<bibRefCitation id="48114B35FFEE8B00FDA6FDB2F32DFDCE" author="Herman, L. H." box="[593,785,562,586]" pageId="12" pageNumber="593" pagination="1 - 7" refId="ref16654" refString="Herman, L. H. (2001) Catalogue of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. Bulletin of the American Museum of Natural History, 265 (1 - 7), i - vi + 1 - 4218. https: // doi. org / 10.1206 / 0003 - 0090.265.1.1" type="journal article" year="2001">Herman 2001: 373</bibRefCitation>
</treatmentCitation>
, 
<treatmentCitation id="AD2110D5FFEE8B00FCEBFDB3F3DDFDCE" author="Smetana, A." box="[796,993,562,586]" page="251" pageId="12" pageNumber="593" year="2004">
<bibRefCitation id="48114B35FFEE8B00FCEBFDB3F3DDFDCE" author="Smetana, A." box="[796,993,562,586]" pageId="12" pageNumber="593" refId="ref17680" refString="Smetana, A. (2004) Staphylinidae, subfamilies Omaliinae - Dasycerinae, Phloeocharinae - Apateticinae, Piestinae - Staphylininae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. II. Hydrophiloidea - Histeroidea - Staphylinoidea. Appolo Books, Stenstrup, 942 pp." type="book" year="2004">Smetana 2004: 251</bibRefCitation>
</treatmentCitation>
, 
<treatmentCitation id="AD2110D5FFEE8B00FC1BFDB3F51FFDCE" author="Schulke, M. &amp; Smetana, A." box="[1004,1315,562,586]" page="326" pageId="12" pageNumber="593" year="2015">
<bibRefCitation id="48114B35FFEE8B00FC1BFDB3F51FFDCE" author="Schulke, M. &amp; Smetana, A." box="[1004,1315,562,586]" pageId="12" pageNumber="593" refId="ref17378" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134." type="book" year="2015">Schülke &amp; Smetana 2015: 326</bibRefCitation>
</treatmentCitation>
, 
<treatmentCitation id="AD2110D5FFEE8B00FADAFDB3F15AFDED" author="Khachikov, E. &amp; Iljina, E. &amp; Gasanova, N. &amp; Japoshvili, G." page="250" pageId="12" pageNumber="593" year="2021">
<bibRefCitation id="48114B35FFEE8B00FADAFDB3F15AFDED" author="Khachikov, E. &amp; Iljina, E. &amp; Gasanova, N. &amp; Japoshvili, G." pageId="12" pageNumber="593" pagination="248 - 266" refId="ref16901" refString="Khachikov, E., Iljina, E., Gasanova, N. &amp; Japoshvili, G. (2021) Preliminary faunistic study on the rove beetles (Coleoptera: Staphylinidae) of Republic of Dagestan, with some additional information from northern Caucasus. Zootaxa, 4950 (2), 248 - 266. https: // doi. org / 10.11646 / zootaxa. 4950.2.2" type="journal article" year="2021">
Khachikov 
<emphasis id="1EF4EAD6FFEE8B00FF30FDD3F0C6FDED" box="[199,250,593,617]" italics="true" pageId="12" pageNumber="593">et al.</emphasis>
2021: 250
</bibRefCitation>
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</treatmentCitationGroup>
</paragraph>
<paragraph id="2C3F36C4FFEE8B00FF60FDF1F33EFD0C" blockId="12.[151,1436,255,648]" box="[151,770,624,648]" pageId="12" pageNumber="593">
<treatmentCitationGroup id="0C9011EAFFEE8B00FF60FDF1F33EFD0C" box="[151,770,624,648]" pageId="12" pageNumber="593">
<taxonomicName id="EB804D47FFEE8B00FF60FDF1F20EFD0C" authorityName="Motschulsky" authorityYear="1860" box="[151,562,624,648]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes" subGenus="Brachydromicus">
<emphasis id="1EF4EAD6FFEE8B00FF60FDF1F123FD0C" box="[151,287,624,648]" italics="true" pageId="12" pageNumber="593">Geodromicus</emphasis>
(
<emphasis id="1EF4EAD6FFEE8B00FED9FDF1F1E9FD0C" box="[302,469,624,648]" italics="true" pageId="12" pageNumber="593">Brachydromicus</emphasis>
) 
<emphasis id="1EF4EAD6FFEE8B00FE14FDF1F20EFD0C" box="[483,562,624,648]" italics="true" pageId="12" pageNumber="593">flavipes</emphasis>
</taxonomicName>
: 
<treatmentCitation id="AD2110D5FFEE8B00FDC8FDF1F33EFD0C" author="Khachikov, E. A." box="[575,770,624,648]" page="8" pageId="12" pageNumber="593" year="1998">
<bibRefCitation id="48114B35FFEE8B00FDC8FDF1F33EFD0C" author="Khachikov, E. A." box="[575,770,624,648]" pageId="12" pageNumber="593" refId="ref16971" refString="Khachikov, E. A. (1998) Materialy k faune zhukov (Coleoptera) Nizhnego Dona i Severnogo Kavkaza. Zhuki-stafiliny. (Staphylinidae). Chast' II. Triba Staphylinini [Materials to the fauna of beetles (Coleoptera) of Lower Don and the North Caucasus. Rove beetles (Staphylinidae). Part II. The tribe Staphylinini]. Rostov Institute of Advanced Training and Retraining of Educators, Rostov-on-Don, 49 pp. [in Russian]" type="book" year="1998">Khachikov 1998: 8</bibRefCitation>
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</paragraph>
</subSubSection>
<subSubSection id="649A654FFFEE8B02FF60FD37F37CF8C9" lastPageId="14" lastPageNumber="595" pageId="12" pageNumber="593" type="materials_examined">
<paragraph id="2C3F36C4FFEE8B00FF60FD37F176FA40" blockId="12.[151,1437,693,2016]" pageId="12" pageNumber="593">
<emphasis id="1EF4EAD6FFEE8B00FF60FD37F180FD4B" bold="true" box="[151,444,693,720]" pageId="12" pageNumber="593">Type material examined.</emphasis>
<typeStatus id="F33B8866FFEE8B00FE35FD37F20EFD54" box="[450,562,694,720]" pageId="12" pageNumber="593" type="lectotype">Lectotype</typeStatus>
of 
<taxonomicName id="EB804D47FFEE8B00FDA3FD37F409FD54" authority="Motschulsky, 1860" authorityName="Motschulsky" authorityYear="1860" box="[596,1077,694,720]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFEE8B00FDA3FD37F360FD54" box="[596,860,694,720]" italics="true" pageId="12" pageNumber="593">Geodromus transversus</emphasis>
<bibRefCitation id="48114B35FFEE8B00FC95FD37F409FD54" author="Motschulsky, V. de" box="[866,1077,694,720]" pageId="12" pageNumber="593" pagination="539 - 588" refId="ref17233" refString="Motschulsky, V. de. (1860) Enumeration des nouvelles especes de coleopteres rapportees de ses voyages. 3 - ieme article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 33 (2), 539 - 588." type="journal article" year="1860">Motschulsky, 1860</bibRefCitation>
</taxonomicName>
<specimenCount id="3A86FD4DFFEE8B00FBCCFD37F46CFD54" box="[1083,1104,694,720]" count="1" pageId="12" pageNumber="593" type="male">♂</specimenCount>
(both antennomeres 2–11 and hind right leg missing; dissected prior to the present study; aedeagus in 
<collectingCountry id="54977654FFEE8B00FC5EFD5BF3C1FD70" box="[937,1021,730,756]" name="Canada" pageId="12" pageNumber="593">Canada</collectingCountry>
balsam was pinned under the beetle): ‘Alp. | Turchetia’ &lt;handwritten&gt;, ‘LECTOTYPUS’ &lt;pink printed label with black thin margins&gt;, [large white card with round plastic, without details of the body], ‘ 
<typeStatus id="F33B8866FFEE8B00FD59FCA3F30CFCB8" box="[686,816,802,828]" pageId="12" pageNumber="593" type="lectotype">Lectotypus</typeStatus>
<specimenCount id="3A86FD4DFFEE8B00FCC1FCA3F377FCB8" box="[822,843,802,828]" count="1" pageId="12" pageNumber="593" type="male">♂</specimenCount>
| 
<taxonomicName id="EB804D47FFEE8B00FCABFCA3F3CFFCB8" authorityName="Redtenbacher" authorityYear="1857" box="[860,1011,802,828]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FBF3FCA3F482FCB8" authority="Mot." authorityName="Motschulsky" authorityYear="1860" box="[1028,1214,802,828]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus Mot.</taxonomicName>
| Bordoni des.1989’ &lt;red handwritten label made by V.I. Gusarov&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FD59FCC7F34FFCE4" authorityName="Bordoni" authorityYear="1985" box="[686,883,838,864]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Neogeodromicus</taxonomicName>
[handwritten] | 
<taxonomicName id="EB804D47FFEE8B00FBD3FCC7F4CDFCE4" authority="(Mot.)" baseAuthorityName="Mot." box="[1060,1265,838,864]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus (Mot.)</taxonomicName>
[handwritten] | det. Bordoni 1989’ &lt;printed&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FE1BFCEBF2B4FC00" authorityName="Redtenbacher" authorityYear="1857" box="[492,648,874,900]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FD6EFCEBF344FC00" authority="Motsch." authorityName="Motschulsky" authorityYear="1860" box="[665,888,874,900]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus Motsch.</taxonomicName>
| V.I. Gusarov det. 199[printed]4’ &lt;handwritten&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FF56FC0FF10BFC2C" authorityName="Redtenbacher" authorityYear="1857" box="[161,311,910,936]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="1EF4EAD6FFEE8B00FF56FC0FF10BFC2C" box="[161,311,910,936]" italics="true" pageId="12" pageNumber="593">Geodromicus</emphasis>
</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FEA7FC0EF1EDFC2C" authorityName="Motschulsky" authorityYear="1860" box="[336,465,911,936]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFEE8B00FEA7FC0EF1EDFC2C" box="[336,465,911,936]" italics="true" pageId="12" pageNumber="593">transversus</emphasis>
</taxonomicName>
| 
<bibRefCitation id="48114B35FFEE8B00FE1BFC0FF2F8FC2C" author="Motschulsky, V. de" box="[492,708,910,936]" pageId="12" pageNumber="593" pagination="539 - 588" refId="ref17233" refString="Motschulsky, V. de. (1860) Enumeration des nouvelles especes de coleopteres rapportees de ses voyages. 3 - ieme article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 33 (2), 539 - 588." type="journal article" year="1860">Motschulsky, 1860</bibRefCitation>
| Shavrin A.V. det. 2014’ &lt;printed&gt; (
<collectionCode id="4A91AE01FFEE8B00FB8FFC0FF4FDFC2C" box="[1144,1217,910,936]" pageId="12" pageNumber="593">ZMM</collectionCode>
). 
