158 lines
22 KiB
XML
158 lines
22 KiB
XML
<document id="0015A7CC397DCFAB8061B2FBCFFE4F37" ID-DOI="http://dx.doi.org/10.3897/zookeys.689.12021" ID-GBIF-Dataset="907713bd-c7f2-4f8e-a75e-b7f9c958865f" ID-PMC="PMC5672588" ID-Pensoft-Pub="1313-2970-689-1" ID-PubMed="29200920" ID-ZooBank="2F40BF4AD35F4CC697D5976EC201E652" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-689-1" ModsDocOrigin="ZooKeys 689" ModsDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" checkinTime="1502744367222" checkinUser="pensoft" docAuthor="Lackner, Tomas & Leschen, Richard A. B." docDate="2017" docId="9A8B4FBCD1AD50F5533CDA9E2738D905" docLanguage="en" docName="ZooKeys 689: 1-263" docOrigin="ZooKeys 689" docSource="http://dx.doi.org/10.3897/zookeys.689.12021" docTitle="Saprinus (Saprinus) artensis Marseul 1862" docType="treatment" docVersion="6" lastPageNumber="83" masterDocId="D302FFB6FFA66345FFBAFF89311EFFDE" masterDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" masterLastPageNumber="263" masterPageNumber="1" pageNumber="79" updateTime="1701316739810" updateUser="plazi">
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<mods:title id="07AF9D4681A4B7D1BE0210C80B9E5994">A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)</mods:title>
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<mods:namePart id="E3C9DEE1F71CB7429B1FCF56A8EE0B11">Lackner, Tomas</mods:namePart>
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<mods:namePart id="A48E895885916D783A0E70F9744C1C25">Leschen, Richard A. B.</mods:namePart>
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<mods:date id="62742AA9A3EAD97D5FD0D272217F1CB3">2017</mods:date>
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<mods:classification id="647AAA1A6315ACE1173E8825A8ADDCC1">journal article</mods:classification>
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<mods:identifier id="CC4B508B2F91FBBE57514017AC241577" type="DOI">http://dx.doi.org/10.3897/zookeys.689.12021</mods:identifier>
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<treatment id="9A8B4FBCD1AD50F5533CDA9E2738D905" ID-GBIF-Taxon="132648676" LSID="urn:lsid:plazi:treatment:9A8B4FBCD1AD50F5533CDA9E2738D905" httpUri="http://treatment.plazi.org/id/9A8B4FBCD1AD50F5533CDA9E2738D905" lastPageId="82" lastPageNumber="83" pageId="78" pageNumber="79">
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<subSubSection id="E0A6EC835C487FF0DA2E7E92299603D8" pageId="78" pageNumber="79" type="nomenclature">
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<paragraph id="058F623C752787DBF5FAF2CA41D87505" pageId="78" pageNumber="79">
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<taxonomicName id="2AD84CD6B603E3DE1BCB81BC9B5C7973" ID-CoL="7WS3J" authority="Marseul, 1862" authorityName="Marseul" authorityYear="1862" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) artensis" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="artensis" subGenus="Saprinus">Saprinus (Saprinus) artensis Marseul, 1862</taxonomicName>
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Figs 320, 321-329, 330-336, 760
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</paragraph>
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</subSubSection>
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<subSubSection id="1EF9541C7BF883EF3AB15EA9A8C14415" pageId="78" pageNumber="79" type="reference_group">
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<paragraph id="A3047447736F3F4FFCDA7E80353DF5CF" pageId="78" pageNumber="79">
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<taxonomicName id="DE573B74B18BB1AF4064566DF5608F02" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus artensis" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="artensis">Saprinus artensis</taxonomicName>
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Marseul, 1862: 445.
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</paragraph>
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</subSubSection>
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<subSubSection id="6847E05AB4846666D4E86F81BE196F7D" pageId="78" pageNumber="79" type="type locality">
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<paragraph id="027610B58A525AD5273652E6F1805416" pageId="78" pageNumber="79">Type locality.</paragraph>
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<paragraph id="56667FF42D98DF70DB8D4F97163FCE51" pageId="78" pageNumber="79">New Caledonia.</paragraph>
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<caption id="2000A05E652C92EE3091045F54FDA989" pageId="78" pageNumber="79">
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<paragraph id="137033A96AA7166B40B4029040220803" pageId="78" pageNumber="79">
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Figure 320.
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<taxonomicName id="BA2D4E51B4EE84A1714E6EC0FF98ABAF" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) artensis" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="artensis" subGenus="Saprinus">Saprinus (Saprinus) artensis</taxonomicName>
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Marseul, 1862 habitus, dorsal view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="2BE3D18E9964A4EC4D719520E40EBC5E" pageId="78" pageNumber="79" type="type material">
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<paragraph id="D865EB116F0529F8FBA4D3D65D387B2C" pageId="78" pageNumber="79">Type material examined.</paragraph>
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<paragraph id="1185721BAB0397FD1369460C106E92E0" pageId="78" pageNumber="79">
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<taxonomicName id="51A32F6FA4456080262ED8C5BA72CAB3" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus artensis" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="artensis">Saprinus artensis</taxonomicName>
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Marseul, 1862: Lectotype, present designation: ♂, side mounted on a triangular card, with terminalia glued on the same triangular card as specimen, right protarsus and left mesotarsus missing, both metatarsal claws broken off, with the following labels: "13 /
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<taxonomicName id="DEFDE0A76847008CD46F0D10BFA94163" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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/
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<taxonomicName id="9AF3F7286173E614F918158FEBD8F5D9" lsidName="artensis" pageId="78" pageNumber="79" rank="species" species="artensis">artensis</taxonomicName>
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m. / illegible text" (round pink label, written); followed by: "
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<taxonomicName id="D192F7B5D396662ACC686342507ADF1F" lsidName="artensis" pageId="78" pageNumber="79" rank="species" species="artensis">artensis</taxonomicName>
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/ type" (pencil-written label added probably by G. Dahlgren); followed by:
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<normalizedToken id="34532846CC46CD5F51D53DBB5CBCB4E9" originalValue="“TYPE”">"TYPE"</normalizedToken>
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(red-typed label); followed by: "MUSEUM PARIS /
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<taxonomicName id="6DE55FF29756D7174517782FEAE0751D" lsidName="artensis" pageId="78" pageNumber="79" rank="species" species="artensis">artensis</taxonomicName>
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/ COLL / DE MARSEUL 1890" (pink label, typed-written); followed by:
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<normalizedToken id="5F99E89800517FAF11DD02EEBB268D8B" originalValue="“09-052”">"09-052"</normalizedToken>
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(yellow, pencil-written label, added by the senior author); followed by: "
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<taxonomicName id="EDED3B814B52F5F0DC77F235F780DD20" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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/
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<taxonomicName id="BE88A7B74C26B362C49612F7282FBEEB" lsidName="artensis" pageId="78" pageNumber="79" rank="species" species="artensis">artensis</taxonomicName>
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/ Marseul, 1862 / LECTOTYPE / Des. T. Lackner
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<normalizedToken id="029A00ADE073E727A93308FFE9434256" originalValue="’11”">'11"</normalizedToken>
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(red label, written) (MNHN). This species was described from an unknown number of specimens and the lectotype designation fixes the identity of species.
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</paragraph>
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</subSubSection>
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<subSubSection id="30541850D566D189D76328D3236A0ACC" pageId="78" pageNumber="79" type="materials_examined">
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<paragraph id="65FB080DDEF60C9949F093089F81A31E" pageId="78" pageNumber="79">Additional material examined.</paragraph>
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<paragraph id="858EDD1DCBA7D2BFBECB58D2CC9330AB" pageId="78" pageNumber="79">
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1 ♂, New Caledonia (MNHN). Apparently this specimen is also from
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<normalizedToken id="DCB25E308DC9198AAEDEE6A4CD6C99D9" originalValue="Marseul’s">Marseul's</normalizedToken>
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collection, but does not originate from the same sampling as the lectotype. There is no
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<normalizedToken id="1C0E4070AC6E7CD35AFC9ED99B1C8AC6" originalValue="“Type”">"Type"</normalizedToken>
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label on it and therefore it is not designated as the paralectotype; 5 ♂♂, "New Caledonia", all from
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<normalizedToken id="EF67104D75EAC7DF87CDB8DDEF3C523E" originalValue="Schmidt’s">Schmidt's</normalizedToken>
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collection, one male has a small label:
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<normalizedToken id="C043DDB6C8213EEA9046AC97FD1BD556" originalValue="“Type”">"Type"</normalizedToken>
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- it is not clear whether it forms a part of the syntype series or not (ZMHUB); 1 ♀, New Caledonia, A. Deyr[olle] (NCB). 1 ♂ & 1 ♀, N. Caledonia, Fauvel (BMNH).
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</paragraph>
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</subSubSection>
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<subSubSection id="52180C166DF32869C0417B0896071A93" pageId="78" pageNumber="79" type="biology_ecology">
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<paragraph id="98F568B7AF7FB970D5E04999D5354820" pageId="78" pageNumber="79">Biology.</paragraph>
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<paragraph id="746E47A30DF86987FEA40430ACF6FC35" pageId="78" pageNumber="79">Unknown, presumably similar to congeners.</paragraph>
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</subSubSection>
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<subSubSection id="9DACD833398518EA42F82F5F1747A980" pageId="78" pageNumber="79" type="distribution">
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<paragraph id="6F18BF3985EE36521FAFC74976237871" pageId="78" pageNumber="79">Distribution.</paragraph>
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<paragraph id="781D3BD41F55F731CACFB8D9386D77F3" pageId="78" pageNumber="79">Endemic to New Caledonia, rarely collected (Fig. 760).</paragraph>
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</subSubSection>
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<subSubSection id="C3485F4233A44F28934E4A73E410B1ED" lastPageId="82" lastPageNumber="83" pageId="78" pageNumber="79" type="description">
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<paragraph id="2FD820B88191FFCEDCA95D3116379C83" pageId="78" pageNumber="79">Re-description.</paragraph>
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<paragraph id="7FA75640FE5059B37766AF9E4AAFA11D" pageId="78" pageNumber="79">Body length: PEL: 3.35 mm; EL: 2.00 mm; APW: 1.25 mm; PPW: 2.50 mm; EW: 2.80 mm (only the type specimen was measured).</paragraph>
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<paragraph id="FA7ADB9382E387A657768AF77B69B231" pageId="78" pageNumber="79">Body (Fig. 320), convex, cuticle dark brown, almost black, shining, without metallic luster; legs, mouthparts and antennal scape dark brown; antennal club black.</paragraph>
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<paragraph id="C7BB7000B2127D44E97A83F5E92528D3" pageId="78" pageNumber="79">Antennal scape (Fig. 321) black, slightly thickened, finely punctuate, with two setae; antennal club (Fig. 322) covered with dense short sensilla intermingled with sparse longer erect setae; sensory structures of antennal club not examined.</paragraph>
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<caption id="5F8B14568801E1C787F238129E3A6306" pageId="78" pageNumber="79">
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<paragraph id="B446F1CA4474901DD14930567C89DBD3" pageId="78" pageNumber="79">
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Figures 321-329. 321
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<taxonomicName id="F56012ECA0AC853DDFC952AFE793B623" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) artensis" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="artensis" subGenus="Saprinus">Saprinus (Saprinus) artensis</taxonomicName>
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Marseul, 1862 head, dorsal view 322 antennal club, ventral view 323 mentum, ventral view 324 propygidium + pygidium 325 prosternum 326 mesoventrite 327 lateral disc of metaventrite + metepisternum 328 protibia, ventral view 329 mesotibia, ventral view.
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</paragraph>
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</caption>
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<paragraph id="F303B42337F7D230DEA42F4E399F9CA5" pageId="79" pageNumber="80">
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<pageBreakToken id="CEF0E68F52C096F80D7512FFBE749430" pageId="79" pageNumber="80" start="start">Mandibles</pageBreakToken>
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dorso-laterally densely punctuate, rounded; labrum finely and sparsely punctuate, convex, with shallow median depression; labral pits present, each with a single labral seta; mentum (Fig. 323) almost quadrate, furnished with sporadic seate, anterior margin with deep triangular notch furnished with several longer setae; other mouthparts not examined.
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</paragraph>
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<paragraph id="D375469F1C6BE018AFDAED9F1102FC70" pageId="79" pageNumber="80">Clypeus (Fig. 321) broad, flattened, sloping down laterally, its punctation denser and coarser than that of labrum; frontal and supra-orbital striae complete, slightly carinate; frontal disc (Fig. 321) finely punctuate, punctures separated by about several times their diameter; eyes convex, well visible from above.</paragraph>
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<paragraph id="93228DD2893C5AD2D29EF981681829AC" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
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Pronotal sides (Fig. 320) moderately narrowing anteriorly, apical angles obtuse, pronotal depressions shallow but present, anterior incision for head shallow; marginal pronotal stria complete, slightly carinate, visible along its entire length from dorsal view, weakened anteriorly; pronotal disc laterally with a band of deep dense elongate punctures originating approximately in pronotal depressions, but not reaching basal angles of pronotum, between it and pronotal margin a narrow smooth band present;
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<pageBreakToken id="7EB95F08D11502CC7A101AF739D2CA6F" pageId="80" pageNumber="81" start="start">rest</pageBreakToken>
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of the pronotal disc almost smooth, with only scattered microscopic punctation; double row of fine ovoid punctures present along pronotal base not reaching ante-scutellar area; pronotal hypomeron glabrous; scutellum small, visible.
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</paragraph>
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<paragraph id="D993769DEB1F55C4380009469FF3804B" pageId="80" pageNumber="81">Elytral epipleura with a double row of fine punctures; marginal epipleural stria complete; marginal elytral stria well impressed and slightly carinate, continued as complete apical elytral stria. Humeral elytral stria well impressed on basal fourth, faintly connected to rather long inner subhumeral stria; four dorsal elytral striae 1-4 well impressed, all about the same length (third elytral stria slightly shorter than others), not reaching elytral half apically; first and second dorsal elytral striae in fine punctures, first elytral interval with elongate strioles; third and fourth striae in large sparsely set punctures, surface of second and third elytral intervals almost smooth, only with microscopic punctation; fourth dorsal elytral stria curved towards sutural elytral stria but not connected with it; sutural elytral stria well-impressed, in fine punctures, abbreviated on basal fourth, apically connected with apical elytral stria; elytral disc on apical 4/7 in dense large punctures; punctures separated by about their own diameter, laterally and especially apically punctures aciculate; elytral flanks almost impunctate.</paragraph>
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<paragraph id="D169467C7F132F986D97E021C2606554" pageId="80" pageNumber="81">Propygidium (Fig. 324) very densely punctuate, punctures separated by less than their diameter; pygidium (Fig. 324) with similar, if somewhat sparser punctation, interspaces in both cases imbricate.</paragraph>
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<paragraph id="5A80073AFD24E783FA1742A618C0376D" pageId="80" pageNumber="81">Anterior margin of median portion of prosternum (Fig. 325) almost straight, rounded laterally; marginal prosternal stria present laterally and also as apical fragment; prosternal process between carinal prosternal striae flat, punctuate; carinal prosternal striae carinate, parallel on basal three-fourths, on apical fourth slightly divergent, united in front (Fig. 325); lateral prosternal striae carinate, rather short, apically attaining carinal prosternal striae at about three-fourths of their length.</paragraph>
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<paragraph id="B13A8739CD9987AE577D78E0943F2251" pageId="80" pageNumber="81">Anterior margin of mesoventrite (Fig. 326) distinctly inwardly arcuate; discal marginal mesoventral stria well impressed, carinate; disc with sparse fine punctation, punctures becoming larger near meso-metaventral suture; meso-metaventral sutural stria indicated by a row of large punctures; intercoxal disc of metaventrite flattened, with slight longitudinal median depression; disc of metaventrite for the most part almost smooth, along posterior margin several rows of punctation appears; lateral metaventral stria (Fig. 327) well impressed, carinate, almost straight, shortened; lateral disc of metaventrite (Fig. 327) slightly concave, with dense shallow large setigerous punctures; metepisternum (Fig. 327) similar, but with deeper and larger punctures without setae, on fused metepimeron punctures becoming much sparser; metepisternal stria present along fused metepimeron, along metepisternum present as short intermittent fragments.</paragraph>
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<paragraph id="A20F40CAC8FD37F5660A30C902786160" pageId="80" pageNumber="81">Intercoxal disc of first abdominal ventrite almost completely striate laterally; disc along basal and lateral margins with shallow punctures of various sizes; rest of sternite with scattered microscopic punctation.</paragraph>
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<paragraph id="F8DE805269FA34FE67ABCBDA3A823233" lastPageId="82" lastPageNumber="83" pageId="80" pageNumber="81">
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Protibia (Fig. 328) slightly dilated, outer margin with five low teeth topped by large denticle, denticles diminishing in size proximally, followed by three minute denticles; setae of outer, median rows, anterior protibial stria and protarsal groove not examined; protibial spur large, bent, growing out from apical margin of protibia; outer
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<pageBreakToken id="B8B3A37B69D8EC86EC4C6215E51CD0F3" pageId="81" pageNumber="82" start="start">part</pageBreakToken>
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of posterior surface (Fig. 328) slightly obscurely variolate, separated from finely punctuate and narrow median part of posterior surface by a definite stria bearing a row of setae; posterior protibial stria complete, bearing almost along its entire length dense row of setae becoming stiffer and longer apically; apical margin of protibia ventrally
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<pageBreakToken id="FCE3BA5EB8A01802B008E394F1057770" pageId="82" pageNumber="83" start="start">with</pageBreakToken>
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three short denticles; inner row of setae single, setae leaf-like, short, diminishing in size basally.
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</paragraph>
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<paragraph id="A49113BA01E0EE76AC4201A304719806" pageId="82" pageNumber="83">Mesotibia (Fig. 329) slender, outer margin with a row of sparse long denticles growing in size apically, another row of much shorter sparser denticles situated on anterior surface of mesotibia; setae of outer row regular, thick, almost as long as denticles themselves; setae of median row shorter and finer; posterior mesotibial stria almost complete; anterior surface of mesotibia sparsely punctuate, interspaces imbricate; anterior mesotibial stria complete; mesotibial spur stout, short; apical margin of mesotibia anteriorly with two short denticles; inner margin of mesotibia with sparse row of short setae; claws of apical tarsomere bent, longer than half its length; metatibia slenderer and longer than mesotibia, in all aspects similar to it, but denticles on outer margin much shorter and sparser.</paragraph>
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<paragraph id="29987D2A2F17C696DC9DC8587CA7CD3A" pageId="82" pageNumber="83">Male genitalia. Eighth sternite (Figs 330-331) with pseudo-pores, fused medially, apex with large velum mesally adorned with dense rows of microscopic setae; laterally apex of eighth sternite with rows of longer dense setae; eighth tergite and eighth sternite not fused laterally (Fig. 332). Ninth tergite (Figs 333-334) typical for the subfamily; tenth tergite inwardly arcuate, apical angles strongly sclerotized, bent; spiculum gastrale (Fig. 333) gradually dilated on most of its apical half; basal end heart-shaped, inwardly arcuate. Aedeagus (Figs 335-336) parallel-sided, with parameres fused along their basal half (roughly), basal piece of aedeagus short, ratio of its length: length of parameres 1: 4; aedeagus slightly curved from lateral view (Fig. 336).</paragraph>
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<caption id="43DB406298C1C22E78E111B0CCC33721" pageId="82" pageNumber="83">
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<paragraph id="39F0266CD70B5BC6A4B63B324C933F8E" pageId="82" pageNumber="83">
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Figures 330-336. 330
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<taxonomicName id="003909F06326EF91D653C711C8A9FDFE" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus (Saprinus) artensis" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="artensis" subGenus="Saprinus">Saprinus (Saprinus) artensis</taxonomicName>
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Marseul, 1862 male terminalia: 8th sternite + 8th tergite, ventral view 331 ditto, dorsal view 332 ditto, lateral view 333 male terminalia: 9th + 10th tergites, dorsal view; spiculum gastrale, ventral view 334 9th + 10th tergites; spiculum gastrale, lateral view 335 male terminalia: aedeagus, dorsal view 336 ditto, lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |