281 lines
38 KiB
XML
281 lines
38 KiB
XML
<document id="3CE04FA33E5776A6AB8B59DEEA89D3A8" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFDA6D6ED6DDDC58BF37078D" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Xanthonomus guineensis Shavrin 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="305" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="302" updateTime="1710501296679" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DF602ECB424C5CF6AFBDABBA1CFE7051">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<mods:namePart id="E4C9D45615977325156216E65B4AAF19">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="3978D8444EC67520E83796CF5C54FAE6">Zootaxa</mods:title>
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<mods:part id="04CDEEAF53E14DD40F53FC70B81B4D4C">
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<mods:date id="53104E5BC784406A13B3190C5864819C">2024</mods:date>
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<mods:number id="C5B74017874114589A74588E4BE42681">2024-03-14</mods:number>
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<mods:number id="CD21EE7A3247C2CB29E5C5BEDF6CD917">5424</mods:number>
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<mods:classification id="BCC5B16BD618EBFE9219CE17FBAFFCBF">journal article</mods:classification>
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<mods:identifier id="A8EC573848C79A9736A99BBEE7F5139D" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
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<mods:identifier id="5FE1EB2DF459D99F75523296C88019B5" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A15C878DFFDA6D6ED6DDDC58BF37078D" ID-DOI="http://doi.org/10.5281/zenodo.10821321" ID-Zenodo-Dep="10821321" LSID="urn:lsid:plazi:treatment:A15C878DFFDA6D6ED6DDDC58BF37078D" httpUri="http://treatment.plazi.org/id/A15C878DFFDA6D6ED6DDDC58BF37078D" lastPageId="54" lastPageNumber="305" pageId="51" pageNumber="302">
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<subSubSection id="61EF6510FFDA6D6BD6DDDC58BC1303C9" box="[151,532,1391,1418]" pageId="51" pageNumber="302" type="nomenclature">
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<paragraph id="294A369BFFDA6D6BD6DDDC58BC1303C9" blockId="51.[151,532,1391,1455]" box="[151,532,1391,1418]" pageId="51" pageNumber="302">
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<heading id="720281F7FFDA6D6BD6DDDC58BC1303C9" bold="true" box="[151,532,1391,1418]" fontSize="11" level="1" pageId="51" pageNumber="302" reason="1">
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<emphasis id="1B81EA89FFDA6D6BD6DDDC58BC1303C9" bold="true" box="[151,532,1391,1418]" pageId="51" pageNumber="302">
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<taxonomicName id="EEF54D18FFDA6D6BD6DDDC58BFCC03C9" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,459,1391,1418]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="guineensis" status="sp. nov.">
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<emphasis id="1B81EA89FFDA6D6BD6DDDC58BFCC03C9" bold="true" box="[151,459,1391,1418]" italics="true" pageId="51" pageNumber="302">Xanthonomus guineensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="00B257F2FFDA6D6BD798DC47BC1303C9" box="[466,532,1392,1418]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="61EF6510FFDA6D6BD6DDDCA3BF7C03EC" box="[151,379,1428,1455]" pageId="51" pageNumber="302" type="description">
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<paragraph id="294A369BFFDA6D6BD6DDDCA3BF7C03EC" blockId="51.[151,532,1391,1455]" box="[151,379,1428,1455]" pageId="51" pageNumber="302">
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(
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<figureCitation id="B1CE2A1EFFDA6D6BD6D5DCA3BF0303ED" box="[159,260,1428,1454]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="51" pageNumber="302">Figs 142</figureCitation>
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,
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<figureCitation id="B1CE2A1EFFDA6D6BD75ADCA3BF7403EC" box="[272,371,1428,1455]" captionStart-0="FIGURES 152–157" captionStart-1="FIGURE 158" captionStartId-0="52.[151,264,1589,1613]" captionStartId-1="53.[151,250,1956,1980]" captionTargetBox-0="[185,1403,415,1547]" captionTargetBox-1="[151,1436,1165,1932]" captionTargetId-0="figure-117@52.[177,1411,415,1566]" captionTargetId-1="figure-421@53.[151,1436,1165,1932]" captionTargetPageId-0="52" captionTargetPageId-1="53" captionText-0="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." captionText-1="FIGURE 158. Distribution of Xanthonomus: X. toxopeanus (circle), X. latus (triangle), X. guineensis (squares), X. thayerae (diamonds)." figureDoi-0="http://doi.org/10.5281/zenodo.10821297" figureDoi-1="http://doi.org/10.5281/zenodo.10821299" httpUri-0="https://zenodo.org/record/10821297/files/figure.png" httpUri-1="https://zenodo.org/record/10821299/files/figure.png" pageId="51" pageNumber="302">152–158</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="61EF6510FFDA6D6BD6DDDCEBBD20008D" pageId="51" pageNumber="302" type="materials_examined">
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<paragraph id="294A369BFFDA6D6BD6DDDCEBBEFE007D" blockId="51.[151,1437,1500,2030]" pageId="51" pageNumber="302">
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<emphasis id="1B81EA89FFDA6D6BD6DDDCEBBF4003B5" bold="true" box="[151,327,1500,1526]" pageId="51" pageNumber="302">Type material.</emphasis>
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<typeStatus id="F64E8839FFDA6D6BD71BDCEBBFBE03B5" box="[337,441,1500,1526]" pageId="51" pageNumber="302" type="holotype">Holotype</typeStatus>
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<specimenCount id="3FF3FD12FFDA6D6BD788DCEBBFD003B5" box="[450,471,1500,1526]" count="1" pageId="51" pageNumber="302" type="male">♂</specimenCount>
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(
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<figureCitation id="B1CE2A1EFFDA6D6BD7A3DCEBBC4803B5" box="[489,591,1500,1526]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="51" pageNumber="302">Fig. 142</figureCitation>
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): ‘
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<collectingRegion id="EB31F879FFDA6D6BD423DCEBBD0B03B5" box="[617,780,1500,1526]" country="Indonesia" name="Papua Barat" pageId="51" pageNumber="302">IRIAN JAYA</collectingRegion>
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: Paniai Prov. | Bilogay,
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<date id="5D4B105BFFDA6D6BD268DCEBBAB403B4" box="[1058,1203,1500,1527]" pageId="51" pageNumber="302" value="1995-12-29">29. 12. 1995</date>
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<quantity id="EE0D9B7EFFDA6D6BD281DCEBBB5D03B5" box="[1227,1370,1500,1526]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="51" pageNumber="302" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100-2200m</quantity>
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| leg. A. Riedel’ <printed>, ‘
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<typeStatus id="F64E8839FFDA6D6BD7D1DF37BC330059" box="[411,564,1536,1562]" pageId="51" pageNumber="302" type="holotype">HOLOTYPE</typeStatus>
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|
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<taxonomicName id="EEF54D18FFDA6D6BD407DF37BCE00059" authorityName="Bernhauer" authorityYear="1926" box="[589,743,1536,1562]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="genus">
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<emphasis id="1B81EA89FFDA6D6BD407DF37BCE00059" box="[589,743,1536,1562]" italics="true" pageId="51" pageNumber="302">Xanthonomus</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="EEF54D18FFDA6D6BD54BDF36BD700059" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[769,887,1537,1562]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="guineensis" status="sp. nov.">
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<emphasis id="1B81EA89FFDA6D6BD54BDF36BD700059" box="[769,887,1537,1562]" italics="true" pageId="51" pageNumber="302">guineensis</emphasis>
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</taxonomicName>
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||
<emphasis id="1B81EA89FFDA6D6BD5CBDF37BDC30059" bold="true" box="[897,964,1536,1562]" pageId="51" pageNumber="302">
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<taxonomicNameLabel id="00B257F2FFDA6D6BD5CBDF37BDC30059" box="[897,964,1536,1562]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.
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||
<collectionCode id="4FE4AE5EFFDA6D6BD21CDF37BA6B0059" box="[1110,1132,1536,1562]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="51" pageNumber="302" type="Museum">V</collectionCode>
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||
. des. 2024’ <red, printed> (
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||
<collectionCode id="4FE4AE5EFFDA6D6BD6D5DF13BEEA007D" box="[159,237,1572,1598]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="51" pageNumber="302">NMW</collectionCode>
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).
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</paragraph>
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<paragraph id="294A369BFFDA6D6BD68DDF7FBD20008D" blockId="51.[151,1437,1500,2030]" pageId="51" pageNumber="302">
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<typeStatus id="F64E8839FFDA6D6BD68DDF7FBF3F0021" box="[199,312,1608,1634]" pageId="51" pageNumber="302" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="3FF3FD12FFDA6D6BD70BDF7FBF780021" box="[321,383,1608,1635]" count="5" pageId="51" pageNumber="302" type="male">5 ♂♂</specimenCount>
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(
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<specimenCount id="3FF3FD12FFDA6D6BD7C6DF7FBC210021" box="[396,550,1608,1634]" count="1" pageId="51" pageNumber="302" type="generic">one specimen</specimenCount>
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dissected),
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<specimenCount id="3FF3FD12FFDA6D6BD4EDDF7FBCE20021" box="[679,741,1608,1634]" count="3" pageId="51" pageNumber="302" type="female">3 ♀♀</specimenCount>
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(
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<specimenCount id="3FF3FD12FFDA6D6BD4B8DF7FBD8B0021" box="[754,908,1608,1634]" count="1" pageId="51" pageNumber="302" type="generic">one specimen</specimenCount>
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dissected): same data as the holotype (
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<specimenCount id="3FF3FD12FFDA6D6BD373DF7FBB660021" box="[1337,1377,1608,1634]" count="1" pageId="51" pageNumber="302" type="male">1 ♂</specimenCount>
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,
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<specimenCount id="3FF3FD12FFDA6D6BD326DF7FBB910021" box="[1388,1430,1608,1634]" count="1" pageId="51" pageNumber="302" type="female">1 ♀</specimenCount>
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: cSh;
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<specimenCount id="3FF3FD12FFDA6D6BD69ADF5BBF1600C5" box="[208,273,1644,1670]" count="4" pageId="51" pageNumber="302" type="male">4 ♂♂</specimenCount>
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,
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<specimenCount id="3FF3FD12FFDA6D6BD757DF5BBF5900C5" box="[285,350,1644,1670]" count="2" pageId="51" pageNumber="302" type="female">2 ♀♀</specimenCount>
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||
:
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<collectionCode id="4FE4AE5EFFDA6D6BD721DF5BBFBF00C5" box="[363,440,1644,1670]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="51" pageNumber="302">NMW</collectionCode>
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);
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||
<specimenCount id="3FF3FD12FFDA6D6BD781DF5BBC0B00C5" box="[459,524,1644,1670]" count="3" pageId="51" pageNumber="302" type="male">3 ♂♂</specimenCount>
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||
,
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<specimenCount id="3FF3FD12FFDA6D6BD452DF5BBC4500C5" box="[536,578,1644,1670]" count="1" pageId="51" pageNumber="302" type="female">1 ♀</specimenCount>
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||
(dissected): ‘
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||
<collectingRegion id="EB31F879FFDA6D6BD49CDF5BBD7000C5" box="[726,887,1644,1670]" country="Indonesia" name="Papua Barat" pageId="51" pageNumber="302">IRIAN JAYA</collectingRegion>
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||
: Paniai Prov. | Sinak, trail to Ilaga |
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||
<quantity id="EE0D9B7EFFDA6D6BD34CDF5BBB9F00C4" box="[1286,1432,1644,1671]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="51" pageNumber="302" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">2100-2500m</quantity>
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||
,
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||
<date id="5D4B105BFFDA6D6BD6DDDFA7BF2F00E8" box="[151,296,1680,1707]" pageId="51" pageNumber="302" value="1995-12-15">15. 12. 1995</date>
|
||
| leg. A. Riedel’ <printed> (
|
||
<collectionCode id="4FE4AE5EFFDA6D6BD438DFA7BCC700E9" box="[626,704,1680,1706]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="51" pageNumber="302">NMW</collectionCode>
|
||
). All
|
||
<typeStatus id="F64E8839FFDA6D6BD54BDFA7BD6B00E9" box="[769,876,1680,1706]" pageId="51" pageNumber="302" type="paratype">paratypes</typeStatus>
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||
with additional red printed label: ‘
|
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<typeStatus id="F64E8839FFDA6D6BD2B7DFA7BB8A00E9" box="[1277,1421,1680,1706]" pageId="51" pageNumber="302" type="paratype">PARATYPE</typeStatus>
|
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|
|
||
<taxonomicName id="EEF54D18FFDA6D6BD6DDDF83BF36008D" authorityName="Bernhauer" authorityYear="1926" box="[151,305,1716,1742]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1B81EA89FFDA6D6BD6DDDF83BF36008D" box="[151,305,1716,1742]" italics="true" pageId="51" pageNumber="302">Xanthonomus</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="EEF54D18FFDA6D6BD70FDF82BFBC008D" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[325,443,1717,1742]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="guineensis" status="sp. nov.">
|
||
<emphasis id="1B81EA89FFDA6D6BD70FDF82BFBC008D" box="[325,443,1717,1742]" italics="true" pageId="51" pageNumber="302">guineensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFDA6D6BD788DF83BC06008D" bold="true" box="[450,513,1716,1742]" pageId="51" pageNumber="302">
|
||
<taxonomicNameLabel id="00B257F2FFDA6D6BD788DF83BC06008D" box="[450,513,1716,1742]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
| Shavrin A.
|
||
<collectionCode id="4FE4AE5EFFDA6D6BD4C1DF83BCA6008D" box="[651,673,1716,1742]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="51" pageNumber="302" type="Museum">V</collectionCode>
|
||
. des. 2024’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61EF6510FFDA6D6ED68DDFEFBF1506BB" lastPageId="54" lastPageNumber="305" pageId="51" pageNumber="302" type="description">
|
||
<paragraph id="294A369BFFDA6D6BD68DDFEFBF8C011D" blockId="51.[151,1437,1500,2030]" pageId="51" pageNumber="302">
|
||
<emphasis id="1B81EA89FFDA6D6BD68DDFEFBF5E00B1" bold="true" box="[199,345,1752,1778]" pageId="51" pageNumber="302">Description.</emphasis>
|
||
Measurements (n=13): HW: 0.40–0.46; HL: 0.20–0.25; OL: 0.12–0.14; TL: 0.02–0.03; AL (
|
||
<typeStatus id="F64E8839FFDA6D6BD6D5DFCBBF0F0155" box="[159,264,1788,1814]" pageId="51" pageNumber="302" type="holotype">holotype</typeStatus>
|
||
): 0.52; PL: 0.29–0.38; PWmax: 0.53–0.57; PWmin: 0.49–0.52; ESL: 0.45–0.52; EW: 0.66–0.73; MTbL (
|
||
<typeStatus id="F64E8839FFDA6D6BD6D5DE17BF0F0179" box="[159,264,1824,1850]" pageId="51" pageNumber="302" type="holotype">holotype</typeStatus>
|
||
): 0.35; MTrL (
|
||
<typeStatus id="F64E8839FFDA6D6BD7E4DE17BC100179" box="[430,535,1824,1850]" pageId="51" pageNumber="302" type="holotype">holotype</typeStatus>
|
||
): 0.24 (MTrL 1–4: 0.07; MTrL 5: 0.17); AW: 0.75–0.80; AedL: 0.45; BL: 1.87– 2.25 (
|
||
<typeStatus id="F64E8839FFDA6D6BD69DDE73BF38011D" box="[215,319,1860,1886]" pageId="51" pageNumber="302" type="holotype">holotype</typeStatus>
|
||
: 2.04).
|
||
</paragraph>
|
||
<paragraph id="294A369BFFDA6D6CD68DDE5FBF9007C4" blockId="51.[151,1437,1500,2030]" lastBlockId="52.[151,1437,151,391]" lastPageId="52" lastPageNumber="303" pageId="51" pageNumber="302">
|
||
Habitus as in
|
||
<figureCitation id="B1CE2A1EFFDA6D6BD714DE5FBFB901C1" box="[350,446,1896,1922]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="51" pageNumber="302">Fig. 142</figureCitation>
|
||
. Body dark brown, with slightly paler elytra and sometimes abdominal tergites VII–VIII; antennomeres 7–11 and femora brown; tibia yellow-brown, with paler apical portion; mouthparts, antennomeres 1–6 and tarsi yellow. Head with very fine, indistinct and dense punctation, finer and more indistinct on infraorbital portions; neck without visible punctures; pronotum with very dense and fine punctation, missing in mediobasal portion; scutellum without punctures; elytra with very fine and dense punctation similar to that in pronotum, usually finer and sparser around scutellum and along suture; abdomen with very dense and fine punctation. Head with dense and coarse microreticulation, transverse on clypeus and isodiametric in middle; neck with moderately fine transverse microreticulation; pronotum with very dense and fine isodiametric meshes about as that in middle portion of head; scutellum with dense and coarse transverse microsculpture; elytra with fine and sometimes indistinct isodiametric microreticulation, sometimes indistinct along suture; abdomen with very dense and moderately coarse isodiametric sculpture.
|
||
</paragraph>
|
||
<caption id="7D8A6613FFDD6D6CD6DDDF02BFD800D6" ID-DOI="http://doi.org/10.5281/zenodo.10821297" ID-Zenodo-Dep="10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="52" pageNumber="303" startId="52.[151,264,1589,1613]" targetBox="[185,1403,415,1547]" targetPageId="52" targetType="figure">
|
||
<paragraph id="294A369BFFDD6D6CD6DDDF02BFD800D6" blockId="52.[151,1437,1589,1686]" pageId="52" pageNumber="303">
|
||
<emphasis id="1B81EA89FFDD6D6CD6DDDF02BF68000E" bold="true" box="[151,367,1589,1613]" pageId="52" pageNumber="303">FIGURES 152–157.</emphasis>
|
||
<taxonomicName id="EEF54D18FFDD6D6CD73DDF02BC72000E" authorityName="Shavrin" authorityYear="2024" box="[375,629,1589,1613]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="303" phylum="Arthropoda" rank="species" species="guineensis">
|
||
<emphasis id="1B81EA89FFDD6D6CD73DDF02BC72000E" box="[375,629,1589,1613]" italics="true" pageId="52" pageNumber="303">Xanthonomus guineensis</emphasis>
|
||
</taxonomicName>
|
||
(paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="294A369BFFDD6D6DD68DDFD0BFE907CB" blockId="52.[151,1436,1767,2044]" lastBlockId="53.[151,1437,151,1142]" lastPageId="53" lastPageNumber="304" pageId="52" pageNumber="303">
|
||
Head distinctly transverse, 1.8 times to twice as broad as long, with slightly elevated infraorbital portions and shallow oval anteriomedian depressions, indistinct in some
|
||
<typeStatus id="F64E8839FFDD6D6CD524DE3CBDDA0166" box="[878,989,1803,1829]" pageId="52" pageNumber="303" type="paratype">paratypes</typeStatus>
|
||
; anteocellar foveae very narrow, linear, reaching level of apical third or about middle length of eyes; postocular parts very short, from indistinct and obtuse postocular carina strongly narrowed posteriad. Eyes very large, convex. Ocelli moderately large, located slightly below level of postocular carina; distance between ocelli slightly longer or about as long as distance between ocellus and posterior margin of eye. Maxillary palpi long; preapical palpomere about twice as long as wide; apical palpomere narrow, slightly longer than preapical segment, from preapical portion gradually narrowed apicad toward rounded or subacute apex. Antennomeres short, reaching basal third of pronotum when reclined, with distinctly transverse antennomere 7 and strongly transverse antennomeres 8–10; antennomeres 6–10 gradually widened apicad; four apical antennomere forming distinct loose club; basal antennomere wide, slightly more than two and a half times as long as wide, antennomere 2 about twice shorter than basal antennomere, 3 slightly shorter and about twice narrower than 2, 4 about twice shorter than 3, 5 slightly longer and distinctly broader than 4, 6 distinctly shorter and broader than 5, 7 distinctly shorter and broader than 6, 8 three times broader than 6, 9–10 about twice longer and distinctly broader than 8, apical antennomere about 1.3–1.5 times as long as 10, from middle gradually rounded toward rounded apex.
|
||
</paragraph>
|
||
<paragraph id="294A369BFFDC6D6DD68DD8A6BCB90455" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">Pronotum slightly convex, 1.5–1.8 times as broad as long, distinctly broader than head, widest in middle, slightly more narrowed posteriad than anteriad; apical and hind angles widely rounded; anterior margin straight, about as long as rounded posterior margin; median part without longitudinal impressions; lateral portions narrowly bordered; laterobasal portion without depression.</paragraph>
|
||
<paragraph id="294A369BFFDC6D6DD68DDB17BC8F04C2" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">Elytra 1.4 times as long as broad, short, reaching basal or middle portion of abdominal tergite III, 1.3–1.5 times as long as pronotum, slightly broadened posteriad, with slightly concave lateral margins and striagth or rounded apical margins. Hind wings fully developed.</paragraph>
|
||
<paragraph id="294A369BFFDC6D6DD68DDBBCBF89048A" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">Metatarsus about 1.4 times as long as metatibia; ventral surface of protarsomeres 1–4 with several rows of modified tenent setae.</paragraph>
|
||
<paragraph id="294A369BFFDC6D6DD68DDBE5BCB40577" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">Abdomen slightly broadened toward andominal tergite IV or V, distinctly broadened than elytra, with two very small and sometimes indistinct oval wing-folding patches in middle of abdominal tergite IV and narrow palisade fringe at apical margin of abdominal tergite VII.</paragraph>
|
||
<paragraph id="294A369BFFDC6D6DD68DDA0ABD43026C" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">
|
||
Male. Femora very wide; tibia significantly broadened in middle; tarsomeres 1–4 slightly broadened, with very long tenent setae. Posterior margin of abdominal tergite VIII slightly sinuate (
|
||
<figureCitation id="B1CE2A1EFFDC6D6DD20DDA56BAAC053F" box="[1095,1195,865,892]" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 154</figureCitation>
|
||
). Posterior margin of abdominal sternite VIII slighly sinuate, with row of very short setae on apical margin (
|
||
<figureCitation id="B1CE2A1EFFDC6D6DD23FDAB2BADE05E3" box="[1141,1241,901,928]" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 155</figureCitation>
|
||
). Aedeagus long, with very wide basal portion, gradually narrowed toward small and rounded apex; parameres narrow, distinctly not reaching apex of median lobe, with two long apical and two preapical setae; internal sac long and moderately wide, with two very long, narrow and curved sclerotized structure, stretching from basal to about middle part of median lobe (
|
||
<figureCitation id="B1CE2A1EFFDC6D6DD69DDD23BF3E026C" box="[215,313,1044,1071]" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 152</figureCitation>
|
||
). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="B1CE2A1EFFDC6D6DD4AADD23BD47026C" box="[736,832,1044,1071]" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 153</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="294A369BFFDC6D6DD68DDD0FBABD0236" blockId="53.[151,1437,151,1142]" pageId="53" pageNumber="304">
|
||
Female. Femora, tibia and tarsomeres 1–4 moderately narrow. Posterior margin of abdominal tergite VIII (
|
||
<figureCitation id="B1CE2A1EFFDC6D6DD325DD0FBEC50235" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 156</figureCitation>
|
||
) and sternite VIII (
|
||
<figureCitation id="B1CE2A1EFFDC6D6DD7D3DD6CBFFE0235" box="[409,505,1115,1142]" captionStart="FIGURES 152–157" captionStartId="52.[151,264,1589,1613]" captionTargetBox="[185,1403,415,1547]" captionTargetId="figure-117@52.[177,1411,415,1566]" captionTargetPageId="52" captionText="FIGURES 152–157. Xanthonomus guineensis (paratypes): 152–aedeagus, parameral view, 153–aedeagus, lateral view, 154– male abdominal tergite VIII, 155–male abdominal sternite VIII, 156–female abdominal tergite VIII, 157–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821297" httpUri="https://zenodo.org/record/10821297/files/figure.png" pageId="53" pageNumber="304">Fig. 157</figureCitation>
|
||
) trunctate. Accessory sclerite and spermatheca not recognized.
|
||
</paragraph>
|
||
<caption id="7D8A6613FFDC6D6DD6DDDE93BF1501A2" ID-DOI="http://doi.org/10.5281/zenodo.10821299" ID-Zenodo-Dep="10821299" httpUri="https://zenodo.org/record/10821299/files/figure.png" pageId="53" pageNumber="304" startId="53.[151,250,1956,1980]" targetBox="[151,1436,1165,1932]" targetPageId="53" targetType="figure">
|
||
<paragraph id="294A369BFFDC6D6DD6DDDE93BF1501A2" blockId="53.[151,1437,1956,2017]" pageId="53" pageNumber="304">
|
||
<emphasis id="1B81EA89FFDC6D6DD6DDDE93BF2801FF" bold="true" box="[151,303,1956,1980]" pageId="53" pageNumber="304">FIGURE 158.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="EEF54D18FFDC6D6DD79DDE92BC6501FE" authorityName="Bernhauer" authorityYear="1926" box="[471,610,1957,1981]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="304" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1B81EA89FFDC6D6DD79DDE92BC6501FE" box="[471,610,1957,1981]" italics="true" pageId="53" pageNumber="304">Xanthonomus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="EEF54D18FFDC6D6DD438DE92BD0401FF" authorityName="Bernhauer" authorityYear="1926" box="[626,771,1957,1980]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="304" phylum="Arthropoda" rank="species" species="toxopeanus">
|
||
<emphasis id="1B81EA89FFDC6D6DD438DE92BD0401FF" box="[626,771,1957,1980]" italics="true" pageId="53" pageNumber="304">X. toxopeanus</emphasis>
|
||
</taxonomicName>
|
||
(circle),
|
||
<taxonomicName id="EEF54D18FFDC6D6DD52EDE92BDB401FE" authorityName="Steel" authorityYear="1955" box="[868,947,1957,1981]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="304" phylum="Arthropoda" rank="species" species="latus">
|
||
<emphasis id="1B81EA89FFDC6D6DD52EDE92BDB401FE" box="[868,947,1957,1981]" italics="true" pageId="53" pageNumber="304">X. latus</emphasis>
|
||
</taxonomicName>
|
||
(triangle),
|
||
<taxonomicName id="EEF54D18FFDC6D6DD262DE92BAB501FF" authorityName="Shavrin" authorityYear="2024" box="[1064,1202,1957,1980]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="304" phylum="Arthropoda" rank="species" species="guineensis">
|
||
<emphasis id="1B81EA89FFDC6D6DD262DE92BAB501FF" box="[1064,1202,1957,1980]" italics="true" pageId="53" pageNumber="304">X. guineensis</emphasis>
|
||
</taxonomicName>
|
||
(squares),
|
||
<taxonomicName id="EEF54D18FFDC6D6DD36FDE92BB9A01FE" authorityName="Shavrin" authorityYear="2024" box="[1317,1437,1957,1981]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="304" phylum="Arthropoda" rank="species" species="thayerae">
|
||
<emphasis id="1B81EA89FFDC6D6DD36FDE92BB9A01FE" box="[1317,1437,1957,1981]" italics="true" pageId="53" pageNumber="304">X. thayerae</emphasis>
|
||
</taxonomicName>
|
||
(diamonds).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="294A369BFFDF6D6ED68DD9A1BF1506BB" blockId="54.[151,1437,150,463]" pageId="54" pageNumber="305">
|
||
<emphasis id="1B81EA89FFDF6D6ED68DD9A1BFB506F3" bold="true" box="[199,434,150,176]" pageId="54" pageNumber="305">Comparative notes.</emphasis>
|
||
Based on the general shape and sculpture of the body
|
||
<taxonomicName id="EEF54D18FFDF6D6ED260D9A0BAC606F3" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1066,1217,151,176]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="guineensis" status="sp. n.">
|
||
<emphasis id="1B81EA89FFDF6D6ED260D9A0BAC606F3" box="[1066,1217,151,176]" italics="true" pageId="54" pageNumber="305">X. guineensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFDF6D6ED280D9A1BB0B06F3" bold="true" box="[1226,1292,150,176]" pageId="54" pageNumber="305">
|
||
<taxonomicNameLabel id="00B257F2FFDF6D6ED280D9A1BB0B06F3" box="[1226,1292,150,176]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="EEF54D18FFDF6D6ED6DDD98CBF1D0697" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,282,186,212]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="thayerae" status="sp. n.">
|
||
<emphasis id="1B81EA89FFDF6D6ED6DDD98CBF1D0697" box="[151,282,186,212]" italics="true" pageId="54" pageNumber="305">X. thayerae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFDF6D6ED768D98DBF640697" bold="true" box="[290,355,186,212]" pageId="54" pageNumber="305">
|
||
<taxonomicNameLabel id="00B257F2FFDF6D6ED768D98DBF640697" box="[290,355,186,212]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, from which it can be distinguished by the slightly larger body, wider median lobe and shorter parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61EF6510FFDF6D6ED68DD836BF850703" pageId="54" pageNumber="305" type="distribution">
|
||
<paragraph id="294A369BFFDF6D6ED68DD836BF850703" blockId="54.[151,1437,150,463]" pageId="54" pageNumber="305">
|
||
<emphasis id="1B81EA89FFDF6D6ED68DD836BF660758" bold="true" box="[199,353,257,283]" pageId="54" pageNumber="305">Distribution.</emphasis>
|
||
<taxonomicName id="EEF54D18FFDF6D6ED720D836BC840758" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[362,643,257,283]" class="Insecta" family="Staphylinidae" genus="Xanthonomus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="305" phylum="Arthropoda" rank="species" species="guineensis" status="sp. nov.">
|
||
<emphasis id="1B81EA89FFDF6D6ED720D836BC840758" box="[362,643,257,283]" italics="true" pageId="54" pageNumber="305">Xanthonomus guineensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFDF6D6ED4C6D835BCC9075F" bold="true" box="[652,718,258,284]" pageId="54" pageNumber="305">
|
||
<taxonomicNameLabel id="00B257F2FFDF6D6ED4C6D835BCC9075F" box="[652,718,258,284]" pageId="54" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from the
|
||
<typeStatus id="F64E8839FFDF6D6ED5A7D835BA1A075F" box="[1005,1053,258,284]" pageId="54" pageNumber="305">type</typeStatus>
|
||
locality in Western New
|
||
<collectingCountry id="51E2760BFFDF6D6ED30ED836BB9F0758" box="[1348,1432,257,283]" name="Guinea-Bissau" pageId="54" pageNumber="305">Guinea</collectingCountry>
|
||
,
|
||
<collectingCountry id="51E2760BFFDF6D6ED6DDD812BF03077C" box="[151,260,293,319]" name="Indonesia" pageId="54" pageNumber="305">Indonesia</collectingCountry>
|
||
(
|
||
<figureCitation id="B1CE2A1EFFDF6D6ED759D812BF710703" box="[275,374,293,320]" captionStart="FIGURE 158" captionStartId="53.[151,250,1956,1980]" captionTargetBox="[151,1436,1165,1932]" captionTargetId="figure-421@53.[151,1436,1165,1932]" captionTargetPageId="53" captionText="FIGURE 158. Distribution of Xanthonomus: X. toxopeanus (circle), X. latus (triangle), X. guineensis (squares), X. thayerae (diamonds)." figureDoi="http://doi.org/10.5281/zenodo.10821299" httpUri="https://zenodo.org/record/10821299/files/figure.png" pageId="54" pageNumber="305">Fig. 158</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61EF6510FFDF6D6ED68DD87EBF37078D" pageId="54" pageNumber="305" type="etymology">
|
||
<paragraph id="294A369BFFDF6D6ED68DD87EBED207C4" blockId="54.[151,1437,150,463]" pageId="54" pageNumber="305">
|
||
<emphasis id="1B81EA89FFDF6D6ED68DD87EBF490720" bold="true" box="[199,334,329,355]" pageId="54" pageNumber="305">Etymology.</emphasis>
|
||
The specific epithet is the latinized adjective derived from the area the specimens were collected from.
|
||
</paragraph>
|
||
<paragraph id="294A369BFFDF6D6ED68DD8A7BF37078D" blockId="54.[151,1437,150,463]" pageId="54" pageNumber="305">
|
||
<emphasis id="1B81EA89FFDF6D6ED68DD8A7BF4C07E9" bold="true" box="[199,331,400,426]" pageId="54" pageNumber="305">Bionomics.</emphasis>
|
||
Specimens were collected at elevations from
|
||
<quantity id="EE0D9B7EFFDF6D6ED512D8A7BA0A07E8" box="[856,1037,400,427]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="54" pageNumber="305" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100 to 2200 m</quantity>
|
||
a.s.l. The detailed bionomical data are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |