143 lines
20 KiB
XML
143 lines
20 KiB
XML
<document id="7BC0308EDF0375AB7D0D3907391A264B" ID-DOI="10.5281/zenodo.170156" ID-GBIF-Dataset="f457a885-11a0-44a9-a283-1e3f3c622874" ID-ISSN="11755326" ID-Zenodo-Dep="170156" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459751259515" checkinUser="plazi" docAuthor="Miller, Kelly B." docDate="2005" docId="9D36878AE831FF8BFEC29E18DDF8FE0B" docLanguage="en" docName="zt01059p059.pdf" docOrigin="Zootaxa 1059" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Agaporomorphus silvaticus Miller, new species" docType="treatment" docVersion="9" lastPageNumber="57" masterDocId="610FFFF2E836FF83FFCA9F04DF14FFDD" masterDocTitle="Two new species of Agaporomorphus Zimmermann (Coleoptera: Dytiscidae) from Peru" masterLastPageNumber="59" masterPageNumber="49" pageNumber="56" updateTime="1698211831608" updateUser="plazi">
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<mods:mods id="E2E2C56B3FCF18E492AA4787307F7C4A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D4BA0C208E3F27E97216C99ED26097F6">
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<mods:title id="5A5657876B0BAD3A2AC5F05192AD7671">Two new species of Agaporomorphus Zimmermann (Coleoptera: Dytiscidae) from Peru</mods:title>
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</mods:titleInfo>
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<mods:name id="FBF3CE9ED76760DB163C050C27370B8F" type="personal">
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<mods:namePart id="514C19FA317DB7F91277AC57F97B8FAD">Miller, Kelly B.</mods:namePart>
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<mods:typeOfResource id="7C449FA617AD42C5F94947639E6A606F">text</mods:typeOfResource>
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<mods:title id="8B65BCB5713E860739727429EB3FD170">Zootaxa</mods:title>
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<mods:part id="42827EC2A4E40BCF7BEC133CBE599773">
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<mods:date id="65703F0F6DE31ED868C377C6C3A3B4F0">2005</mods:date>
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<mods:number id="92FB15845E3CE90F8EF09E6C352611E3">1059</mods:number>
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<mods:start id="7B8665B2EAC711AE035348389763B82B">49</mods:start>
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<mods:classification id="FE670B699E6C68757523C0E8E57BED4D">journal article</mods:classification>
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<mods:identifier id="C3925D37BBBB049C5DB7254F639AB006" type="DOI">10.5281/zenodo.170156</mods:identifier>
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<mods:identifier id="41109C5011AA0D07B500CAB35C45A737" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="7FDE32704E2F5E79B4A9F77CECAF5358" type="Zenodo-Dep">170156</mods:identifier>
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<treatment id="9D36878AE831FF8BFEC29E18DDF8FE0B" ID-DOI="http://doi.org/10.5281/zenodo.6265560" ID-GBIF-Taxon="119337775" ID-Zenodo-Dep="6265560" LSID="urn:lsid:plazi:treatment:9D36878AE831FF8BFEC29E18DDF8FE0B" httpUri="http://treatment.plazi.org/id/9D36878AE831FF8BFEC29E18DDF8FE0B" lastPageId="8" lastPageNumber="57" pageId="7" pageNumber="56">
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<subSubSection id="5D856517E831FF84FEC29E18DECBFE84" pageId="7" pageNumber="56" type="nomenclature">
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<paragraph id="1520369CE831FF84FEC29E18DC51FEE8" blockId="7.[264,837,283,345]" box="[264,837,283,310]" pageId="7" pageNumber="56">
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<heading id="4E6881F0E831FF84FEC29E18DC51FEE8" bold="true" box="[264,837,283,310]" fontSize="11" level="1" pageId="7" pageNumber="56" reason="1">
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<emphasis id="27EBEA8EE831FF84FEC29E18DC51FEE8" bold="true" box="[264,837,283,310]" pageId="7" pageNumber="56">
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<taxonomicName id="D29F4D1FE831FF84FEC29E18DDBCFEE8" ID-CoL="65MG6" authority="Miller" authorityName="Miller" box="[264,680,283,310]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="silvaticus" status="sp. nov.">
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<emphasis id="27EBEA8EE831FF84FEC29E18DD47FEEB" bold="true" box="[264,595,284,310]" italics="true" pageId="7" pageNumber="56">Agaporomorphus silvaticus</emphasis>
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Miller