<typeStatus id="F33B8866FFEE8B00FB20FC0FF546FC2C" box="[1239,1402,910,936]" pageId="12" pageNumber="593" type="paralectotype">Paralectotypes</typeStatus>
of 
<taxonomicName id="EB804D47FFEE8B00FF60FC33F294FC48" authority="Motschulsky, 1860: 1" authorityName="Motschulsky" authorityPageNumber="1" authorityYear="1860" box="[151,680,946,972]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFEE8B00FF60FC33F198FC48" box="[151,420,946,972]" italics="true" pageId="12" pageNumber="593">Geodromus transversus</emphasis>
<bibRefCitation id="48114B35FFEE8B00FE58FC33F294FC48" author="Motschulsky, V. de" box="[431,680,946,972]" pageId="12" pageNumber="593" pagination="539 - 588" refId="ref17233" refString="Motschulsky, V. de. (1860) Enumeration des nouvelles especes de coleopteres rapportees de ses voyages. 3 - ieme article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 33 (2), 539 - 588." type="journal article" year="1860">
Motschulsky, 1860: 
<specimenCount id="3A86FD4DFFEE8B00FD6DFC33F294FC48" box="[666,680,946,972]" count="1" pageId="12" pageNumber="593" type="male">1</specimenCount>
</bibRefCitation>
</taxonomicName>
♂ (head and pronotum missing), 
<specimenCount id="3A86FD4DFFEE8B00FBC0FC33F445FC48" box="[1079,1145,946,972]" count="2" pageId="12" pageNumber="593" type="female">2 ♀♀</specimenCount>
(
<specimenCount id="3A86FD4DFFEE8B00FB7BFC33F530FC48" box="[1164,1292,946,972]" count="1" pageId="12" pageNumber="593" type="female">one female</specimenCount>
without left middle leg): ‘Alp. Turchet’ &lt;pink label, handwritten in black Indian ink&gt;, ‘transversus’ &lt;handwritten in black Indian ink&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FF2BFC7BF158FB90" box="[220,356,1018,1044]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FE81FC7BF27BFB90" authority="Motsc." authorityName="Motschulsky" authorityYear="1860" box="[374,583,1018,1044]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus Motsc.</taxonomicName>
| Alp. Cauc.Turchetia’ &lt;large label, with black margins, handwritten in black Indian ink&gt; ‘ 
<typeStatus id="F33B8866FFEE8B00FED3FB9FF1F2FBBC" box="[292,462,1054,1080]" pageId="12" pageNumber="593" type="paralectotype">Paralectotypus</typeStatus>
| 
<taxonomicName id="EB804D47FFEE8B00FE16FB9FF244FBBC" authorityName="Redtenbacher" authorityYear="1857" box="[481,632,1054,1080]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FD7DFB9FF379FBBC" authority="Mot." authorityName="Motschulsky" authorityYear="1860" box="[650,837,1054,1080]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus Mot.</taxonomicName>
| Bordoni des. 1989’ &lt;red handwritten label made by V.I. Gusarov&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FEBEFBC3F232FBD8" authorityName="Bordoni" authorityYear="1985" box="[329,526,1090,1116]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Neogeodromicus</taxonomicName>
[handwritten] | 
<taxonomicName id="EB804D47FFEE8B00FD33FBC3F3AFFBD8" authority="(Mot.)" baseAuthorityName="Mot." box="[708,915,1090,1116]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus (Mot.)</taxonomicName>
[handwritten] | det. Bordoni 1989’ &lt;printed&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FF6CFBE7F10BFB04" authorityName="Redtenbacher" authorityYear="1857" box="[155,311,1126,1152]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FEBBFBE7F212FB04" authority="Motsch." authorityName="Motschulsky" authorityYear="1860" box="[332,558,1126,1152]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus Motsch.</taxonomicName>
| V.I. Gusarov det. 199[printed]4’ &lt;handwritten&gt; (
<collectionCode id="4A91AE01FFEE8B00FB91FBE7F493FB04" box="[1126,1199,1126,1152]" pageId="12" pageNumber="593">ZMM</collectionCode>
); 
<specimenCount id="3A86FD4DFFEE8B00FB34FBE7F4CCFB04" box="[1219,1264,1126,1152]" count="1" pageId="12" pageNumber="593" type="female">1 ♀</specimenCount>
: ‘Alp. Turchet’ &lt;pink label, handwritten in black Indian ink&gt;, ‘ 
<typeStatus id="F33B8866FFEE8B00FD6BFB0BF37AFB20" box="[668,838,1162,1188]" pageId="12" pageNumber="593" type="paralectotype">Paralectotypus</typeStatus>
| 
<taxonomicName id="EB804D47FFEE8B00FCA0FB0BF3D2FB20" authorityName="Redtenbacher" authorityYear="1857" box="[855,1006,1162,1188]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FC08FB0BF485FB20" authority="Mot." authorityName="Motschulsky" authorityYear="1860" box="[1023,1209,1162,1188]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus Mot.</taxonomicName>
| Bordoni des. 1989’ &lt;red handwritten label made by V.I. Gusarov&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FD59FB2FF34FFB4C" authorityName="Bordoni" authorityYear="1985" box="[686,883,1198,1224]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Neogeodromicus</taxonomicName>
[handwritten] | 
<taxonomicName id="EB804D47FFEE8B00FBD3FB2FF4CDFB4C" authority="(Mot.)" baseAuthorityName="Mot." box="[1060,1265,1198,1224]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus (Mot.)</taxonomicName>
[handwritten] | det. Bordoni 1989’ &lt;printed&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FE19FB53F2B6FB68" authorityName="Redtenbacher" authorityYear="1857" box="[494,650,1234,1260]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FD6BFB53F340FB68" authority="Motsch." authorityName="Motschulsky" authorityYear="1860" box="[668,892,1234,1260]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus Motsch.</taxonomicName>
| V.I. Gusarov det. 199[printed]4’ &lt;handwritten&gt; (
<collectionCode id="4A91AE01FFEE8B00FF68FB77F0D4FA94" box="[159,232,1270,1296]" pageId="12" pageNumber="593">ZMM</collectionCode>
); 
<specimenCount id="3A86FD4DFFEE8B00FF0AFB77F114FA94" box="[253,296,1270,1296]" count="1" pageId="12" pageNumber="593" type="female">1 ♀</specimenCount>
(left antenna and right antennomeres 7–11 missing): 
<taxonomicName id="EB804D47FFEE8B00FC7EFB77F42CFA94" authorityName="Motschulsky" authorityYear="1860" box="[905,1040,1270,1296]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">‘transversus</taxonomicName>
| m.’ &lt;handwritten in black Indian ink&gt;, ‘246?’ &lt;handwritten in black Indian ink&gt;, ‘ 
<typeStatus id="F33B8866FFEE8B00FD50FA9BF36DFAB0" box="[679,849,1306,1332]" pageId="12" pageNumber="593" type="paralectotype">Paralectotypus</typeStatus>
| 
<taxonomicName id="EB804D47FFEE8B00FCA8FA9BF3CAFAB0" authorityName="Redtenbacher" authorityYear="1857" box="[863,1014,1306,1332]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FBF2FA9BF482FAB0" authority="Mot." authorityName="Motschulsky" authorityYear="1860" box="[1029,1214,1306,1332]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus Mot.</taxonomicName>
| Bordoni des. 1989’ &lt;red handwritten label made by V.I. Gusarov&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FD59FABFF34FFADC" authorityName="Bordoni" authorityYear="1985" box="[686,883,1342,1368]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Neogeodromicus</taxonomicName>
[handwritten] | 
<taxonomicName id="EB804D47FFEE8B00FBD3FABFF4CDFADC" authority="(Mot.)" baseAuthorityName="Mot." box="[1060,1265,1342,1368]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus (Mot.)</taxonomicName>
[handwritten] | det. Bordoni 1989’ &lt;printed&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FE19FAE3F2B6FAF8" authorityName="Redtenbacher" authorityYear="1857" box="[494,650,1378,1404]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FD6BFAE3F340FAF8" authority="Motsch." authorityName="Motschulsky" authorityYear="1860" box="[668,892,1378,1404]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus Motsch.</taxonomicName>
| V.I. Gusarov det. 199[printed]4’ &lt;handwritten&gt; (
<collectionCode id="4A91AE01FFEE8B00FF68FA07F0D4FA24" box="[159,232,1414,1440]" pageId="12" pageNumber="593">ZMM</collectionCode>
). All 
<typeStatus id="F33B8866FFEE8B00FED3FA07F1FAFA24" box="[292,454,1414,1440]" pageId="12" pageNumber="593" type="paralectotype">paralectotypes</typeStatus>
with additional printed label: ‘ 
<taxonomicName id="EB804D47FFEE8B00FCD7FA07F38AFA24" authorityName="Redtenbacher" authorityYear="1857" box="[800,950,1414,1440]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="1EF4EAD6FFEE8B00FCD7FA07F38AFA24" box="[800,950,1414,1440]" italics="true" pageId="12" pageNumber="593">Geodromicus</emphasis>
</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FC3EFA06F476FA24" authorityName="Motschulsky" authorityYear="1860" box="[969,1098,1415,1440]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFEE8B00FC3EFA06F476FA24" box="[969,1098,1415,1440]" italics="true" pageId="12" pageNumber="593">transversus</emphasis>
</taxonomicName>
| 
<bibRefCitation id="48114B35FFEE8B00FBAAFA07F50EFA24" author="Motschulsky, V. de" box="[1117,1330,1414,1440]" pageId="12" pageNumber="593" pagination="539 - 588" refId="ref17233" refString="Motschulsky, V. de. (1860) Enumeration des nouvelles especes de coleopteres rapportees de ses voyages. 3 - ieme article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 33 (2), 539 - 588." type="journal article" year="1860">Motschulsky, 1860</bibRefCitation>
| Shavrin A.V. det. 2014’.
</paragraph>
<paragraph id="2C3F36C4FFEE8B00FF30FA4FF390F944" blockId="12.[151,1437,693,2016]" pageId="12" pageNumber="593">
<typeStatus id="F33B8866FFEE8B00FF30FA4FF10BFA6C" box="[199,311,1486,1512]" pageId="12" pageNumber="593" type="lectotype">Lectotype</typeStatus>
of 
<taxonomicName id="EB804D47FFEE8B00FE91FA4FF313FA6C" authority="Motschulsky, 1860" authorityName="Motschulsky" authorityYear="1860" box="[358,815,1486,1512]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFEE8B00FE91FA4FF276FA6C" box="[358,586,1486,1512]" italics="true" pageId="12" pageNumber="593">Geodromus flavipes</emphasis>
<bibRefCitation id="48114B35FFEE8B00FDA2FA4FF313FA6C" author="Motschulsky, V. de" box="[597,815,1486,1512]" pageId="12" pageNumber="593" pagination="539 - 588" refId="ref17233" refString="Motschulsky, V. de. (1860) Enumeration des nouvelles especes de coleopteres rapportees de ses voyages. 3 - ieme article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 33 (2), 539 - 588." type="journal article" year="1860">Motschulsky, 1860</bibRefCitation>
</taxonomicName>
<specimenCount id="3A86FD4DFFEE8B00FCCCFA4FF36CFA6C" box="[827,848,1486,1512]" count="1" pageId="12" pageNumber="593" type="male">♂</specimenCount>
(dissected prior to the present study; aedeagus in 
<collectingCountry id="54977654FFEE8B00FF60FA73F0D7F988" box="[151,235,1522,1548]" name="Canada" pageId="12" pageNumber="593">Canada</collectingCountry>
balsam was pinned under the beetle): ‘Alp. Cauc.’ &lt;handwritten&gt;, ‘LECTOTYPUS’ &lt;pink printed label with black thin margins&gt;, [large white card with round plastic, without details of the body], [large white card with round plastic, without details of the body], ‘ 
<typeStatus id="F33B8866FFEE8B00FD37F9BBF37EF9D0" box="[704,834,1594,1620]" pageId="12" pageNumber="593" type="lectotype">Lectotypus</typeStatus>
<specimenCount id="3A86FD4DFFEE8B00FCBEF9BBF362F9D0" box="[841,862,1594,1620]" count="1" pageId="12" pageNumber="593" type="male">♂</specimenCount>
| 
<taxonomicName id="EB804D47FFEE8B00FC83F9BBF437F9D0" authorityName="Redtenbacher" authorityYear="1857" box="[884,1035,1594,1620]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FBD6F9BBF484F9D0" authority="Mot." authorityName="Motschulsky" authorityYear="1860" box="[1057,1208,1594,1620]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes">flavipes Mot.</taxonomicName>
| Bordoni des.1989’ &lt;red handwritten label made by V.I. Gusarov&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FD35F9DFF3BBF9FC" authorityName="Bordoni" authorityYear="1985" box="[706,903,1630,1656]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Neogeodromicus</taxonomicName>
[handwritten] | 
<taxonomicName id="EB804D47FFEE8B00FBB6F9DFF4D7F9FC" authority="(Mot.)" baseAuthorityName="Mot." box="[1089,1259,1630,1656]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes">flavipes (Mot.)</taxonomicName>
[handwritten] | det. Bordoni 1989’ &lt;printed&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FE0FF903F2A8F918" authorityName="Redtenbacher" authorityYear="1857" box="[504,660,1666,1692]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FD5DF903F35BF918" authority="Motsch." authorityName="Motschulsky" authorityYear="1860" box="[682,871,1666,1692]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes">flavipes Motsch.</taxonomicName>
| V.I. Gusarov det. 199[printed]4’ &lt;handwritten&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FF56F927F10BF944" authorityName="Redtenbacher" authorityYear="1857" box="[161,311,1702,1728]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="1EF4EAD6FFEE8B00FF56F927F10BF944" box="[161,311,1702,1728]" italics="true" pageId="12" pageNumber="593">Geodromicus</emphasis>
</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FEBDF927F19DF944" authorityName="Motschulsky" authorityYear="1860" box="[330,417,1702,1728]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFEE8B00FEBDF927F19DF944" box="[330,417,1702,1728]" italics="true" pageId="12" pageNumber="593">flavipes</emphasis>
</taxonomicName>
| 
<bibRefCitation id="48114B35FFEE8B00FE42F927F2B5F944" author="Motschulsky, V. de" box="[437,649,1702,1728]" pageId="12" pageNumber="593" pagination="539 - 588" refId="ref17233" refString="Motschulsky, V. de. (1860) Enumeration des nouvelles especes de coleopteres rapportees de ses voyages. 3 - ieme article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 33 (2), 539 - 588." type="journal article" year="1860">Motschulsky, 1860</bibRefCitation>
| Shavrin A.V. det. 2014’.
</paragraph>
<paragraph id="2C3F36C4FFEE8B00FF30F94BF2FFF864" blockId="12.[151,1437,693,2016]" pageId="12" pageNumber="593">
<typeStatus id="F33B8866FFEE8B00FF30F94BF156F960" box="[199,362,1738,1764]" pageId="12" pageNumber="593" type="paralectotype">Paralectotypes</typeStatus>
of 
<taxonomicName id="EB804D47FFEE8B00FE65F94BF352F960" authority="Motschulsky, 1860: 1" authorityName="Motschulsky" authorityPageNumber="1" authorityYear="1860" box="[402,878,1738,1764]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFEE8B00FE65F94BF24EF960" box="[402,626,1738,1764]" italics="true" pageId="12" pageNumber="593">Geodromus flavipes</emphasis>
<bibRefCitation id="48114B35FFEE8B00FD8DF94BF352F960" author="Motschulsky, V. de" box="[634,878,1738,1764]" pageId="12" pageNumber="593" pagination="539 - 588" refId="ref17233" refString="Motschulsky, V. de. (1860) Enumeration des nouvelles especes de coleopteres rapportees de ses voyages. 3 - ieme article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 33 (2), 539 - 588." type="journal article" year="1860">
Motschulsky, 1860: 
<specimenCount id="3A86FD4DFFEE8B00FC97F94BF352F960" box="[864,878,1738,1764]" count="1" pageId="12" pageNumber="593" type="male">1</specimenCount>
</bibRefCitation>
</taxonomicName>
♂ (dissected prior to the present study; aedeagus and apical abdominal segments in 
<collectingCountry id="54977654FFEE8B00FDE7F96FF258F88C" box="[528,612,1774,1800]" name="Canada" pageId="12" pageNumber="593">Canada</collectingCountry>
balsam was pinned under the beetle), 
<specimenCount id="3A86FD4DFFEE8B00FBF3F96FF412F88C" box="[1028,1070,1774,1800]" count="1" pageId="12" pageNumber="593" type="female">1 ♀</specimenCount>
: [empty long original transparent card], ‘Alp. Cauc’ &lt;pink label, handwritten in black Indian ink&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FC64F893F427F8A8" box="[915,1051,1810,1836]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromus</taxonomicName>
| flavipes | Motsch. | Alp. Cauc.’ &lt;handwritten in black Indian ink&gt;, ‘ 
<typeStatus id="F33B8866FFEE8B00FDCCF8B7F2D9F8D4" box="[571,741,1846,1872]" pageId="12" pageNumber="593" type="paralectotype">Paralectotypus</typeStatus>
<specimenCount id="3A86FD4DFFEE8B00FD07F8B7F339F8D4" box="[752,773,1846,1872]" count="1" pageId="12" pageNumber="593" type="male">♂</specimenCount>
| 
<taxonomicName id="EB804D47FFEE8B00FCD7F8B7F38BF8D4" authorityName="Redtenbacher" authorityYear="1857" box="[800,951,1846,1872]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FC25F8B7F457F8D4" authority="Mot." authorityName="Motschulsky" authorityYear="1860" box="[978,1131,1846,1872]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes">flavipes Mot.</taxonomicName>
| Bordoni des.1989’ &lt;two similar red labels; handwritten by V.I. Gusarov&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FD49F8DBF3BFF8F0" authorityName="Bordoni" authorityYear="1985" box="[702,899,1882,1908]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Neogeodromicus</taxonomicName>
[handwritten] | 
<taxonomicName id="EB804D47FFEE8B00FBC5F8DBF4C2F8F0" authority="(Mot.)" baseAuthorityName="Mot." box="[1074,1278,1882,1908]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">transversus (Mot.)</taxonomicName>
[handwritten] | det. Bordoni 1989’ &lt;two similar labels, printed&gt;, ‘ 
<taxonomicName id="EB804D47FFEE8B00FD31F8FFF35EF81C" authorityName="Redtenbacher" authorityYear="1857" box="[710,866,1918,1944]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FC85F8FFF410F81C" authority="Motsch." authorityName="Motschulsky" authorityYear="1860" box="[882,1068,1918,1944]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="flavipes">flavipes Motsch.</taxonomicName>
| V.I. Gusarov det. 199[printed]4’ &lt;handwritten&gt; (
<collectionCode id="4A91AE01FFEE8B00FEBCF823F1A8F838" box="[331,404,1954,1980]" pageId="12" pageNumber="593">ZMM</collectionCode>
). All 
<typeStatus id="F33B8866FFEE8B00FE38F823F24DF838" box="[463,625,1954,1980]" pageId="12" pageNumber="593" type="paralectotype">paralectotypes</typeStatus>
with additional printed label: ‘ 
<taxonomicName id="EB804D47FFEE8B00FC3DF823F45CF838" authorityName="Redtenbacher" authorityYear="1857" box="[970,1120,1954,1980]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="1EF4EAD6FFEE8B00FC3DF823F45CF838" box="[970,1120,1954,1980]" italics="true" pageId="12" pageNumber="593">Geodromicus</emphasis>
</taxonomicName>
| 
<taxonomicName id="EB804D47FFEE8B00FB84F822F4C8F838" box="[1139,1268,1955,1980]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFEE8B00FB84F822F4C8F838" box="[1139,1268,1955,1980]" italics="true" pageId="12" pageNumber="593">transversus</emphasis>
</taxonomicName>
| 
<bibRefCitation id="48114B35FFEE8B00FAF1F823F0F3F864" author="Motschulsky, V. de" pageId="12" pageNumber="593" pagination="539 - 588" refId="ref17233" refString="Motschulsky, V. de. (1860) Enumeration des nouvelles especes de coleopteres rapportees de ses voyages. 3 - ieme article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 33 (2), 539 - 588." type="journal article" year="1860">Motschulsky, 1860</bibRefCitation>
| Shavrin A.V. det. 2014’ &lt;printed&gt; (
<collectionCode id="4A91AE01FFEE8B00FD9AF847F28AF864" box="[621,694,1990,2016]" pageId="12" pageNumber="593">ZMM</collectionCode>
).
</paragraph>
<caption id="78FF664CFFEF8B01FF60F9FEF49AF95B" ID-DOI="http://doi.org/10.5281/zenodo.12189114" ID-Zenodo-Dep="12189114" httpUri="https://zenodo.org/record/12189114/files/figure.png" pageId="13" pageNumber="594" startId="13.[151,264,1663,1687]" targetBox="[159,1424,190,1630]" targetPageId="13" targetType="figure">
<paragraph id="2C3F36C4FFEF8B01FF60F9FEF49AF95B" blockId="13.[151,1437,1663,1759]" pageId="13" pageNumber="594">
<emphasis id="1EF4EAD6FFEF8B01FF60F9FEF16BF913" bold="true" box="[151,343,1663,1687]" pageId="13" pageNumber="594">FIGURES 16–23.</emphasis>
Aedeagus of 
<taxonomicName id="EB804D47FFEF8B01FE1DF9FEF2D1F913" authorityName="Shavrin" authorityYear="2024" box="[490,749,1663,1687]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="species" species="alborzensis">
<emphasis id="1EF4EAD6FFEF8B01FE1DF9FEF2D1F913" box="[490,749,1663,1687]" italics="true" pageId="13" pageNumber="594">Geodromicus alborzensis</emphasis>
</taxonomicName>
(Figs. 16–17: holotype) and 
<taxonomicName id="EB804D47FFEF8B01FBD6F901F48BF913" box="[1057,1207,1664,1687]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFEF8B01FBD6F901F48BF913" box="[1057,1207,1664,1687]" italics="true" pageId="13" pageNumber="594">G. transversus</emphasis>
</taxonomicName>
(Figs. 18–19: Russia, Krasnaya Polyana; Fig. 20: lectotype of 
<taxonomicName id="EB804D47FFEF8B01FDCAF925F2EFF93F" box="[573,723,1700,1723]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFEF8B01FDCAF925F273F93F" box="[573,591,1700,1723]" italics="true" pageId="13" pageNumber="594">G</emphasis>
. 
<emphasis id="1EF4EAD6FFEF8B01FDA9F925F2EFF93F" box="[606,723,1700,1723]" italics="true" pageId="13" pageNumber="594">transversus</emphasis>
</taxonomicName>
; Fig. 21: lectotype of 
<taxonomicName id="EB804D47FFEF8B01FC49F925F413F93F" authorityName="Motschulsky" authorityYear="1860" box="[958,1071,1699,1723]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFEF8B01FC49F925F413F93F" box="[958,1071,1699,1723]" italics="true" pageId="13" pageNumber="594">G. flavipes</emphasis>
</taxonomicName>
; Fig. 22–23: Karachay-Cherkessia, Epchik Pass): 16, 18–22—parameral view, 17, 19—lateral view, 23—internal sac. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="2C3F36C4FFEF8B01FF30F8A8F0C1F873" blockId="13.[151,1437,1833,2039]" pageId="13" pageNumber="594">
[
<specimenCount id="3A86FD4DFFEF8B01FF39F8A8F168F8C7" box="[206,340,1833,1859]" count="1" pageId="13" pageNumber="594" type="female">One female</specimenCount>
of 
<taxonomicName id="EB804D47FFEF8B01FE89F8ABF1C5F8C7" authorityName="Motschulsky" authorityYear="1860" box="[382,505,1833,1860]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFEF8B01FE89F8ABF1C5F8C7" box="[382,505,1833,1860]" italics="true" pageId="13" pageNumber="594">G. flavipes</emphasis>
</taxonomicName>
was labeled as 
<typeStatus id="F33B8866FFEF8B01FD45F8A8F370F8C7" box="[690,844,1833,1859]" pageId="13" pageNumber="594" type="paralectotype">paralectotype</typeStatus>
, but this specimen belongs to the genus 
<taxonomicName id="EB804D47FFEF8B01FAE8F8ABF549F8C7" box="[1311,1397,1834,1859]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="genus">
<emphasis id="1EF4EAD6FFEF8B01FAE8F8ABF549F8C7" box="[1311,1397,1834,1859]" italics="true" pageId="13" pageNumber="594">Lesteva</emphasis>
</taxonomicName>
: 
<emphasis id="1EF4EAD6FFEF8B01FA71F8ABF5AAF8C7" box="[1414,1430,1834,1859]" italics="true" pageId="13" pageNumber="594">L</emphasis>
. (s.str.) sp. Labels of this specimen are: ‘Daghestan’ &lt;pink, handwritten&gt;, ‘rufipes’ &lt;handwritten in black Indian ink&gt;, ‘248.’ &lt;handwritten in black Indian ink&gt;,‘? 
<specimenCount id="3A86FD4DFFEF8B01FD23F8F0F2D5F80F" box="[724,745,1905,1931]" count="1" pageId="13" pageNumber="594" type="female">♀</specimenCount>
’ &lt;handwritten&gt;, ‘ 
<typeStatus id="F33B8866FFEF8B01FC43F8F0F462F80F" box="[948,1118,1905,1931]" pageId="13" pageNumber="594" type="paralectotype">Paralectotypus</typeStatus>
<specimenCount id="3A86FD4DFFEF8B01FB90F8F3F440F808" box="[1127,1148,1906,1932]" count="1" pageId="13" pageNumber="594" type="male">♂</specimenCount>
| 
<taxonomicName id="EB804D47FFEF8B01FB63F8F0F517F80F" authorityName="Redtenbacher" authorityYear="1857" box="[1172,1323,1905,1931]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="genus">Geodromicus</taxonomicName>
| 
<taxonomicName id="EB804D47FFEF8B01FAB3F8F0F0F1F834" authority="Mot." authorityName="Motschulsky" authorityYear="1860" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="species" species="flavipes">flavipes Mot.</taxonomicName>
| Bordoni des.1989’ &lt;red label handwritten by V.I. Gusarov&gt;, ‘ 
<taxonomicName id="EB804D47FFEF8B01FC56F814F45AF82B" authorityName="Bordoni" authorityYear="1985" box="[929,1126,1941,1967]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="genus">Neogeodromicus</taxonomicName>
[handwritten] | 
<taxonomicName id="EB804D47FFEF8B01FAE9F817F0E3F857" authority="(Mot.)" baseAuthorityName="Mot." class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="species" species="transversus">transversus (Mot.)</taxonomicName>
[handwritten] | det. Bordoni 1989’ &lt;printed&gt;, ‘ 
<taxonomicName id="EB804D47FFEF8B01FCFEF838F359F857" box="[777,869,1977,2003]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="genus">Lesteva</taxonomicName>
<specimenCount id="3A86FD4DFFEF8B01FC87F83BF3B9F850" box="[880,901,1978,2004]" count="1" pageId="13" pageNumber="594" type="female">♀</specimenCount>
| sp. | V.I. Gusarov det. 1994’ &lt;handwritten&gt; (
<collectionCode id="4A91AE01FFEF8B01FF68F85CF0D6F873" box="[159,234,2013,2039]" pageId="13" pageNumber="594">ZMM</collectionCode>
).]
</paragraph>
<paragraph id="2C3F36C4FFEC8B02FF30FF17F37CF8C9" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="595">
<emphasis id="1EF4EAD6FFEC8B02FF30FF17F3EEFF34" bold="true" box="[199,978,150,176]" pageId="14" pageNumber="595">
Additional material examined. 
<collectingCountry id="54977654FFEC8B02FDCEFF17F298FF34" box="[569,676,150,176]" name="Russia" pageId="14" pageNumber="595">RUSSIA</collectingCountry>
: 
<collectingRegion id="EE44F826FFEC8B02FD58FF17F37CFF34" box="[687,832,150,176]" country="Russia" name="Krasnodar" pageId="14" pageNumber="595">KRASNODAR</collectingRegion>
TERRITORY:
</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FC2FFF16F426FF35" box="[984,1050,151,177]" count="2" pageId="14" pageNumber="595" type="male">2 ♂♂</specimenCount>
: Sochi District. Krasnaya Polyana. 
<quantity id="EB789B21FFEC8B02FF60FF3AF0D0FF51" box="[151,236,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="595" unit="m" value="1200.0">1200 m</quantity>
a.s.l. 
<date id="583E1004FFEC8B02FED9FF3BF193FF51" box="[302,431,186,213]" pageId="14" pageNumber="595" value="2002-06-25">25.06.2002</date>
. V. Filippov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FD4DFF3AF2DAFF51" box="[698,742,187,213]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FD05FF3AF322FF51" box="[754,798,187,213]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: Achishkno Mts. 
<date id="583E1004FFEC8B02FC14FF3BF458FF51" box="[995,1124,186,213]" pageId="14" pageNumber="595" value="1996-07-25">25.07.1996</date>
. P.V. Romantsov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FF60FF5EF0FAFF7D" box="[151,198,223,249]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: Pseashkha Pass. 
<quantity id="EB789B21FFEC8B02FE65FF5EF1D5FF7D" box="[402,489,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="595" unit="m" value="2000.0">2000 m</quantity>
a.s.l., under stones. 
<date id="583E1004FFEC8B02FD24FF5EF368FF7D" box="[723,852,223,249]" pageId="14" pageNumber="595" value="1994-03-08">03.08.1994</date>
. A. Solodovnikov leg. (
<collectionCode id="4A91AE01FFEC8B02FB95FF5EF485FF7D" box="[1122,1209,223,249]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="595">ZMUC</collectionCode>
); 
<specimenCount id="3A86FD4DFFEC8B02FB39FF5EF4C1FF7D" box="[1230,1277,223,249]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: NW slope of Pseakhsha Mts. near north and south Pseakhsha. 
<quantity id="EB789B21FFEC8B02FD26FE82F315FE99" box="[721,809,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="14" pageNumber="595" unit="m" value="2400.0">2400 m</quantity>
a.s.l., under stones. 
<date id="583E1004FFEC8B02FBEEFE83F4A6FE99" box="[1049,1178,258,285]" pageId="14" pageNumber="595" value="1994-05-08">05.08.1994</date>
. A. Solodovnikov leg. (
<collectionCode id="4A91AE01FFEC8B02FF68FEA6F0CAFEC5" box="[159,246,295,321]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="595">ZMUC</collectionCode>
); 
<specimenCount id="3A86FD4DFFEC8B02FEFBFEA6F16CFEC5" box="[268,336,295,321]" count="4" pageId="14" pageNumber="595" type="male">4 ♂♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FEA9FEA6F19EFEC5" box="[350,418,295,321]" count="7" pageId="14" pageNumber="595" type="female">7 ♀♀</specimenCount>
: Lagonaki Plateau. 
<date id="583E1004FFEC8B02FD70FEA6F335FEC5" box="[647,777,295,321]" pageId="14" pageNumber="595" value="1991-07-20">20.07.1991</date>
. E.A. Khachikov leg. (cKh, cSh); 
<specimenCount id="3A86FD4DFFEC8B02FB60FEA6F4E0FEC5" box="[1175,1244,295,321]" count="2" pageId="14" pageNumber="595" type="male">2 ♂♂</specimenCount>
: same data, near snow field. 
<date id="583E1004FFEC8B02FEEAFECAF1A7FEE1" box="[285,411,331,357]" pageId="14" pageNumber="595" value="1996-06-28">28.06.1996</date>
(cSh); 
<emphasis id="1EF4EAD6FFEC8B02FE07FECBF2FEFEE0" bold="true" box="[496,706,330,356]" pageId="14" pageNumber="595">
<collectingRegion id="EE44F826FFEC8B02FE07FECBF24EFEE0" box="[496,626,330,356]" country="Russia" name="Stavropol'" pageId="14" pageNumber="595">STAVROPOL</collectingRegion>
AREA:
</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FD3DFECAF2CBFEE1" box="[714,759,331,357]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FCF3FECAF30EFEE1" box="[772,818,331,357]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: Teberda State Reserve, Kizgich Canyon near Arkhyz. 
<quantity id="EB789B21FFEC8B02FF60FEEEF0D2FE0D" box="[151,238,367,393]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="595" unit="m" value="1400.0">1400 m</quantity>
a.s.l., swamp with 
<taxonomicName id="EB804D47FFEC8B02FE3CFEEEF27BFE0D" box="[459,583,367,393]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="14" pageNumber="595" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="1EF4EAD6FFEC8B02FE3CFEEEF232FE0D" box="[459,526,367,393]" italics="true" pageId="14" pageNumber="595">Carex</emphasis>
spp.
</taxonomicName>
, 
<taxonomicName id="EB804D47FFEC8B02FDA3FEEEF2D6FE0D" box="[596,746,367,393]" class="Liliopsida" family="Juncaceae" genus="Juncus" kingdom="Plantae" order="Poales" pageId="14" pageNumber="595" phylum="Tracheophyta" rank="species" species="undefined-A">
<emphasis id="1EF4EAD6FFEC8B02FDA3FEEEF29EFE0C" box="[596,674,367,392]" italics="true" pageId="14" pageNumber="595">Juncus</emphasis>
sp. A
</taxonomicName>
.B. Ryvkin leg. (cSh); 
<emphasis id="1EF4EAD6FFEC8B02FC12FEEFF472FE0C" bold="true" box="[997,1102,366,392]" pageId="14" pageNumber="595">
<collectingRegion id="EE44F826FFEC8B02FC12FEEFF476FE0C" box="[997,1098,366,392]" country="Russia" name="Adygeya" pageId="14" pageNumber="595">ADYGEA</collectingRegion>
:
</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FBA1FEEEF4B9FE0D" box="[1110,1157,367,393]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: Lagonaki Plateau, karst near Psenodakh Lake. 
<quantity id="EB789B21FFEC8B02FE57FE12F203FE29" box="[416,575,403,429]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="14" pageNumber="595" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200 m</quantity>
a.s.l., alpica, under stones. 
<date id="583E1004FFEC8B02FC70FE12F434FE29" box="[903,1032,403,429]" pageId="14" pageNumber="595" value="1994-08-29">29.08.1994</date>
. A. Solodovnikov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FA90FE12F5AAFE29" box="[1383,1430,403,429]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: same Plateau, Oshten Mt. 07– 
<date id="583E1004FFEC8B02FE1AFE36F252FE55" box="[493,622,439,465]" pageId="14" pageNumber="595" value="1992-10-08">10.08.1992</date>
. E.A. Khachikov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FC40FE37F3C5FE55" box="[951,1017,438,465]" count="5" pageId="14" pageNumber="595" type="male">5 ♂♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FBF0FE37F475FE55" box="[1031,1097,438,465]" count="5" pageId="14" pageNumber="595" type="female">5 ♀♀</specimenCount>
: same data. 
<date id="583E1004FFEC8B02FB20FE36F569FE55" box="[1239,1365,439,465]" pageId="14" pageNumber="595" value="1996-06-28">28.06.1996</date>
(cKh, cSh); 
<emphasis id="1EF4EAD6FFEC8B02FF2EFE5BF1C4FE70" bold="true" box="[217,504,474,500]" pageId="14" pageNumber="595">
<collectingRegion id="EE44F826FFEC8B02FF2EFE5BF1CFFE70" box="[217,499,474,500]" country="Russia" name="Karachayevo-Cherkesiya" pageId="14" pageNumber="595">KARACHAY-CHERKESSIA</collectingRegion>
:
</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FE08FE5AF27CFE71" box="[511,576,475,501]" count="3" pageId="14" pageNumber="595" type="male">3 ♂♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FDBBFE5AF2B1FE71" box="[588,653,475,501]" count="4" pageId="14" pageNumber="595" type="female">4 ♀♀</specimenCount>
: Teberda Nature Reserve, Alibek Gorge. 
<date id="583E1004FFEC8B02FBACFE5AF4E0FE71" box="[1115,1244,475,501]" pageId="14" pageNumber="595" value="1996-10-07">10.07.1996</date>
. E.A. Khachikov leg. (cKh, сSh); 
<specimenCount id="3A86FD4DFFEC8B02FEA6FE7EF1AFFD9D" box="[337,403,511,537]" count="2" pageId="14" pageNumber="595" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FE57FE7EF1DDFD9D" box="[416,481,511,537]" count="3" pageId="14" pageNumber="595" type="female">3 ♀♀</specimenCount>
: same data, Dvuyazychnyi Glacier. 
<date id="583E1004FFEC8B02FC8FFE7EF3CAFD9D" box="[888,1014,511,537]" pageId="14" pageNumber="595" value="1996-12-07">12.07.1996</date>
(cKh, cSh); 
<specimenCount id="3A86FD4DFFEC8B02FB70FE7EF48FFD9D" box="[1159,1203,511,537]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FB37FE7EF53DFD9D" box="[1216,1281,511,537]" count="2" pageId="14" pageNumber="595" type="female">2 ♀♀</specimenCount>
: same Nature Reserve, Malaya Khatipara Mt. Alpic meadows, traps. 11.08– 
<date id="583E1004FFEC8B02FCB8FDA3F3ECFDB9" box="[847,976,546,573]" pageId="14" pageNumber="595" value="2007-08-15">15.08.2007</date>
. F.A. Martynovchenko leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FAA2FDA2F5BEFDB9" box="[1365,1410,547,573]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FA79FDA2F0FFFDE5" count="2" pageId="14" pageNumber="595" type="female">2 ♀♀</specimenCount>
: same data. 
<quantity id="EB789B21FFEC8B02FEBDFDC6F1A2FDE5" box="[330,414,583,609]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="14" pageNumber="595" unit="m" value="2800.0">2800 m</quantity>
a.s.l. 22– 
<date id="583E1004FFEC8B02FDFFFDC6F2B5FDE5" box="[520,649,583,609]" pageId="14" pageNumber="595" value="2007-07-23">23.07.2007</date>
.A.S. Prosvirov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FC41FDC6F3DDFDE5" box="[950,993,583,609]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: about 
<quantity id="EB789B21FFEC8B02FBC6FDC6F44BFDE5" box="[1073,1143,583,609]" metricMagnitude="4" metricUnit="m" metricValue="3.6" pageId="14" pageNumber="595" unit="km" value="36.0">36 km</quantity>
from Teberdinskiy Nature Reserve, Dombai Mt. 
<quantity id="EB789B21FFEC8B02FE67FDEAF217FD01" box="[400,555,619,645]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="14" pageNumber="595" unit="m" value="2950.0" valueMax="3000.0" valueMin="2900.0">2900–3000 m</quantity>
a.s.l., alpica. 
<date id="583E1004FFEC8B02FD32FDEAF37AFD01" box="[709,838,619,645]" pageId="14" pageNumber="595" value="2007-07-29">29.07.2007</date>
. A.S. Prosvirov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FB8FFDEAF480FD01" box="[1144,1212,619,645]" count="2" pageId="14" pageNumber="595" type="male">2 ♂♂</specimenCount>
: Teberda, Khatipara Pass. 
<date id="583E1004FFEC8B02FF2FFD0FF165FD2D" box="[216,345,654,681]" pageId="14" pageNumber="595" value="1962-07-15">15.07.1962</date>
. A.L. Tichomirova leg. (
<collectionCode id="4A91AE01FFEC8B02FD86FD0EF286FD2D" box="[625,698,655,681]" pageId="14" pageNumber="595">ZMM</collectionCode>
); 
<specimenCount id="3A86FD4DFFEC8B02FD38FD0EF32DFD2D" box="[719,785,655,681]" count="4" pageId="14" pageNumber="595" type="male">4 ♂♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FCE9FD0EF35DFD2D" box="[798,865,655,681]" count="4" pageId="14" pageNumber="595" type="female">4 ♀♀</specimenCount>
: camp near Bolshaya Khatipara Mt. Alpica, under stones on a slope near camp. 
<date id="583E1004FFEC8B02FE1DFD32F250FD49" box="[490,620,691,717]" pageId="14" pageNumber="595" value="2007-04-07">04.07.2007</date>
. A.S. Prosvirov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FC50FD32F3D6FD49" box="[935,1002,691,717]" count="2" pageId="14" pageNumber="595" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FC0FFD32F407FD49" box="[1016,1083,691,717]" count="4" pageId="14" pageNumber="595" type="female">4 ♀♀</specimenCount>
: same Gorge, road to Dombai. 
<date id="583E1004FFEC8B02FF60FD56F124FD75" box="[151,280,727,753]" pageId="14" pageNumber="595" value="1996-07-14">14.07.1996</date>
. E.A. Khachikov leg. (cKh, cSh); 
<specimenCount id="3A86FD4DFFEC8B02FD6EFD56F2E0FD75" box="[665,732,727,753]" count="2" pageId="14" pageNumber="595" type="male">2 ♂♂</specimenCount>
: same Gorge, environs of Dombai. 
<date id="583E1004FFEC8B02FB9DFD56F4D7FD75" box="[1130,1259,727,753]" pageId="14" pageNumber="595" value="1994-07-17">17.07.1994</date>
. Yu.G. Arzanov &amp; I.V. Shokhin leg. (cKh, cSh); 
<specimenCount id="3A86FD4DFFEC8B02FDF2FD7AF20EFC91" box="[517,562,763,789]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FDC9FD7AF2BCFC91" box="[574,640,763,789]" count="4" pageId="14" pageNumber="595" type="female">4 ♀♀</specimenCount>
: Dombai, Musa-Lchitara Mt., 
<quantity id="EB789B21FFEC8B02FC2DFD7AF40CFC91" box="[986,1072,763,789]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="595" unit="m" value="3000.0">3000 m</quantity>
a.s.l., under stones. 
<date id="583E1004FFEC8B02FAE0FD7AF5A5FC91" box="[1303,1433,763,789]" pageId="14" pageNumber="595" value="2008-07-26">26.07.2008</date>
. F.V. Ivanov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FE60FC9EF1E5FCBD" box="[407,473,799,825]" count="2" pageId="14" pageNumber="595" type="male">2 ♂♂</specimenCount>
: 
<quantity id="EB789B21FFEC8B02FE14FC9FF20BFCBD" box="[483,567,798,825]" metricMagnitude="3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="14" pageNumber="595" unit="km" value="4.5" valueMax="5.0" valueMin="4.0">4–5 km</quantity>
NE of Dombai, SW slopes of Musat-Cheri Range, 
<geoCoordinate id="49B45003FFEC8B02FB98FC9FF53CFCBD" box="[1135,1280,798,825]" degrees="43" direction="north" minutes="17.254" orientation="latitude" pageId="14" pageNumber="595" precision="1" value="43.287567">43°17.254´N</geoCoordinate>
<geoCoordinate id="49B45003FFEC8B02FAF0FC9EF5A5FCBD" box="[1287,1433,799,825]" degrees="41" direction="east" minutes="40.919" orientation="longitude" pageId="14" pageNumber="595" precision="1" value="41.681984">41°40.919´E</geoCoordinate>
. 
<quantity id="EB789B21FFEC8B02FF60FCC2F0D2FCD9" box="[151,238,835,861]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="595" unit="m" value="3000.0">3000 m</quantity>
a.s.l., alpine zone, under stones near snow field. 
<date id="583E1004FFEC8B02FCD3FCC3F399FCD9" box="[804,933,834,861]" pageId="14" pageNumber="595" value="2011-07-25">25.07.2011</date>
. A. Solodovnikov leg. (
<collectionCode id="4A91AE01FFEC8B02FB43FCC2F537FCD9" box="[1204,1291,835,861]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="595">ZMUC</collectionCode>
); 
<specimenCount id="3A86FD4DFFEC8B02FAD7FCC2F573FCD9" box="[1312,1359,835,861]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: upper flow of Beskes River. 2900 a.s.l. 
<date id="583E1004FFEC8B02FDEEFCE6F2A6FC05" box="[537,666,871,897]" pageId="14" pageNumber="595" value="1997-08-14">14.08.1997</date>
. I.A. Solodovnikov leg. (
<collectionCode id="4A91AE01FFEC8B02FC4AFCE6F437FC05" box="[957,1035,871,897]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="595">NMW</collectionCode>
); 
<specimenCount id="3A86FD4DFFEC8B02FBD7FCE6F472FC05" box="[1056,1102,871,897]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: Tumbashi tract. 
<date id="583E1004FFEC8B02FAE0FCE7F5A5FC05" box="[1303,1433,870,897]" pageId="14" pageNumber="595" value="1991-09-15">15.09.1991</date>
. 
<quantity id="EB789B21FFEC8B02FF60FC0AF0D0FC21" box="[151,236,907,933]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="595" unit="m" value="2000.0">2000 m</quantity>
a.s.l., stream. V. Grebennikov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FD37FC0AF2D0FC21" box="[704,748,907,933]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FD0FFC0AF318FC21" box="[760,804,907,933]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: Epchik Pass. 06.1992. Yu.G. Arzanov leg. (cKh); 
<specimenCount id="3A86FD4DFFEC8B02FAAAFC0AF5ABFC21" box="[1373,1431,907,933]" count="10" pageId="14" pageNumber="595" type="male">10 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FF60FC2EF0D4FC4D" box="[151,232,943,969]" count="10" pageId="14" pageNumber="595" type="female">10 ♀♀</specimenCount>
: upper flow of Dukka R., Dukka Mt. Pass. 
<date id="583E1004FFEC8B02FD2DFC2EF367FC4D" box="[730,859,943,969]" pageId="14" pageNumber="595" value="1996-07-17">17.07.1996</date>
. E.A. Khachikov leg. (cKh, cSh); 
<specimenCount id="3A86FD4DFFEC8B02FB14FC2EF52DFC4D" box="[1251,1297,943,969]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FAE9FC2EF55CFC4D" box="[1310,1376,943,969]" count="2" pageId="14" pageNumber="595" type="female">2 ♀♀</specimenCount>
: env. Nizhyaya Teberda, bank of Azchek River. 
<quantity id="EB789B21FFEC8B02FD83FC52F32CFC69" box="[628,784,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="14" pageNumber="595" unit="m" value="2650.0" valueMax="2700.0" valueMin="2600.0">2600–2700 m</quantity>
a.s.l. 
<date id="583E1004FFEC8B02FCA4FC52F3E8FC69" box="[851,980,979,1005]" pageId="14" pageNumber="595" value="2007-01-08">01.08.2007</date>
. A.S. Prosvirov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FAF0FC52F575FC69" box="[1287,1353,979,1005]" count="2" pageId="14" pageNumber="595" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FAA2FC52F5AAFC69" box="[1365,1430,979,1005]" count="4" pageId="14" pageNumber="595" type="female">4 ♀♀</specimenCount>
: same data. 
<quantity id="EB789B21FFEC8B02FEE3FC76F192FB95" box="[276,430,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="14" pageNumber="595" unit="m" value="2700.0" valueMax="2800.0" valueMin="2600.0">2600–2800 m</quantity>
a.s.l. 
<date id="583E1004FFEC8B02FE18FC76F24DFB95" box="[495,625,1015,1041]" pageId="14" pageNumber="595" value="2007-07-26">26.07.2007</date>
. A.S. Prosvirov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FC57FC76F3F7FB95" box="[928,971,1015,1041]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FC20FC76F43FFB95" box="[983,1027,1015,1041]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: env. of Nizhnyaya Teberda, bank of Azgek R. 
<quantity id="EB789B21FFEC8B02FEF2FB9AF1A3FBB1" box="[261,415,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="14" pageNumber="595" unit="m" value="2650.0" valueMax="2700.0" valueMin="2600.0">2600–2700 m</quantity>
a.s.l. 
<date id="583E1004FFEC8B02FE29FB9AF263FBB1" box="[478,607,1051,1077]" pageId="14" pageNumber="595" value="2007-02-08">02.08.2007</date>
. A.S. Prosvirov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FC7EFB9AF388FBB1" box="[905,948,1051,1077]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: upper flow of Burnaya R. 
<date id="583E1004FFEC8B02FB29FB9BF55CFBB1" box="[1246,1376,1050,1077]" pageId="14" pageNumber="595" value="1995-04-08">04.08.1995</date>
. E.A. Khachikov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FE6FFBBEF1FAFBDD" box="[408,454,1087,1113]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: Daut Gorge. 
<date id="583E1004FFEC8B02FD92FBBEF2DAFBDD" box="[613,742,1087,1113]" pageId="14" pageNumber="595" value="1994-11-07">11.07.1994</date>
. Shokhin I. leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FC1AFBBEF426FBDD" box="[1005,1050,1087,1113]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FBD0FBBEF455FBDD" box="[1063,1129,1087,1113]" count="3" pageId="14" pageNumber="595" type="female">3 ♀♀</specimenCount>
: Arkasara Mts. environs of Pkhiya. 
<date id="583E1004FFEC8B02FF05FBE2F14FFBF9" box="[242,371,1123,1149]" pageId="14" pageNumber="595" value="1996-08-14">14.08.1996</date>
. E.A. Khachikov leg. (cKh, cSh); 
<emphasis id="1EF4EAD6FFEC8B02FD18FBE3F43DFBF8" bold="true" box="[751,1025,1122,1148]" pageId="14" pageNumber="595">
<collectingRegion id="EE44F826FFEC8B02FD18FBE3F3C1FBF8" box="[751,1021,1122,1148]" country="Russia" name="Kabardino-Balkariya" pageId="14" pageNumber="595">KABARDINO-BALKARIA</collectingRegion>
:
</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FBF0FBE2F408FBF9" box="[1031,1076,1123,1149]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: Bezenchuk, Cherek-Balkarskiy R. 
<date id="583E1004FFEC8B02FF4DFB06F106FB25" box="[186,314,1159,1185]" pageId="14" pageNumber="595" value="1992-11-08">11.08.1992</date>
. V. Grebennikov leg. (cKh); 
<specimenCount id="3A86FD4DFFEC8B02FD72FB06F2F6FB25" box="[645,714,1159,1185]" count="2" pageId="14" pageNumber="595" type="male">2 ♂♂</specimenCount>
: upper reaches of Chechenskiy Gorge. 29.07– 
<date id="583E1004FFEC8B02FB2EFB06F566FB25" box="[1241,1370,1159,1185]" pageId="14" pageNumber="595" value="1998-07-08">07.08.1998</date>
. O.G. Brekhov leg. (cKh); 
<emphasis id="1EF4EAD6FFEC8B02FE89FB2BF29EFB40" bold="true" box="[382,674,1194,1220]" pageId="14" pageNumber="595">
<collectingRegion id="EE44F826FFEC8B02FE89FB2BF2A2FB40" box="[382,670,1194,1220]" country="Russia" name="North Ossetia" pageId="14" pageNumber="595">NORTH OSSETIA-ALANIA</collectingRegion>
:
</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FD5EFB2AF2D7FB41" box="[681,747,1195,1221]" count="4" pageId="14" pageNumber="595" type="male">4 ♂♂</specimenCount>
: Tsey Gorge. 
<date id="583E1004FFEC8B02FC73FB2AF439FB41" box="[900,1029,1195,1221]" pageId="14" pageNumber="595" value="1990-09-19">19.09.1990</date>
. A. Klimenko leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FAD5FB2AF573FB41" box="[1314,1359,1195,1221]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: 
<collectingRegion id="EE44F826FFEC8B02FAACFB2AF0D5FB6D" country="Russia" name="North Ossetia" pageId="14" pageNumber="595">North Ossetia</collectingRegion>
Nature Reserve, basin of Ardon River. 1982. S.K. Alekseev leg. (
<collectionCode id="4A91AE01FFEC8B02FC42FB4EF3C2FB6D" box="[949,1022,1231,1257]" pageId="14" pageNumber="595">ZMM</collectionCode>
); 
<specimenCount id="3A86FD4DFFEC8B02FBF8FB4EF406FB6D" box="[1039,1082,1231,1257]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: basin of Ardon R., Tsei R. 
<quantity id="EB789B21FFEC8B02FA93FB4FF091FA89" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="595" unit="m" value="2500.0">2500 m</quantity>
a.s.l. 
<date id="583E1004FFEC8B02FF07FB72F14DFA89" box="[240,369,1267,1293]" pageId="14" pageNumber="595" value="1990-09-19">19.09.1990</date>
. A. Klimenko leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FD67FB72F280FA89" box="[656,700,1267,1293]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FD3EFB72F2C9FA89" box="[713,757,1267,1293]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: upper reaches of Fiagdon R., Khilak Gorge, alpic mountain path. 
<quantity id="EB789B21FFEC8B02FF21FA96F14FFAB5" box="[214,371,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.1" metricValueMin="2.8" pageId="14" pageNumber="595" unit="m" value="2950.0" valueMax="3100.0" valueMin="2800.0">2800–3100 m</quantity>
a.s.l. 10– 
<date id="583E1004FFEC8B02FE17FA96F25FFAB5" box="[480,611,1303,1329]" pageId="14" pageNumber="595" value="1984-07-13">13.07.1984</date>
. S.K. Alekseev leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FC63FA96F3E4FAB5" box="[916,984,1303,1329]" count="2" pageId="14" pageNumber="595" type="male">2 ♂♂</specimenCount>
: same data. 
<date id="583E1004FFEC8B02FB94FA96F4D9FAB5" box="[1123,1253,1303,1329]" pageId="14" pageNumber="595" value="1984-10-08">10.08.1984</date>
. S. Alexeev leg. (
<collectionCode id="4A91AE01FFEC8B02FF68FABAF0D4FAD1" box="[159,232,1339,1365]" pageId="14" pageNumber="595">ZMM</collectionCode>
); 
<specimenCount id="3A86FD4DFFEC8B02FF0BFABAF115FAD1" box="[252,297,1339,1365]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: Zaramag, Karmadon Gorge. 
<date id="583E1004FFEC8B02FD8FFABBF2C5FAD1" box="[632,761,1338,1365]" pageId="14" pageNumber="595" value="1999-06-05">06.05.1999</date>
. Murashov leg. (cSh); 
<emphasis id="1EF4EAD6FFEC8B02FC0EFABBF445FAD0" bold="true" box="[1017,1145,1338,1364]" pageId="14" pageNumber="595">
<collectingRegion id="EE44F826FFEC8B02FC0EFABBF449FAD0" box="[1017,1141,1338,1364]" country="Russia" name="Dagestan" pageId="14" pageNumber="595">DAGESTAN</collectingRegion>
:
</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FB77FABAF490FAD1" box="[1152,1196,1339,1365]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: Kasan (camp). 
<quantity id="EB789B21FFEC8B02FA93FABAF091FAFD" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="595" unit="m" value="2000.0">2000 m</quantity>
a.s.l. 
<date id="583E1004FFEC8B02FF18FADEF14DFAFD" box="[239,369,1375,1401]" pageId="14" pageNumber="595" value="1998-07-13">13.07.1998</date>
. E.V. Iljina leg. (cKh); 
<specimenCount id="3A86FD4DFFEC8B02FD83FADEF29DFAFD" box="[628,673,1375,1401]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: Bogos mts. 
<quantity id="EB789B21FFEC8B02FCC6FADFF3F0FAFD" box="[817,972,1374,1401]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="14" pageNumber="595" unit="m" value="2250.0" valueMax="3000.0" valueMin="1500.0">1500–3000 m</quantity>
a.s.l. 
<date id="583E1004FFEC8B02FBF9FADEF4B3FAFD" box="[1038,1167,1375,1401]" pageId="14" pageNumber="595" value="1994-07-28">28.07.1994</date>
. E.V. Iljina leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FA79FADEF0FFFA19" count="4" pageId="14" pageNumber="595" type="male">4 ♂♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FF38FA02F0C6FA19" box="[207,250,1411,1437]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: env. Makhachkala S salt-marsh. 
<date id="583E1004FFEC8B02FD98FA02F2CCFA19" box="[623,752,1411,1437]" pageId="14" pageNumber="595" value="1996-07-07">07.07.1996</date>
. E.V. Iljina leg. (cKh, cSh); 
<specimenCount id="3A86FD4DFFEC8B02FBDFFA02F456FA19" box="[1064,1130,1411,1437]" count="2" pageId="14" pageNumber="595" type="male">2 ♂♂</specimenCount>
: Knyukh Mts. 
<quantity id="EB789B21FFEC8B02FAF9FA02F55EFA19" box="[1294,1378,1411,1437]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="595" unit="m" value="2000.0">2000 m</quantity>
a.s.l. 10– 
<date id="583E1004FFEC8B02FF37FA26F17EFA45" box="[192,322,1447,1473]" pageId="14" pageNumber="595" value="2004-11-07">11.07.2004</date>
. E.V. Iljina leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FDB4FA26F253FA45" box="[579,623,1447,1473]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: Tarki-Tau. 
<date id="583E1004FFEC8B02FD0FFA27F346FA45" box="[760,890,1446,1473]" pageId="14" pageNumber="595" value="1998-01-05">01.05.1998</date>
. E.V. Iljina leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FB8CFA26F494FA45" box="[1147,1192,1447,1473]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FB43FA26F4CAFA45" box="[1204,1270,1447,1473]" count="4" pageId="14" pageNumber="595" type="female">4 ♀♀</specimenCount>
: upper flow of Chiragshay R., Alakhundag Mt. 
<quantity id="EB789B21FFEC8B02FDFEFA4AF25CFA61" box="[521,608,1483,1509]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="595" unit="m" value="3000.0">3000 m</quantity>
a.s.l. 
<date id="583E1004FFEC8B02FD52FA4BF31BFA61" box="[677,807,1482,1509]" pageId="14" pageNumber="595" value="2006-07-15">15.07.2006</date>
. E.V. Iljina leg. (cKh, cSh); 
<emphasis id="1EF4EAD6FFEC8B02FB9AFA4BF5B9FA61" bold="true" box="[1133,1413,1482,1509]" pageId="14" pageNumber="595">
<collectingCountry id="54977654FFEC8B02FB9AFA4BF4C5FA60" box="[1133,1273,1482,1508]" name="Georgia" pageId="14" pageNumber="595">GEORGIA</collectingCountry>
: 
<collectingRegion id="EE44F826FFEC8B02FAF1FA4BF540FA60" box="[1286,1404,1482,1508]" country="Georgia" name="Abkhazia" pageId="14" pageNumber="595">ABKHASIA</collectingRegion>
:
</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FA79FA4AF092F98D" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: NNW Sukhumi, Bzybskiy Mts., 
<quantity id="EB789B21FFEC8B02FDDBFA6FF2B4F98D" box="[556,648,1518,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="14" pageNumber="595" unit="km" value="1.75">1.75 km</quantity>
SSW Gudaut Pass, 
<geoCoordinate id="49B45003FFEC8B02FC9CFA6FF3C7F98D" box="[875,1019,1518,1545]" degrees="43" direction="north" minutes="16" orientation="latitude" pageId="14" pageNumber="595" precision="15" seconds="50" value="43.280556">43°16´50´´N</geoCoordinate>
<geoCoordinate id="49B45003FFEC8B02FBF5FA6FF4AEF98D" box="[1026,1170,1518,1545]" degrees="40" direction="east" minutes="48" orientation="longitude" pageId="14" pageNumber="595" precision="15" seconds="05" value="40.801388">40°48´05´´E</geoCoordinate>
. 
<quantity id="EB789B21FFEC8B02FB6AFA6FF4C9F98D" box="[1181,1269,1518,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.4949999999999999" pageId="14" pageNumber="595" unit="m" value="1495.0">1495 m</quantity>
. 
<quantity id="EB789B21FFEC8B02FAF6FA6EF5A1F98D" box="[1281,1437,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="14" pageNumber="595" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">1200–1400 m</quantity>
a.s.l. A.S. Prosvirov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FE0DF992F21AF9A9" box="[506,550,1555,1581]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: western part of Bzybskiy Mts., about 
<quantity id="EB789B21FFEC8B02FC2DF992F42FF9A9" box="[986,1043,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="595" unit="km" value="1.0">1 km</quantity>
NW Narpa. 
<quantity id="EB789B21FFEC8B02FB55F992F501F9A9" box="[1186,1341,1555,1581]" metricMagnitude="3" metricUnit="m" metricValue="2.18" metricValueMax="2.2" metricValueMin="2.16" pageId="14" pageNumber="595" unit="m" value="2180.0" valueMax="2200.0" valueMin="2160.0">2160–2200 m</quantity>
a.s.l., in flight. 
<date id="583E1004FFEC8B02FF16F9B6F15FF9D5" box="[225,355,1591,1617]" pageId="14" pageNumber="595" value="2010-07-29">29.07.2010</date>
. A.S. Prosvirov leg. (cSh); 
<specimenCount id="3A86FD4DFFEC8B02FD65F9B6F283F9D5" box="[658,703,1591,1617]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: northern slope of Bzybskiy Mts., 
<quantity id="EB789B21FFEC8B02FBB7F9B6F444F9D5" box="[1088,1144,1591,1617]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="595" unit="km" value="4.0">4 km</quantity>
NW Dzyshra. 
<quantity id="EB789B21FFEC8B02FAE9F9B7F091F9F1" metricMagnitude="3" metricUnit="m" metricValue="1.845" metricValueMax="1.85" metricValueMin="1.8399999999999999" pageId="14" pageNumber="595" unit="m" value="1845.0" valueMax="1850.0" valueMin="1840.0">1840–1850 m</quantity>
a.s.l. 
<date id="583E1004FFEC8B02FF1AF9DAF153F9F1" box="[237,367,1627,1653]" pageId="14" pageNumber="595" value="2010-04-08">04.08.2010</date>
. A.S. Prosvirov leg. (cSh); 
<emphasis id="1EF4EAD6FFEC8B02FD6DF9DAF2D2F9F1" bold="true" box="[666,750,1627,1653]" pageId="14" pageNumber="595">KHEVI:</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FD03F9DAF308F9F1" box="[756,820,1627,1653]" count="2" pageId="14" pageNumber="595" type="female">2 ♀♀</specimenCount>
: ’Cauc[asus]. Centr. Kobi’, ’ 
<taxonomicName id="EB804D47FFEC8B02FB99F9DAF0DEF91D" authority="Mnnh." authorityName="Mnnh." class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="595" phylum="Arthropoda" rank="species" species="globulicollis">Geodromicus globulicollis Mnnh.</taxonomicName>
det. Roubal’ (
<collectionCode id="4A91AE01FFEC8B02FE89F9FEF1FBF91D" box="[382,455,1663,1689]" pageId="14" pageNumber="595">ZMM</collectionCode>
); 
<emphasis id="1EF4EAD6FFEC8B02FE2DF9FFF27CF91C" bold="true" box="[474,576,1662,1688]" pageId="14" pageNumber="595">SVANETI:</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FDB1F9FEF24EF91D" box="[582,626,1663,1689]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: ‘Cauc. Svanetia Latpari 
<quantity id="EB789B21FFEC8B02FC7AF9FEF3DEF91D" box="[909,994,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="3.09" pageId="14" pageNumber="595" unit="m" value="3090.0">3090 m</quantity>
Dr. Veselý’ (
<collectionCode id="4A91AE01FFEC8B02FB85F9FEF4F4F91D" box="[1138,1224,1663,1689]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="14" pageNumber="595">NMPC</collectionCode>
); 
<specimenCount id="3A86FD4DFFEC8B02FB2CF9FEF53CF91D" box="[1243,1280,1663,1689]" count="1" pageId="14" pageNumber="595" type="male">1♂</specimenCount>
: Latpari Pass. 
<date id="583E1004FFEC8B02FF60F922F124F939" box="[151,280,1699,1725]" pageId="14" pageNumber="595" value="1930-11-08">11.08.1930</date>
. Bogatchov leg. (
<collectionCode id="4A91AE01FFEC8B02FE29F922F21BF939" box="[478,551,1699,1725]" pageId="14" pageNumber="595">ZMM</collectionCode>
); 
<specimenCount id="3A86FD4DFFEC8B02FDCBF922F255F939" box="[572,617,1699,1725]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
, 
<specimenCount id="3A86FD4DFFEC8B02FD80F922F298F939" box="[631,676,1699,1725]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: ‘CAUCASUS Svanetia [1]913 Dr. Veselý’ (
<collectionCode id="4A91AE01FFEC8B02FB56F922F4CBF939" box="[1185,1271,1699,1725]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="14" pageNumber="595">NMPC</collectionCode>
); 
<emphasis id="1EF4EAD6FFEC8B02FAFDF923F0C5F964" bold="true" pageId="14" pageNumber="595">
<collectingCountry id="54977654FFEC8B02FAFDF923F5ABF938" box="[1290,1431,1698,1724]" name="Armenia" pageId="14" pageNumber="595">ARMENIA</collectingCountry>
: 
<collectingRegion id="EE44F826FFEC8B02FF60F947F0C9F964" box="[151,245,1734,1760]" country="Armenia" name="Tavush" pageId="14" pageNumber="595">TAVUSH</collectingRegion>
:
</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FF09F946F115F965" box="[254,297,1735,1761]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: Dilijan Reserve, Akhnabat valley near Salakh. 
<quantity id="EB789B21FFEC8B02FCCFF947F3EDF965" box="[824,977,1734,1761]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="14" pageNumber="595" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400–1500 m</quantity>
a.s.l., forest with 
<taxonomicName id="EB804D47FFEC8B02FB63F946F4E6F964" box="[1172,1242,1735,1760]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="595" phylum="Tracheophyta" rank="genus">
<emphasis id="1EF4EAD6FFEC8B02FB63F946F4E6F964" box="[1172,1242,1735,1760]" italics="true" pageId="14" pageNumber="595">Fagus</emphasis>
</taxonomicName>
, 
<taxonomicName id="EB804D47FFEC8B02FB10F946F51BF964" box="[1255,1319,1735,1760]" class="Mammalia" family="Mustelidae" genus="Taxus" kingdom="Animalia" order="Carnivora" pageId="14" pageNumber="595" phylum="Chordata" rank="genus">
<emphasis id="1EF4EAD6FFEC8B02FB10F946F51BF964" box="[1255,1319,1735,1760]" italics="true" pageId="14" pageNumber="595">Taxus</emphasis>
</taxonomicName>
, 
<taxonomicName id="EB804D47FFEC8B02FAC3F946F5A0F965" box="[1332,1436,1735,1761]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="595" phylum="Tracheophyta" rank="genus">
<emphasis id="1EF4EAD6FFEC8B02FAC3F946F5A0F965" box="[1332,1436,1735,1761]" italics="true" pageId="14" pageNumber="595">Carpinus</emphasis>
</taxonomicName>
etc., in litter. 
<date id="583E1004FFEC8B02FEDBF96AF191F881" box="[300,429,1771,1797]" pageId="14" pageNumber="595" value="1983-04-17">17.04.1983</date>
. S. Golovatch leg. (
<collectionCode id="4A91AE01FFEC8B02FD7DF96AF2EFF881" box="[650,723,1771,1797]" pageId="14" pageNumber="595">ZMM</collectionCode>
); 
<specimenCount id="3A86FD4DFFEC8B02FD10F96AF32FF881" box="[743,787,1771,1797]" count="1" pageId="14" pageNumber="595" type="female">1 ♀</specimenCount>
: ‘Caucasus. Armen.Geb. Leder Reitter’, ‘brevicolle Fauv. det. Luze’, ‘Geodr. 
<taxonomicName id="EB804D47FFEC8B02FE98F88EF2C7F8AD" authority="Fvl. Coll." authorityName="Fvl. Coll." box="[367,763,1807,1833]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="595" phylum="Arthropoda" rank="variety" species="globulicollis" variety="brevicolle">globulicollis v. brevicolle Fvl. Coll.</taxonomicName>
Reitter’ (
<collectionCode id="4A91AE01FFEC8B02FC94F88EF383F8AD" box="[867,959,1807,1833]" pageId="14" pageNumber="595">NHHM</collectionCode>
); 
<emphasis id="1EF4EAD6FFEC8B02FC26F88FF55AF8AC" bold="true" box="[977,1382,1806,1832]" pageId="14" pageNumber="595">REGION WITHOUT EXACT LOCALITY:</emphasis>
<specimenCount id="3A86FD4DFFEC8B02FA9CF88EF5ABF8AD" box="[1387,1431,1807,1833]" count="1" pageId="14" pageNumber="595" type="male">1 ♂</specimenCount>
: ‘Caucasus 4252 -5.’, ‘4252. macroelytron Fourcr.?’ (
<collectionCode id="4A91AE01FFEC8B02FD1CF8B2F308F8C9" box="[747,820,1843,1869]" pageId="14" pageNumber="595">ZMM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="649A654FFFEC8B03FF30F8D7F500FA19" lastPageId="15" lastPageNumber="596" pageId="14" pageNumber="595" type="description">
<paragraph id="2C3F36C4FFEC8B02FF30F8D7F133F859" blockId="14.[151,1437,150,2013]" pageId="14" pageNumber="595">
<emphasis id="1EF4EAD6FFEC8B02FF30F8D7F149F8F4" bold="true" box="[199,373,1878,1904]" pageId="14" pageNumber="595">Redescription.</emphasis>
Measurements (n=100): HW: 0.65–0.85; HL: 0.47–0.63; OL: 0.15–0.17; TL: 0.12–0.15; AL (averaged): 2.30; PL: 0.52–0.78; PWmax: 0.78–1.07; PWmin: 0.64–0.80; ESL: 0.88–1.25; EW: 1.06–1.52; MTbL (averaged): 0.77; MTrL (averaged): 0.42 (MTrL 1–4: 0.22; MTrL 5: 0.20); AW: 1.08–1.62; AedL: 0.75–0.85; BL: 3.05–5.30.
</paragraph>
<paragraph id="2C3F36C4FFED8B03FF30FF16F300FE0D" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="596">
Habitus as in 
<figureCitation id="B4BB2A41FFED8B03FEADFF16F1A0FF35" box="[346,412,151,177]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1587,1611]" captionTargetBox="[183,1404,185,1555]" captionTargetId="figure-30@4.[177,1410,179,1561]" captionTargetPageId="4" captionText="FIGURES 1–6. Habitus of Geodromicus: 1—G. brevicollis (Russia: Karachay-Cherkessia), 2—G. brevicollis (Turkey: Ardahan), 3—G. jarrigeanus (holotype), 4—G. ziganensis (holotype), 5—G. alborzensis (holotype), 6—G. transversus (Russia: Karachay-Cherkessia). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12189107" httpUri="https://zenodo.org/record/12189107/files/figure.png" pageId="15" pageNumber="596">Fig. 6</figureCitation>
. Body brown to reddish-brown, usually with darker head and abdomen and paler elytra (some specimens with indistinct large yellowish spots in middle portion of each elytron); mouthparts, antennae and legs yellow-brown (antennomeres 1–2 or 1–3 sometimes paler); maxillary palpomeres 2 and 4, and tarsi yellowish. Head with irregular fine microsculpture: sparser, finer and transverse in middle, and denser, isodiametric or transverse on infraorbital portions (some specimens without visible meshes); neck with very dense and coarse isodiametric microreticulation; pronotum without or with very fine transverse or isodiametric microsculpture in middle portion; scutellum with very fine transverse meshes or without them.
</paragraph>
<paragraph id="2C3F36C4FFED8B03FF30FE12F142FD49" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="596">Head 1.3 times as broad as long, with very deep and wide anterio-median depression, sometimes slightly narrowed and extended basad; interocellar depression deep and moderately narrow, separated from infraorbital portions by diagonally convergent latero-anteriad anteocellar foveae, reaching level of anterior margin of eyes. Distance between ocelli slightly shorter than distance between ocellus and posterior margin of eyes. Punctation small and sparse, finer and sometimes indistinct in middle between anteocellar foveae. Mediobasal portion between ocelli usually with dense irregular transverse elevations. Apical segment of maxillary palpus short, from basal portion gradually narrowed toward subacute apex. Antenna reaching apical third of elytra when reclined; antennomeres 4–7 slightly broader and longer than 3, 8–10 slightly broader than 7, apical antennomere 1.3–1.7 times as long as preceding segement.</paragraph>
<paragraph id="2C3F36C4FFED8B03FF30FD56F512FC05" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="596">Pronotum 1.3–1.5 times as broad as long, widest slightly in front of middle or in anterior third, gradually or sharply narrowed basad toward obtuse or subacute hind angles; laterobasal margins moderately short, widely concave; mediobasal depression shallow or very deep, oval or transverse; anterior and posterior margins distinctly concave in middle. Punctation about as that on head, but more regular, denser and sometimes larger and deeper, sparser in middle. Some specimens with irregular transverse elevations between punctures in front of basal margin.</paragraph>
<paragraph id="2C3F36C4FFED8B03FF30FC0AF574FC21" blockId="15.[151,1437,151,2013]" box="[199,1352,907,933]" pageId="15" pageNumber="596">Elytra 1.2 times as broad as long. Punctation about as that on pronotum, dense, usually finer in middle.</paragraph>
<paragraph id="2C3F36C4FFED8B03FF30FC2EF085FC69" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="596">Abdomen slightly broader than elytra, with two large transverse tomentose spots in middle of abdominal tergite IV.</paragraph>
<paragraph id="2C3F36C4FFED8B03FF30FC76F3C2FB6D" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="596">
Male. Protarsomeres 1–4 wide. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII concave. Aedeagus with wide basal part and very wide median lobe, gradually narrowed toward rounded apex; parameres narrow, slightly broadened apically, reaching or slightly exceeding apex of median lobe, with 2–3 moderately short apical and 1–2 preapical setae; parameral surface of median lobe distinctly elevated, usually with narrow sclerotized lateral edges; internal sac moderately short and narrow, with paired narrow elongate sclerotized structures in basal portion, with moderately long narrow flagellum between them (
<figureCitation id="B4BB2A41FFED8B03FB20FB2AF5ADFB41" box="[1239,1425,1195,1221]" captionStart="FIGURES 16–23" captionStartId="13.[151,264,1663,1687]" captionTargetBox="[159,1424,190,1630]" captionTargetId="figure-15@13.[151,1436,181,1639]" captionTargetPageId="13" captionText="FIGURES 16–23. Aedeagus of Geodromicus alborzensis (Figs. 16–17: holotype) and G. transversus (Figs. 18–19: Russia, Krasnaya Polyana; Fig. 20: lectotype of G. transversus; Fig. 21: lectotype of G. flavipes; Fig. 22–23: Karachay-Cherkessia, Epchik Pass): 16, 18–22—parameral view, 17, 19—lateral view, 23—internal sac. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12189114" httpUri="https://zenodo.org/record/12189114/files/figure.png" pageId="15" pageNumber="596">Figs. 18, 20–22</figureCitation>
). Lateral aspect of the aedeagus as in 
<figureCitation id="B4BB2A41FFED8B03FDDCFB4EF241FB6D" box="[555,637,1231,1257]" captionStart="FIGURES 16–23" captionStartId="13.[151,264,1663,1687]" captionTargetBox="[159,1424,190,1630]" captionTargetId="figure-15@13.[151,1436,181,1639]" captionTargetPageId="13" captionText="FIGURES 16–23. Aedeagus of Geodromicus alborzensis (Figs. 16–17: holotype) and G. transversus (Figs. 18–19: Russia, Krasnaya Polyana; Fig. 20: lectotype of G. transversus; Fig. 21: lectotype of G. flavipes; Fig. 22–23: Karachay-Cherkessia, Epchik Pass): 16, 18–22—parameral view, 17, 19—lateral view, 23—internal sac. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12189114" httpUri="https://zenodo.org/record/12189114/files/figure.png" pageId="15" pageNumber="596">Fig. 19</figureCitation>
; inflated internal sac as in 
<figureCitation id="B4BB2A41FFED8B03FC5FFB4EF3C6FB6D" box="[936,1018,1231,1257]" captionStart="FIGURES 16–23" captionStartId="13.[151,264,1663,1687]" captionTargetBox="[159,1424,190,1630]" captionTargetId="figure-15@13.[151,1436,181,1639]" captionTargetPageId="13" captionText="FIGURES 16–23. Aedeagus of Geodromicus alborzensis (Figs. 16–17: holotype) and G. transversus (Figs. 18–19: Russia, Krasnaya Polyana; Fig. 20: lectotype of G. transversus; Fig. 21: lectotype of G. flavipes; Fig. 22–23: Karachay-Cherkessia, Epchik Pass): 16, 18–22—parameral view, 17, 19—lateral view, 23—internal sac. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12189114" httpUri="https://zenodo.org/record/12189114/files/figure.png" pageId="15" pageNumber="596">Fig. 23</figureCitation>
.
</paragraph>
<paragraph id="2C3F36C4FFED8B03FF30FB72F0C7FAB5" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="596">Female. Protarsomeres 1–4 moderately narrow. Apical margins of abdominal tergite VIII and sternite VIII rounded.</paragraph>
<paragraph id="2C3F36C4FFED8B03FF30FABBF500FA19" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="596">
<emphasis id="1EF4EAD6FFED8B03FF30FABBF18CFAD1" bold="true" box="[199,432,1338,1365]" pageId="15" pageNumber="596">Comparative notes.</emphasis>
<taxonomicName id="EB804D47FFED8B03FE41FABAF2E8FAD0" authorityName="Motschulsky" authorityYear="1860" box="[438,724,1339,1365]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFED8B03FE41FABAF2E8FAD0" box="[438,724,1339,1365]" italics="true" pageId="15" pageNumber="596">Geodromicus transversus</emphasis>
</taxonomicName>
can be distinguished from other species of the 
<taxonomicName id="EB804D47FFED8B03FB28FABAF568FAD1" box="[1247,1364,1339,1365]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="brevicollis">
<emphasis id="1EF4EAD6FFED8B03FB28FABAF568FAD1" box="[1247,1364,1339,1365]" italics="true" pageId="15" pageNumber="596">brevicollis</emphasis>
</taxonomicName>
group by the shape of the slightly more transverse pronotum, with moderately short and widely concave laterobasal margins in front of hind angles, the distinctly broader median lobe and the morphology of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="649A654FFFED8B03FF30FA27F2DDF9A9" pageId="15" pageNumber="596" type="distribution">
<paragraph id="2C3F36C4FFED8B03FF30FA27F3C1FA45" blockId="15.[151,1437,151,2013]" box="[199,1021,1446,1473]" pageId="15" pageNumber="596">
<emphasis id="1EF4EAD6FFED8B03FF30FA27F15DFA44" bold="true" box="[199,353,1446,1472]" pageId="15" pageNumber="596">Distribution.</emphasis>
The species is widely distributed in the Caucasus (
<figureCitation id="B4BB2A41FFED8B03FC6BFA26F3CCFA45" box="[924,1008,1447,1473]" captionStart="FIGURE 24" captionStartId="16.[151,250,1258,1282]" captionTargetBox="[151,1435,309,1235]" captionTargetId="figure-90@16.[151,1435,309,1235]" captionTargetPageId="16" captionText="FIGURE 24. Distribution of Geodromicus transversus in the Caucasus. Black circles: material studied by author, white circles: previously published material not seen by author." figureDoi="http://doi.org/10.5281/zenodo.12189116" httpUri="https://zenodo.org/record/12189116/files/figure.png" pageId="15" pageNumber="596">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="2C3F36C4FFED8B03FF30FA4BF2DDF9A9" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="596">
<emphasis id="1EF4EAD6FFED8B03FF30FA4BF177FA60" bold="true" box="[199,331,1482,1508]" pageId="15" pageNumber="596">Bionomics.</emphasis>
Specimens were collected at elevations from 
<quantity id="EB789B21FFED8B03FCA0FA4AF431FA61" box="[855,1037,1483,1509]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="3.1" metricValueMin="1.2" pageId="15" pageNumber="596" unit="m" value="2150.0" valueMax="3100.0" valueMin="1200.0">1200 to 3100 m</quantity>
a.s.l. and were taken under stones, in gravel and litter near rivers and streams. Some specimens were collected in the subnival zone under stones near snow fields, occasionally near swamps and in flight.
</paragraph>
</subSubSection>
<subSubSection id="649A654FFFED8B1CFF30F9B6F3B0FE99" lastPageId="16" lastPageNumber="597" pageId="15" pageNumber="596" type="discussion">
<paragraph id="2C3F36C4FFED8B1CFF30F9B6F3DFFF7D" blockId="15.[151,1437,151,2013]" lastBlockId="16.[151,1436,151,285]" lastPageId="16" lastPageNumber="597" pageId="15" pageNumber="596">
<emphasis id="1EF4EAD6FFED8B03FF30F9B6F107F9D5" bold="true" box="[199,315,1591,1617]" pageId="15" pageNumber="596">Remarks.</emphasis>
<taxonomicName id="EB804D47FFED8B03FEB4F9B6F272F9D4" box="[323,590,1591,1617]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFED8B03FEB4F9B6F272F9D4" box="[323,590,1591,1617]" italics="true" pageId="15" pageNumber="596">Geodromus transversus</emphasis>
</taxonomicName>
and 
<taxonomicName id="EB804D47FFED8B03FD71F9B6F33DF9D5" authorityName="Motschulsky" authorityYear="1860" box="[646,769,1591,1617]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFED8B03FD71F9B6F33DF9D5" box="[646,769,1591,1617]" italics="true" pageId="15" pageNumber="596">G. flavipes</emphasis>
</taxonomicName>
were originally described based on an unspecified number of 
<typeStatus id="F33B8866FFED8B03FF4FF9DAF126F9F1" box="[184,282,1627,1653]" pageId="15" pageNumber="596" type="syntype">syntypes</typeStatus>
from “...des Alpes du Caucase en Touchétie”. 
<bibRefCitation id="48114B35FFED8B03FCCEF9DAF3ECF9F1" author="Fauvel, A." box="[825,976,1627,1653]" pageId="15" pageNumber="596" pagination="27 - 192" refId="ref16241" refString="Fauvel, A. (1871) Faune Gallo-Rhenane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhenanes et la Valais avec tableaux synoptiques et planches gravees. Bulletin de la Societe Linneenne de Normandie, Series 2, 5 (1869 - 1870), 27 - 192." type="journal article" year="1871">Fauvel (1871</bibRefCitation>
a [=1872]) regarded 
<taxonomicName id="EB804D47FFED8B03FB48F9DAF559F9F0" box="[1215,1381,1627,1653]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFED8B03FB48F9DAF559F9F0" box="[1215,1381,1627,1653]" italics="true" pageId="15" pageNumber="596">G. transversus</emphasis>
</taxonomicName>
as a synonym of 
<taxonomicName id="EB804D47FFED8B03FEDFF9FEF30EF91D" authority="Mannerheim, 1830" authorityName="Mannerheim" authorityYear="1830" box="[296,818,1663,1689]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="globulicollis">
<emphasis id="1EF4EAD6FFED8B03FEDFF9FEF26DF91D" box="[296,593,1663,1689]" italics="true" pageId="15" pageNumber="596">Anthophagus globulicollis</emphasis>
Mannerheim, 1830
</taxonomicName>
(now synonym of 
<taxonomicName id="EB804D47FFED8B03FBF8F9FEF556F91D" authority="(Fabricius, 1798)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1039,1386,1663,1689]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="plagiatus">
<emphasis id="1EF4EAD6FFED8B03FBF8F9FEF4A7F91D" box="[1039,1179,1663,1689]" italics="true" pageId="15" pageNumber="596">G. plagiatus</emphasis>
(Fabricius, 1798)
</taxonomicName>
, see details in 
<bibRefCitation id="48114B35FFED8B03FEF2F922F191F939" author="Shavrin, A. V." box="[261,429,1699,1725]" pageId="15" pageNumber="596" pagination="401 - 429" refId="ref17547" refString="Shavrin, A. V. (2022 a) The plagiatus species group of the genus Geodromicus Redtenbacher, 1857 (Insecta: Coleoptera: Staphylinidae: Omaliinae), and additional data on some species from Japan. Zootaxa, 5188 (5), 401 - 429. https: // doi. org / 10.11646 / zootaxa. 5188.5.1" type="journal article" year="2022">Shavrin 2022a</bibRefCitation>
). 
<bibRefCitation id="48114B35FFED8B03FE37F922F336F939" author="Gemminger, M. &amp; Harold, B. de" box="[448,778,1699,1725]" pageId="15" pageNumber="596" pagination="425 - 752" refId="ref16410" refString="Gemminger, M. &amp; Harold, B. de. (1868) n. k. In: Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom II. Dytiscidae, Gyrinidae, Hydrophilidae, Staphylinidae, Pselaphidae, Gnostidae, Paussidae, Scydmaenidae, Silphidae, Trichopterygidae, Scaphidiidae. E. H. Gummi, Monachii, pp. 425 - 752 + (6). https: // doi. org / 10.5962 / bhl. title. 9089" type="journal article" year="1868">Gemminger &amp; Harold (1868)</bibRefCitation>
and 
<bibRefCitation id="48114B35FFED8B03FCB6F922F3DCF939" author="Fauvel, A." box="[833,992,1699,1725]" pageId="15" pageNumber="596" pagination="479 - 494" refId="ref16207" refString="Fauvel, A. (1869) Remarques synonymiques sur les staphylinides du Catalogus Coleopterorum de MM v. Harold et Gemminger. L'Abeille. Memoires d'Entomologie, 5, 479 - 494." type="journal article" year="1869">Fauvel (1869)</bibRefCitation>
regarded 
<taxonomicName id="EB804D47FFED8B03FBA6F922F4F6F939" authorityName="Motschulsky" authorityYear="1860" box="[1105,1226,1699,1725]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFED8B03FBA6F922F4F6F939" box="[1105,1226,1699,1725]" italics="true" pageId="15" pageNumber="596">G. flavipes</emphasis>
</taxonomicName>
as 
<taxonomicName id="EB804D47FFED8B03FB06F922F5A0F939" authorityName="Mannerheim" authorityYear="1830" box="[1265,1436,1699,1725]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="globulicollis">
<emphasis id="1EF4EAD6FFED8B03FB06F922F5A0F939" box="[1265,1436,1699,1725]" italics="true" pageId="15" pageNumber="596">A. globulicollis</emphasis>
</taxonomicName>
too. 
<bibRefCitation id="48114B35FFED8B03FF3CF946F145F965" author="Bordoni, A." box="[203,377,1735,1761]" pageId="15" pageNumber="596" pagination="157 - 160" refId="ref15928" refString="Bordoni, A. (1993) Note intorno ad alcuni Omaliinae. Atti Museo Civico di Storia Naturale, Grosseto, 15, 157 - 160." type="journal article" year="1993">Bordoni (1993)</bibRefCitation>
cited both species as valid taxa within the genus 
<taxonomicName id="EB804D47FFED8B03FC45F946F527F965" authority="Bordoni, 1985" authorityName="Bordoni" authorityYear="1985" box="[946,1307,1734,1761]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="genus">
<emphasis id="1EF4EAD6FFED8B03FC45F946F453F965" box="[946,1135,1735,1761]" italics="true" pageId="15" pageNumber="596">Neogeodromicus</emphasis>
<bibRefCitation id="48114B35FFED8B03FB8FF946F527F965" author="Bordoni, A." box="[1144,1307,1734,1761]" pageId="15" pageNumber="596" pagination="371 - 375" refId="ref15894" refString="Bordoni, A. (1985) Neogeodromicus n. gen., ketmeniensis n. sp. del Turkestan (Col. Staphylinidae). Redia, 68, 371 - 375." type="journal article" year="1985">Bordoni, 1985</bibRefCitation>
</taxonomicName>
which was originally established for the Middle Asian 
<taxonomicName id="EB804D47FFED8B03FD77F96AF3E0F881" authority="Bordoni, 1985" authorityName="Bordoni" authorityYear="1985" box="[640,988,1770,1797]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="ketmeniensis">
<emphasis id="1EF4EAD6FFED8B03FD77F96AF30CF881" box="[640,816,1771,1797]" italics="true" pageId="15" pageNumber="596">N. ketmeniensis</emphasis>
<bibRefCitation id="48114B35FFED8B03FCC0F96AF3E0F881" author="Bordoni, A." box="[823,988,1770,1797]" pageId="15" pageNumber="596" pagination="371 - 375" refId="ref15894" refString="Bordoni, A. (1985) Neogeodromicus n. gen., ketmeniensis n. sp. del Turkestan (Col. Staphylinidae). Redia, 68, 371 - 375." type="journal article" year="1985">Bordoni, 1985</bibRefCitation>
</taxonomicName>
. Bordoni designated 
<typeStatus id="F33B8866FFED8B03FB3DF96AF500F881" box="[1226,1340,1771,1797]" pageId="15" pageNumber="596" type="lectotype">lectotypes</typeStatus>
for both species, briefly redescribed them, and provided figures of the aedeagi; the author had possibly overlooked 
<bibRefCitation id="48114B35FFED8B03FAB8F88EF0DEF8C9" author="Zerche, L." pageId="15" pageNumber="596" pagination="105 - 142" refId="ref17790" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche (1992)</bibRefCitation>
who synonymized 
<taxonomicName id="EB804D47FFED8B03FE32F8B2F2BEF8C9" authorityName="Bordoni" authorityYear="1985" box="[453,642,1843,1869]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="genus">
<emphasis id="1EF4EAD6FFED8B03FE32F8B2F2BEF8C9" box="[453,642,1843,1869]" italics="true" pageId="15" pageNumber="596">Neogeodromicus</emphasis>
</taxonomicName>
with 
<taxonomicName id="EB804D47FFED8B03FD3FF8B2F3C2F8C9" authority="Luze, 1903" authorityName="Luze" authorityYear="1903" box="[712,1022,1843,1869]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="genus">
<emphasis id="1EF4EAD6FFED8B03FD3FF8B2F34DF8C9" box="[712,881,1843,1869]" italics="true" pageId="15" pageNumber="596">Trichodromeus</emphasis>
<bibRefCitation id="48114B35FFED8B03FC8DF8B2F3C2F8C9" author="Luze" box="[890,1022,1843,1869]" pageId="15" pageNumber="596" pagination="103 - 117" refId="ref17205" refString="Luze G. 1903. Revision der palaarktischen Arten der Staphyliniden-gattung Geodromicus Redtenb. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 53, 103 - 117." type="journal article" year="1903">Luze, 1903</bibRefCitation>
</taxonomicName>
. Thus, 
<taxonomicName id="EB804D47FFED8B03FBA3F8B2F5A0F8C8" box="[1108,1436,1843,1869]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFED8B03FBA3F8B2F5A0F8C8" box="[1108,1436,1843,1869]" italics="true" pageId="15" pageNumber="596">Neogeodromicus transversus</emphasis>
</taxonomicName>
and 
<taxonomicName id="EB804D47FFED8B03FF30F8D6F102F8F5" authorityName="Motschulsky" authorityYear="1860" box="[199,318,1879,1905]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFED8B03FF30F8D6F102F8F5" box="[199,318,1879,1905]" italics="true" pageId="15" pageNumber="596">N. flavipes</emphasis>
</taxonomicName>
were formally transferred to the latter genus. Based on the study of the type and additional material of 
<taxonomicName id="EB804D47FFED8B03FF43F8FAF16AF810" box="[180,342,1915,1941]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFED8B03FF43F8FAF16AF810" box="[180,342,1915,1941]" italics="true" pageId="15" pageNumber="596">G. transversus</emphasis>
</taxonomicName>
and 
<taxonomicName id="EB804D47FFED8B03FE7CF8FAF23FF811" authorityName="Motschulsky" authorityYear="1860" box="[395,515,1915,1941]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFED8B03FE7CF8FAF23FF811" box="[395,515,1915,1941]" italics="true" pageId="15" pageNumber="596">G. flavipes</emphasis>
</taxonomicName>
, I found that these species undoubtedly belong to the 
<taxonomicName id="EB804D47FFED8B03FBA4F8FAF4F4F811" box="[1107,1224,1915,1941]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="brevicollis">
<emphasis id="1EF4EAD6FFED8B03FBA4F8FAF4F4F811" box="[1107,1224,1915,1941]" italics="true" pageId="15" pageNumber="596">brevicollis</emphasis>
</taxonomicName>
group of the genus 
<taxonomicName id="EB804D47FFED8B03FF60F81EF111F83D" authorityName="Redtenbacher" authorityYear="1857" box="[151,301,1951,1977]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="genus">
<emphasis id="1EF4EAD6FFED8B03FF60F81EF111F83D" box="[151,301,1951,1977]" italics="true" pageId="15" pageNumber="596">Geodromicus</emphasis>
</taxonomicName>
. Species of the genus 
<taxonomicName id="EB804D47FFED8B03FDD0F81EF2ECF83D" authorityName="Luze" authorityYear="1903" box="[551,720,1951,1977]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="genus">
<emphasis id="1EF4EAD6FFED8B03FDD0F81EF2ECF83D" box="[551,720,1951,1977]" italics="true" pageId="15" pageNumber="596">Trichodromeus</emphasis>
</taxonomicName>
which are widely distributed in Middle Asia and the Himalayan Region are completely different in the morphology of the habitus, mouthparts and the aedeagus (e.g. 
<bibRefCitation id="48114B35FFED8B03FAFFF842F5A4F859" author="Zerche, L." box="[1288,1432,1987,2013]" pageId="15" pageNumber="596" pagination="105 - 142" refId="ref17790" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche 1992</bibRefCitation>
, 
<bibRefCitation id="48114B35FFF28B1CFF60FF16F17FFF35" author="Shavrin, A. V." box="[151,323,151,177]" pageId="16" pageNumber="597" pagination="353 - 362" refId="ref17490" refString="Shavrin, A. V. (2021 b) Four new species of the genus Trichodromeus Luze, 1903 from Nepal and Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 4991 (2), 353 - 362. https: // doi. org / 10.11646 / zootaxa. 4991.2.8" type="journal article" year="2021">Shavrin 2021b</bibRefCitation>
). Based on the study of external and internal morphological features, I could not find sufficient differences between 
<taxonomicName id="EB804D47FFF28B1CFE8AFF3AF223FF50" box="[381,543,187,213]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="597" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFF28B1CFE8AFF3AF223FF50" box="[381,543,187,213]" italics="true" pageId="16" pageNumber="597">G. transversus</emphasis>
</taxonomicName>
and 
<taxonomicName id="EB804D47FFF28B1CFDA3FF3AF2F1FF51" authorityName="Motschulsky" authorityYear="1860" box="[596,717,187,213]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="597" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFF28B1CFDA3FF3AF2F1FF51" box="[596,717,187,213]" italics="true" pageId="16" pageNumber="597">G. flavipes</emphasis>
</taxonomicName>
. Thus, I synonymize 
<taxonomicName id="EB804D47FFF28B1CFC4BFF3AF409FF51" authorityName="Motschulsky" authorityYear="1860" box="[956,1077,187,213]" class="Insecta" family="Carabidae" genus="Geodromus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="597" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="1EF4EAD6FFF28B1CFC4BFF3AF409FF51" box="[956,1077,187,213]" italics="true" pageId="16" pageNumber="597">G. flavipes</emphasis>
</taxonomicName>
with 
<taxonomicName id="EB804D47FFF28B1CFB84FF3AF52AFF50" box="[1139,1302,187,213]" class="Insecta" family="Staphylinidae" genus="Neogeodromicus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="597" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="1EF4EAD6FFF28B1CFB84FF3AF52AFF50" box="[1139,1302,187,213]" italics="true" pageId="16" pageNumber="597">G. transversus</emphasis>
</taxonomicName>
, the species which has page priority (page 550) in the same article (
<bibRefCitation id="48114B35FFF28B1CFCF7FF5EF3EAFF7D" author="Motschulsky, V. de" box="[768,982,223,249]" pageId="16" pageNumber="597" pagination="539 - 588" refId="ref17233" refString="Motschulsky, V. de. (1860) Enumeration des nouvelles especes de coleopteres rapportees de ses voyages. 3 - ieme article. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 33 (2), 539 - 588." type="journal article" year="1860">Motschulsky 1860</bibRefCitation>
).
</paragraph>
<paragraph id="2C3F36C4FFF28B1CFF30FE82F3B0FE99" blockId="16.[151,1436,151,285]" box="[199,908,259,285]" pageId="16" pageNumber="597">
It is here recorded from 
<collectingCountry id="54977654FFF28B1CFE20FE82F20DFE99" box="[471,561,259,285]" name="Georgia" pageId="16" pageNumber="597">Georgia</collectingCountry>
and 
<collectingCountry id="54977654FFF28B1CFD92FE82F2FBFE99" box="[613,711,259,285]" name="Armenia" pageId="16" pageNumber="597">Armenia</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>