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</taxonomicName>
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,
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<taxonomicNameLabel id="3CD857F5E831FF84FD7F9E1FDC51FEE8" box="[693,837,283,309]" pageId="7" pageNumber="56" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="1520369CE831FF84FEC29E3BDECBFE84" blockId="7.[264,837,283,345]" box="[264,479,319,345]" pageId="7" pageNumber="56">(Figs. 3, 6, 16–18)</paragraph>
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</subSubSection>
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<subSubSection id="5D856517E831FF84FEC29E8ADD0BFD44" pageId="7" pageNumber="56" type="diagnosis">
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<paragraph id="1520369CE831FF84FEC29E8ADD0BFD44" blockId="7.[264,1324,398,1905]" pageId="7" pageNumber="56">
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<emphasis id="27EBEA8EE831FF84FEC29E8ADE9CFE75" bold="true" box="[264,392,398,424]" pageId="7" pageNumber="56">Diagnosis.</emphasis>
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This species lacks conspicuous modifications to the antennae (Fig. 3). The protarsal claws are long and robust with the ventral margins slightly sinuate and the bases strongly curved (Fig. 6). The mesotarsal claws are not long or sinuate. Males lack a triangular process along the posterior margin of abdominal sternite VI and lack a series of oblique rugae on each side of abdominal ventrite III. The male median lobe is very robust and strongly curved in lateral aspect (
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<figureCitation id="8DA42A19E831FF84FD1B9D53DC3FFDAC" box="[721,811,599,625]" captionStart="FIGURES 8 – 18" captionStartId="6.[264,382,1711,1735]" captionTargetBox="[316,1265,289,1710]" captionTargetId="figure@6.[316,1276,274,1710]" captionTargetPageId="6" captionText="FIGURES 8 – 18. Agaporomorphus spp., male genitalia. 8 – 11 — A. knischi; 8 — right lateral lobe, right lateral aspect; 9 — median lobe, right lateral aspect; 10 — median lobe, ventral aspect; 11 — right lateral lobe, left lateral aspect. 12 – 15 — A. tambopatensis; 12 — right lateral lobe, right lateral aspect; 13 — median lobe, right lateral aspect; 14 — median lobe, ventral aspect; 15 — right lateral lobe, left lateral aspect. 16 – 18 — A. silvaticus; 16 — right lateral lobe, right lateral aspect; 17 — median lobe, right lateral aspect; 18 — median lobe, ventral aspect." httpUri="https://zenodo.org/record/170157/files/figure.png" pageId="7" pageNumber="56">Fig. 17</figureCitation>
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). It is convoluted with folds and lobes in ventral aspect (
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<figureCitation id="8DA42A19E831FF84FE709D7BDD05FD44" box="[442,529,639,665]" captionStart="FIGURES 8 – 18" captionStartId="6.[264,382,1711,1735]" captionTargetBox="[316,1265,289,1710]" captionTargetId="figure@6.[316,1276,274,1710]" captionTargetPageId="6" captionText="FIGURES 8 – 18. Agaporomorphus spp., male genitalia. 8 – 11 — A. knischi; 8 — right lateral lobe, right lateral aspect; 9 — median lobe, right lateral aspect; 10 — median lobe, ventral aspect; 11 — right lateral lobe, left lateral aspect. 12 – 15 — A. tambopatensis; 12 — right lateral lobe, right lateral aspect; 13 — median lobe, right lateral aspect; 14 — median lobe, ventral aspect; 15 — right lateral lobe, left lateral aspect. 16 – 18 — A. silvaticus; 16 — right lateral lobe, right lateral aspect; 17 — median lobe, right lateral aspect; 18 — median lobe, ventral aspect." httpUri="https://zenodo.org/record/170157/files/figure.png" pageId="7" pageNumber="56">Fig. 18</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="5D856517E831FF84FEF29DA2DBF9F984" pageId="7" pageNumber="56" type="description">
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<paragraph id="1520369CE831FF84FEF29DA2DBF9F984" blockId="7.[264,1324,398,1905]" pageId="7" pageNumber="56">
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<emphasis id="27EBEA8EE831FF84FEF29DA2DEC4FD1D" bold="true" box="[312,464,678,704]" pageId="7" pageNumber="56">Description.</emphasis>
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<emphasis id="27EBEA8EE831FF84FE219DA3DD81FD1D" box="[491,661,679,704]" italics="true" pageId="7" pageNumber="56">Measurements</emphasis>
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. TL = 3.48 mm, GW = 1.70 mm, TL/GW = 2.04.
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<emphasis id="27EBEA8EE831FF84FEC29DCBDE9DFD35" box="[264,393,719,744]" italics="true" pageId="7" pageNumber="56">Coloration</emphasis>
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. Redbrown on all dorsal surfaces, broadly light yellow along anterior margin of elytron. All ventral surfaces and appendages yellow except abdominal sternites, medial portion of prosternum and prosternal process redyellow.
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<emphasis id="27EBEA8EE831FF84FC309C1BDA36FCE5" box="[1018,1314,799,824]" italics="true" pageId="7" pageNumber="56">Sculpture and structure</emphasis>
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. Pronotum with microsculpture consisting of fine, slightly longitudinally lengthened cells, with very fine and short longitudinal striae dispersed randomly and moderately densely; lateral pronotal bead obscured in anterior onefourth. Prosternum medially strongly and sharply carinate, carina extending onto prosternal process; prosternal process medially with a distinct longitudinal carina extending to apex, laterally with strongly beaded margins, apex pointed. Elytron covered with extremely fine, evenly spaced, short striae, striae more punctiform laterally and apically. Metafemur moderately broad, length about 3.2 times greatest width.
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<emphasis id="27EBEA8EE831FF84FD839B5BDDEFFBA5" box="[585,763,1119,1144]" italics="true" pageId="7" pageNumber="56">Male genitalia</emphasis>
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. Median lobe in lateral aspect very robust, broadly expanded dorsad, strongly curved, apex narrowed with small hyaline apicoventral lobe (
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<figureCitation id="8DA42A19E831FF84FE9B9BABDEAEFB14" box="[337,442,1199,1225]" captionStart="FIGURES 8 – 18" captionStartId="6.[264,382,1711,1735]" captionTargetBox="[316,1265,289,1710]" captionTargetId="figure@6.[316,1276,274,1710]" captionTargetPageId="6" captionText="FIGURES 8 – 18. Agaporomorphus spp., male genitalia. 8 – 11 — A. knischi; 8 — right lateral lobe, right lateral aspect; 9 — median lobe, right lateral aspect; 10 — median lobe, ventral aspect; 11 — right lateral lobe, left lateral aspect. 12 – 15 — A. tambopatensis; 12 — right lateral lobe, right lateral aspect; 13 — median lobe, right lateral aspect; 14 — median lobe, ventral aspect; 15 — right lateral lobe, left lateral aspect. 16 – 18 — A. silvaticus; 16 — right lateral lobe, right lateral aspect; 17 — median lobe, right lateral aspect; 18 — median lobe, ventral aspect." httpUri="https://zenodo.org/record/170157/files/figure.png" pageId="7" pageNumber="56">Figs. 17</figureCitation>
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); in ventral aspect very robust, broad, with complicated folding and structures (
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<figureCitation id="8DA42A19E831FF84FE409BD3DEF5FB2C" box="[394,481,1239,1265]" captionStart="FIGURES 8 – 18" captionStartId="6.[264,382,1711,1735]" captionTargetBox="[316,1265,289,1710]" captionTargetId="figure@6.[316,1276,274,1710]" captionTargetPageId="6" captionText="FIGURES 8 – 18. Agaporomorphus spp., male genitalia. 8 – 11 — A. knischi; 8 — right lateral lobe, right lateral aspect; 9 — median lobe, right lateral aspect; 10 — median lobe, ventral aspect; 11 — right lateral lobe, left lateral aspect. 12 – 15 — A. tambopatensis; 12 — right lateral lobe, right lateral aspect; 13 — median lobe, right lateral aspect; 14 — median lobe, ventral aspect; 15 — right lateral lobe, left lateral aspect. 16 – 18 — A. silvaticus; 16 — right lateral lobe, right lateral aspect; 17 — median lobe, right lateral aspect; 18 — median lobe, ventral aspect." httpUri="https://zenodo.org/record/170157/files/figure.png" pageId="7" pageNumber="56">Fig. 18</figureCitation>
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). Lateral lobe broad basally, narrowed medially, apical portion slender, apicomedially with large membranous lobes bearing series of few long setae (
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<figureCitation id="8DA42A19E831FF84FB5F9BFBDBFDFAC4" box="[1173,1257,1279,1305]" captionStart="FIGURES 8 – 18" captionStartId="6.[264,382,1711,1735]" captionTargetBox="[316,1265,289,1710]" captionTargetId="figure@6.[316,1276,274,1710]" captionTargetPageId="6" captionText="FIGURES 8 – 18. Agaporomorphus spp., male genitalia. 8 – 11 — A. knischi; 8 — right lateral lobe, right lateral aspect; 9 — median lobe, right lateral aspect; 10 — median lobe, ventral aspect; 11 — right lateral lobe, left lateral aspect. 12 – 15 — A. tambopatensis; 12 — right lateral lobe, right lateral aspect; 13 — median lobe, right lateral aspect; 14 — median lobe, ventral aspect; 15 — right lateral lobe, left lateral aspect. 16 – 18 — A. silvaticus; 16 — right lateral lobe, right lateral aspect; 17 — median lobe, right lateral aspect; 18 — median lobe, ventral aspect." httpUri="https://zenodo.org/record/170157/files/figure.png" pageId="7" pageNumber="56">Fig. 16</figureCitation>
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) also with setae apicomedially.
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<emphasis id="27EBEA8EE831FF84FDF69A23DD80FA9D" box="[572,660,1319,1344]" italics="true" pageId="7" pageNumber="56">Female</emphasis>
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. Unknown.
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<emphasis id="27EBEA8EE831FF84FCE59A23DB05FA9D" box="[815,1041,1319,1344]" italics="true" pageId="7" pageNumber="56">Sexual dimorphism</emphasis>
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. Female unknown, but male protarsal claws modified, long, basally strongly curved, slightly sinuate (Fig. 6); pro and mesotarsal claws a little less than half length of mesotarsomere V; without apical lobe on mesotarsomere V; protarsomeres I and II broadened, protarsomere I with two large adhesive setae, protarsomere II without adhesive setae; mesotarsomeres I and II slightly broadened, mesotarsomere I with one large, medial adhesive seta and two large, apical adhesive setae, mesotarsomere II with two smaller, apical adhesive setae. Male antennomeres V and VI slightly broader and flatter than other antennomeres (Fig. 3).
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</paragraph>
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</subSubSection>
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<subSubSection id="5D856517E831FF84FEF29962DD3AF8AC" pageId="7" pageNumber="56" type="etymology">
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<paragraph id="1520369CE831FF84FEF29962DE9DF90C" blockId="7.[264,1324,398,1905]" pageId="7" pageNumber="56">
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<emphasis id="27EBEA8EE831FF84FEF29962DED0F95D" bold="true" box="[312,452,1638,1664]" pageId="7" pageNumber="56">Etymology.</emphasis>
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The specific epithet is a Latin adjective
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<taxonomicName id="D29F4D1FE831FF84FC6C9963DB01F95D" box="[934,1045,1639,1664]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="silvaticus">
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<emphasis id="27EBEA8EE831FF84FC6C9963DB01F95D" box="[934,1045,1639,1664]" italics="true" pageId="7" pageNumber="56">silvaticus</emphasis>
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</taxonomicName>
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,
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<emphasis id="27EBEA8EE831FF84FBE79963DB2FF95D" box="[1069,1083,1639,1664]" italics="true" pageId="7" pageNumber="56">a</emphasis>
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,
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<emphasis id="27EBEA8EE831FF84FB9E9963DB6CF95D" box="[1108,1144,1639,1664]" italics="true" pageId="7" pageNumber="56">um</emphasis>
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, meaning of or belonging to a wood or to trees, referring to the dense, virgin forest in which this species was found.
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</paragraph>
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<paragraph id="1520369CE831FF84FEF299DADD3AF8AC" blockId="7.[264,1324,398,1905]" pageId="7" pageNumber="56">
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<emphasis id="27EBEA8EE831FF84FEF299DADD9AF925" bold="true" box="[312,654,1758,1784]" pageId="7" pageNumber="56">Phylogenetic relationships.</emphasis>
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<taxonomicName id="D29F4D1FE831FF84FD6799DBDCECF925" box="[685,1016,1759,1784]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="silvaticus">
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<emphasis id="27EBEA8EE831FF84FD6799DBDCECF925" box="[685,1016,1759,1784]" italics="true" pageId="7" pageNumber="56">Agaporomorphus silvaticus</emphasis>
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</taxonomicName>
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is the sister to a clade containing
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<taxonomicName id="D29F4D1FE831FF84FE589803DD77F8FD" box="[402,611,1799,1824]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="tambopatensis">
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<emphasis id="27EBEA8EE831FF84FE589803DD77F8FD" box="[402,611,1799,1824]" italics="true" pageId="7" pageNumber="56">A. tambopatensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="D29F4D1FE831FF84FD639803DC30F8FD" box="[681,804,1799,1824]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="56" phylum="Arthropoda" rank="species" species="knischi">
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<emphasis id="27EBEA8EE831FF84FD639803DC30F8FD" box="[681,804,1799,1824]" italics="true" pageId="7" pageNumber="56">A. knischi</emphasis>
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</taxonomicName>
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based on the common presence in these species of a small series of setae on each side of the dorsal surface of the male median lobe (Char. 7,
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<figureCitation id="8DA42A19E831FF84FEBF9853DD0BF8AC" box="[373,543,1879,1905]" captionStart="FIGURES 8 – 18" captionStartId="6.[264,382,1711,1735]" captionTargetBox="[316,1265,289,1710]" captionTargetId="figure@6.[316,1276,274,1710]" captionTargetPageId="6" captionText="FIGURES 8 – 18. Agaporomorphus spp., male genitalia. 8 – 11 — A. knischi; 8 — right lateral lobe, right lateral aspect; 9 — median lobe, right lateral aspect; 10 — median lobe, ventral aspect; 11 — right lateral lobe, left lateral aspect. 12 – 15 — A. tambopatensis; 12 — right lateral lobe, right lateral aspect; 13 — median lobe, right lateral aspect; 14 — median lobe, ventral aspect; 15 — right lateral lobe, left lateral aspect. 16 – 18 — A. silvaticus; 16 — right lateral lobe, right lateral aspect; 17 — median lobe, right lateral aspect; 18 — median lobe, ventral aspect." httpUri="https://zenodo.org/record/170157/files/figure.png" pageId="7" pageNumber="56">Figs. 9, 13, 17</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="5D856517E83EFF8BFEF29E1FDCAFFEEB" box="[312,955,283,310]" pageId="8" pageNumber="57" type="distribution">
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<paragraph id="1520369CE83EFF8BFEF29E1FDCAFFEEB" blockId="8.[264,1323,283,470]" box="[312,955,283,310]" pageId="8" pageNumber="57">
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<emphasis id="27EBEA8EE83EFF8BFEF29E1FDECCFEE8" bold="true" box="[312,472,283,309]" pageId="8" pageNumber="57">Distribution.</emphasis>
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Known only from Madre de Dios,
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<collectingCountry id="6D88760CE83EFF8BFCB49E18DCA3FEEB" box="[894,951,284,310]" name="Peru" pageId="8" pageNumber="57">Peru</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="5D856517E83EFF8BFEF29E47DDF8FE0B" pageId="8" pageNumber="57" type="materials_examined">
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<paragraph id="1520369CE83EFF8BFEF29E47DDF8FE0B" blockId="8.[264,1323,283,470]" pageId="8" pageNumber="57">
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<emphasis id="27EBEA8EE83EFF8BFEF29E47DD2CFE80" bold="true" box="[312,568,323,349]" pageId="8" pageNumber="57">Material examined.</emphasis>
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<typeStatus id="CA24883EE83EFF8BFD959E40DDEAFE83" box="[607,766,324,350]" pageId="8" pageNumber="57" type="holotype">HOLOTYPE</typeStatus>
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: ɗ labeled, “
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<collectingCountry id="6D88760CE83EFF8BFC7C9E40DB1CFE83" box="[950,1032,324,350]" name="Peru" pageId="8" pageNumber="57">PERU</collectingCountry>
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: Madre de Dios, Rio Tambopata, Posadas Amazonas,
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<date id="6121105CE83EFF8BFD7C9E68DCA2FE5B" box="[694,950,364,390]" pageId="8" pageNumber="57" value="2003-12-13">13 December 2003</date>
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, K.B. Miller/
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<typeStatus id="CA24883EE83EFF8BFB4D9E68DA36FE5B" box="[1159,1314,364,390]" pageId="8" pageNumber="57" type="holotype">HOLOTYPE</typeStatus>
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:
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<taxonomicName id="D29F4D1FE83EFF8BFEC29E90DDF3FE73" authority="Miller, 2005" authorityName="Miller" authorityYear="2005" box="[264,743,404,430]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="silvaticus">
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<emphasis id="27EBEA8EE83EFF8BFEC29E90DD58FE70" box="[264,588,404,429]" italics="true" pageId="8" pageNumber="57">Agaporomorphus silvaticus</emphasis>
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Miller, 2005
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</taxonomicName>
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[red label with double black line border].” This species is known only from the
|
||
<typeStatus id="CA24883EE83EFF8BFDB79EB8DDFCFE0B" box="[637,744,444,470]" pageId="8" pageNumber="57" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |