262 lines
60 KiB
XML
262 lines
60 KiB
XML
<document id="6C4EE8C86773E179FF419F003A508FDA" ID-CLB-Dataset="36889" ID-DOI="10.11646/zootaxa.3774.1.1" ID-GBIF-Dataset="41d311ee-850e-4125-9a40-5a96a2aa9065" ID-ISSN="1175-5326" ID-Zenodo-Dep="285693" ID-ZooBank="B5A2EF46-2BF6-4ED3-A5F4-5F9951400545" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460391910470" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2014" docId="9A6687E64F10302EFF34FD14FDDCFE66" docLanguage="en" docName="zt03774p030.pdf" docOrigin="Zootaxa 3774 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microscydmus Saulcy & Croissandeau" docType="treatment" docVersion="8" lastPageNumber="5" masterDocId="665FFF9E4F13302AFFA3FFEEFFFEFFCA" masterDocTitle="The smallest Cyrtoscydmini of Australia: revision of Microscydmus Saulcy & Croissandeau and Penicillidmus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="4" updateTime="1698298320569" updateUser="ExternalLinkService">
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<mods:title id="2D2D79C61D724B6DC2DDD38988CC65E4">The smallest Cyrtoscydmini of Australia: revision of Microscydmus Saulcy & Croissandeau and Penicillidmus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="3644FEC2874B799603C400E990432321">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="39F3EC7CFED1210C22833C5E97DCCED2">Zootaxa</mods:title>
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<mods:date id="876B579D429305840DAEBA5FD614B6DC">2014</mods:date>
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<mods:number id="19376F5F29B166A6BFAF36A5222A9717">3774</mods:number>
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<treatment id="9A6687E64F10302EFF34FD14FDDCFE66" ID-DOI="http://doi.org/10.5281/zenodo.5664958" ID-GBIF-Taxon="127667219" ID-Zenodo-Dep="5664958" LSID="urn:lsid:plazi:treatment:9A6687E64F10302EFF34FD14FDDCFE66" httpUri="http://treatment.plazi.org/id/9A6687E64F10302EFF34FD14FDDCFE66" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<subSubSection id="5AD5657B4F103029FF34FD14FD8DFCDE" box="[151,627,762,788]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="127036F04F103029FF34FD14FD8DFCDE" blockId="3.[151,627,762,788]" box="[151,627,762,788]" pageId="3" pageNumber="4">
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<heading id="4938819C4F103029FF34FD14FD8DFCDE" bold="true" box="[151,627,762,788]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<taxonomicName id="D5CF4D734F103029FF34FD14FD8DFCDE" ID-CoL="8VRV9" authority="Saulcy & Croissandeau" authorityName="Saulcy & Croissandeau" box="[151,627,762,788]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="20BBEAE24F103029FF34FD14FEB9FCDE" bold="true" box="[151,327,762,788]" italics="true" pageId="3" pageNumber="4">Microscydmus</emphasis>
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Saulcy & Croissandeau
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="5AD5657B4F10302EFF34FCAFFDDCFE66" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="reference_group">
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<paragraph id="127036F04F103029FF34FCAFFD23FCBC" blockId="3.[151,1436,833,886]" pageId="3" pageNumber="4">
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<treatmentCitationGroup id="32DF11DE4F103029FF34FCAFFD23FCBC" pageId="3" pageNumber="4">
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<taxonomicName id="D5CF4D734F103029FF34FCAFFD47FC9D" ID-CoL="8VRV9" authority="Saulcy & Croissandeau, 1893: 225" authorityName="Saulcy & Croissandeau" authorityPageNumber="225" authorityYear="1893" box="[151,697,833,855]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="20BBEAE24F103029FF34FCAFFED4FC9D" box="[151,298,833,855]" italics="true" pageId="3" pageNumber="4">Microscydmus</emphasis>
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<treatmentCitation id="936E10E14F103029FE9BFCAFFD47FC9D" author="Saulcy" box="[312,697,833,855]" page="225" pageId="3" pageNumber="4" year="1893">
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<bibRefCitation id="765E4B014F103029FE9BFCAFFD7FFC9D" author="Saulcy" box="[312,641,833,855]" pageId="3" pageNumber="4" refString="Saulcy, F. & Croissandeau, J. (1893) Tableau des genres des Scydmaenidae. Annales de la Societe Entomologique de France, 62, 225 - 238." type="journal article" year="1893">Saulcy & Croissandeau, 1893</bibRefCitation>
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: 225
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</treatmentCitation>
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</taxonomicName>
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(without species);
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<typeStatus id="CD7488524F103029FC30FCACFC40FC9D" box="[915,958,834,855]" pageId="3" pageNumber="4">type</typeStatus>
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species:
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<taxonomicName id="D5CF4D734F103029FB8FFCAFFA62FC9D" authority="Schaum, 1844" authorityName="Schaum" authorityYear="1844" box="[1068,1436,833,855]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nanus">
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<emphasis id="20BBEAE24F103029FB8FFCAFFB0AFC9D" box="[1068,1268,833,855]" italics="true" pageId="3" pageNumber="4">Scydmaenus nanus</emphasis>
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<bibRefCitation id="765E4B014F103029FAA0FCAFFA62FC9D" author="Schaum" box="[1283,1436,833,855]" pageId="3" pageNumber="4" refString="Schaum, H. R. (1844) Nachtrage zur Monographie der Gattung Scydmaenus. Zeitschrift fur die Entomologie, 5, 459 - 472." type="journal article" year="1844">Schaum, 1844</bibRefCitation>
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</taxonomicName>
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(subsequent monotypy by
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<treatmentCitation id="936E10E14F103029FE71FC8EFD2EFCBC" author="Croissandeau" box="[466,720,864,886]" page="105" pageId="3" pageNumber="4" year="1898">
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<bibRefCitation id="765E4B014F103029FE71FC8EFD60FCBC" author="Croissandeau" box="[466,670,864,886]" pageId="3" pageNumber="4" refString="Croissandeau, J. (1898) Monographie des Scydmaenidae. Annales de la Societe Entomologique de France, 67, 105 - 167." type="journal article" year="1898">Croissandeau, 1898</bibRefCitation>
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: 105
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</treatmentCitation>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="127036F04F103029FF34FC4CFA6DFA31" blockId="3.[151,1436,930,2035]" pageId="3" pageNumber="4">
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<emphasis id="20BBEAE24F103029FF34FC4CFD1AFC71" bold="true" box="[151,740,930,955]" pageId="3" pageNumber="4">
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Morphology and diagnosis of
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<taxonomicName id="D5CF4D734F103029FE54FC4CFD61FC71" ID-CoL="8VRV9" box="[503,671,930,955]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="20BBEAE24F103029FE54FC4CFD61FC71" bold="true" box="[503,671,930,955]" italics="true" pageId="3" pageNumber="4">Microscydmus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3B8857994F103029FD05FC4CFD1AFC71" box="[678,740,930,955]" pageId="3" pageNumber="4" sensu="stricto">s. str.</taxonomicNameLabel>
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</emphasis>
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The nominotypical subgenus of
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<taxonomicName id="D5CF4D734F103029FBF2FC4DFB0DFC71" ID-CoL="8VRV9" box="[1105,1267,931,955]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="20BBEAE24F103029FBF2FC4DFB0DFC71" box="[1105,1267,931,955]" italics="true" pageId="3" pageNumber="4">Microscydmus</emphasis>
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</taxonomicName>
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can be defined as a cyrtoscydmine member comprising beetles with typical "ant-like" appearance (i.e., with an elongate body and distinct constrictions between the head and prothorax and between prothorax and pterothorax + abdomen) and extremely small bodies (below
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<quantity id="D5379B154F103029FDA9FBE1FDAEFBEC" box="[522,592,1039,1063]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="mm" value="1.0">1 mm</quantity>
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in length, typically
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<quantity id="D5379B154F103029FCE7FBE0FC31FBED" box="[836,975,1038,1063]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="4" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6-0.8 mm</quantity>
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), sharing the following combination of characters: i) tempora, genae and sides of pronotum densely covered with straight, long and thick bristles (
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<figureCitation id="8AF42A754F103029FAE4FBDCFA69FB81" box="[1351,1431,1074,1099]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1640,1663]" captionTargetBox="[151,1436,497,1619]" captionTargetId="figure@4.[151,1436,479,1619]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Dorsal habitus (1); head in antero-dorsal view (2); left mandible in dorsal view (3); head in ventral view (4). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lb, labrum; md, mandible; mn, mentum; mst, mesostipes; mxp, maxillary palpus; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285694/files/figure.png" pageId="3" pageNumber="4">Figs. 4</figureCitation>
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,
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<figureCitation id="8AF42A754F103029FF34FBB9FF58FBA5" box="[151,166,1111,1135]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">8</figureCitation>
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); ii) head short, subtriangular or subtrapezoidal (
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<figureCitation id="8AF42A754F103029FD74FBB8FC9EFBA5" box="[727,864,1110,1135]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1640,1663]" captionTargetBox="[151,1436,497,1619]" captionTargetId="figure@4.[151,1436,479,1619]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Dorsal habitus (1); head in antero-dorsal view (2); left mandible in dorsal view (3); head in ventral view (4). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lb, labrum; md, mandible; mn, mentum; mst, mesostipes; mxp, maxillary palpus; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285694/files/figure.png" pageId="3" pageNumber="4">Figs. 1–2, 4</figureCitation>
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) with weakly convex vertex (
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<figureCitation id="8AF42A754F103029FB15FBB8FB00FBA5" box="[1206,1278,1110,1135]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1640,1663]" captionTargetBox="[151,1436,497,1619]" captionTargetId="figure@4.[151,1436,479,1619]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Dorsal habitus (1); head in antero-dorsal view (2); left mandible in dorsal view (3); head in ventral view (4). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lb, labrum; md, mandible; mn, mentum; mst, mesostipes; mxp, maxillary palpus; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285694/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
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;
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<emphasis id="20BBEAE24F103029FAA8FBB9FAE1FBA4" box="[1291,1311,1111,1134]" italics="true" pageId="3" pageNumber="4">vt</emphasis>
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) and frons slightly projecting anteriorly with its narrow anterior part narrowly separating antennal insertions; iii) antennae with distinct club composed of antennomeres IX–XI; iv) submentum short and broad, without lateral sutures (
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<figureCitation id="8AF42A754F103029FACCFB70FF56FB11" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1640,1663]" captionTargetBox="[151,1436,497,1619]" captionTargetId="figure@4.[151,1436,479,1619]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Dorsal habitus (1); head in antero-dorsal view (2); left mandible in dorsal view (3); head in ventral view (4). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lb, labrum; md, mandible; mn, mentum; mst, mesostipes; mxp, maxillary palpus; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285694/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
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;
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<emphasis id="20BBEAE24F103029FF17FB2DFF1FFB10" box="[180,225,1219,1242]" italics="true" pageId="3" pageNumber="4">smn</emphasis>
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); v) posterior tentorial pits visible in ventral view and located in distinct transverse groove (
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<figureCitation id="8AF42A754F103029FB4CFB2CFACBFB11" box="[1263,1333,1218,1243]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1640,1663]" captionTargetBox="[151,1436,497,1619]" captionTargetId="figure@4.[151,1436,479,1619]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Dorsal habitus (1); head in antero-dorsal view (2); left mandible in dorsal view (3); head in ventral view (4). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lb, labrum; md, mandible; mn, mentum; mst, mesostipes; mxp, maxillary palpus; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285694/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
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;
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<emphasis id="20BBEAE24F103029FAE2FB2DFA9BFB10" box="[1345,1381,1219,1242]" italics="true" pageId="3" pageNumber="4">ptp</emphasis>
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); vi) occipital constriction only slightly narrower than vertex (
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<figureCitation id="8AF42A754F103029FC99FB08FC7AFB35" box="[826,900,1254,1279]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1640,1663]" captionTargetBox="[151,1436,497,1619]" captionTargetId="figure@4.[151,1436,479,1619]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Dorsal habitus (1); head in antero-dorsal view (2); left mandible in dorsal view (3); head in ventral view (4). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lb, labrum; md, mandible; mn, mentum; mst, mesostipes; mxp, maxillary palpus; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285694/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
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;
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<emphasis id="20BBEAE24F103029FC36FB09FC3BFB34" box="[917,965,1255,1278]" italics="true" pageId="3" pageNumber="4">occ)</emphasis>
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; vii) prothorax without lateral marginal carinae, with complete hypomeral ridges (
|
||
<figureCitation id="8AF42A754F103029FDCFFAE4FD4EFAE9" box="[620,688,1290,1315]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FD1FFAE5FD1FFAE9" box="[700,737,1291,1315]" italics="true" pageId="3" pageNumber="4">hyr</emphasis>
|
||
) and basisternal part of prosternum (
|
||
<figureCitation id="8AF42A754F103029FBDFFAE4FB3EFAE9" box="[1148,1216,1290,1315]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FB6FFAE5FB13FAE9" box="[1228,1261,1291,1315]" italics="true" pageId="3" pageNumber="4">bst</emphasis>
|
||
) laterally fused with internal (adcoxal) part of each hypomeron; viii) pronotum with two pairs of ante-basal pits (
|
||
<figureCitation id="8AF42A754F103029FAACFAC0FAA9FA8D" box="[1295,1367,1326,1351]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FACBFAC0FA72FA8C" box="[1384,1420,1326,1350]" italics="true" pageId="3" pageNumber="4">abf</emphasis>
|
||
), internal pair connected by variously distinct transverse impression or groove (
|
||
<figureCitation id="8AF42A754F103029FBB2FABCFBA9FAA1" box="[1041,1111,1362,1387]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FBC6FABDFB71FAA1" box="[1125,1167,1363,1387]" italics="true" pageId="3" pageNumber="4">abg</emphasis>
|
||
); ix) mesoventrite with asetose impressions and setose impressions (
|
||
<figureCitation id="8AF42A754F103029FD33FA98FCE4FA45" box="[656,794,1398,1423]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Figs. 10, 13</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FC84FA99FCC3FA44" box="[807,829,1399,1422]" italics="true" pageId="3" pageNumber="4">ai</emphasis>
|
||
,
|
||
<emphasis id="20BBEAE24F103029FCEFFA99FCA1FA44" box="[844,863,1399,1422]" italics="true" pageId="3" pageNumber="4">si</emphasis>
|
||
) and mesoventral intercoxal process (
|
||
<figureCitation id="8AF42A754F103029FAAEFA98FA69FA45" box="[1293,1431,1398,1423]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Figs. 10, 13</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FF34FA75FF2FFA78" box="[151,209,1435,1458]" italics="true" pageId="3" pageNumber="4">msvp</emphasis>
|
||
) carinate, moderately expanding ventrally and not connected anteriorly to anterior ridge of mesoventrite (
|
||
<figureCitation id="8AF42A754F103029FF3CFA50FED2FA1D" box="[159,300,1470,1495]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Figs. 10, 13</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FE98FA51FEAAFA1C" box="[315,340,1471,1494]" italics="true" pageId="3" pageNumber="4">ar</emphasis>
|
||
); x) metaventral intercoxal process (
|
||
<figureCitation id="8AF42A754F103029FD5FFA50FCACFA1D" box="[764,850,1470,1495]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Fig. 10</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FCC2FA51FC69FA1C" box="[865,919,1471,1494]" italics="true" pageId="3" pageNumber="4">mtvp</emphasis>
|
||
) with two long spines; xi) each elytron with short but distinct and deep basal impression bearing one large basal elytral fovea filled with sparse setae (
|
||
<figureCitation id="8AF42A754F103029FA97FA0CFA7BFA31" box="[1332,1413,1506,1531]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="127036F04F103029FF64F9E8FA62F839" blockId="3.[151,1436,930,2035]" pageId="3" pageNumber="4">
|
||
Additionally, species of
|
||
<taxonomicName id="D5CF4D734F103029FE42F9E8FD7DF9D4" ID-CoL="8VRV9" box="[481,643,1542,1566]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="20BBEAE24F103029FE42F9E8FD7DF9D4" box="[481,643,1542,1566]" italics="true" pageId="3" pageNumber="4">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B8857994F103029FD2EF9E9FD35F9D4" box="[653,715,1543,1566]" pageId="3" pageNumber="4" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
are characterized by distinct tempora (
|
||
<figureCitation id="8AF42A754F103029FB30F9E8FB25F9D5" box="[1171,1243,1542,1567]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1640,1663]" captionTargetBox="[151,1436,497,1619]" captionTargetId="figure@4.[151,1436,479,1619]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Dorsal habitus (1); head in antero-dorsal view (2); left mandible in dorsal view (3); head in ventral view (4). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lb, labrum; md, mandible; mn, mentum; mst, mesostipes; mxp, maxillary palpus; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285694/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FB49F9E9FAF8F9D4" box="[1258,1286,1543,1566]" italics="true" pageId="3" pageNumber="4">tm</emphasis>
|
||
); absence of fronto-clypeal groove; antennal insertions located on anterior surface of head; mandibles (
|
||
<figureCitation id="8AF42A754F103029FB44F9C4FAA3F989" box="[1255,1373,1578,1603]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 7" captionStartId-0="4.[151,264,1640,1663]" captionStartId-1="5.[151,264,1876,1899]" captionTargetBox-0="[151,1436,497,1619]" captionTargetBox-1="[170,1413,796,1836]" captionTargetId-0="figure@4.[151,1436,479,1619]" captionTargetId-1="figure@5.[170,1414,796,1836]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 1 – 4. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Dorsal habitus (1); head in antero-dorsal view (2); left mandible in dorsal view (3); head in ventral view (4). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lb, labrum; md, mandible; mn, mentum; mst, mesostipes; mxp, maxillary palpus; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." captionText-1="FIGURES 5 – 7. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Mouthparts in ventral view (5); details of anterior part of labium in ventral view (6); left maxillary palpus in ventral view (7). Abbreviations: gal, galea; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere I – III; md, mandible; mn, mentum; mst, mesostipes; mxp 1 – 4, maxillary palpomere I – IV." httpUri-0="https://zenodo.org/record/285694/files/figure.png" httpUri-1="https://zenodo.org/record/285695/files/figure.png" pageId="3" pageNumber="4">Figs. 2–5</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FAD3F9C5FA6CF989" box="[1392,1426,1579,1603]" italics="true" pageId="3" pageNumber="4">md</emphasis>
|
||
) subtriangular, with indistinct, small and often visible only in ventral view sub-apical tooth (
|
||
<figureCitation id="8AF42A754F103029FB36F9A0FB29F9AD" box="[1173,1239,1614,1639]" captionStart="FIGURES 5 – 7" captionStartId="5.[151,264,1876,1899]" captionTargetBox="[170,1413,796,1836]" captionTargetId="figure@5.[170,1414,796,1836]" captionTargetPageId="5" captionText="FIGURES 5 – 7. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Mouthparts in ventral view (5); details of anterior part of labium in ventral view (6); left maxillary palpus in ventral view (7). Abbreviations: gal, galea; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere I – III; md, mandible; mn, mentum; mst, mesostipes; mxp 1 – 4, maxillary palpomere I – IV." httpUri="https://zenodo.org/record/285695/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
|
||
) and large dorsomesal prostheca (
|
||
<figureCitation id="8AF42A754F103029FEC3F99CFE56F941" box="[352,424,1650,1675]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1640,1663]" captionTargetBox="[151,1436,497,1619]" captionTargetId="figure@4.[151,1436,479,1619]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Dorsal habitus (1); head in antero-dorsal view (2); left mandible in dorsal view (3); head in ventral view (4). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lb, labrum; md, mandible; mn, mentum; mst, mesostipes; mxp, maxillary palpus; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285694/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FE1BF99DFE27F940" box="[440,473,1651,1674]" italics="true" pageId="3" pageNumber="4">pst</emphasis>
|
||
); hypostomal ridges (
|
||
<figureCitation id="8AF42A754F103029FD7AF99CFCDFF941" box="[729,801,1650,1675]" captionStart="FIGURES 1 – 4" captionStartId="4.[151,264,1640,1663]" captionTargetBox="[151,1436,497,1619]" captionTargetId="figure@4.[151,1436,479,1619]" captionTargetPageId="4" captionText="FIGURES 1 – 4. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Dorsal habitus (1); head in antero-dorsal view (2); left mandible in dorsal view (3); head in ventral view (4). Abbreviations: bst, basistipes; cd, cardo; cl, clypeus; fr, frons; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lb, labrum; md, mandible; mn, mentum; mst, mesostipes; mxp, maxillary palpus; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; smn, submentum; tm, temple; vt, vertex." httpUri="https://zenodo.org/record/285694/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FC93F99DFCB7F941" box="[816,841,1651,1675]" italics="true" pageId="3" pageNumber="4">hr</emphasis>
|
||
) short and extending postero-mesally, not reaching anterior margin of gular plate; maxillary palpomere III (
|
||
<figureCitation id="8AF42A754F103029FCB4F978FCA3F964" box="[791,861,1686,1711]" captionStart="FIGURES 5 – 7" captionStartId="5.[151,264,1876,1899]" captionTargetBox="[170,1413,796,1836]" captionTargetId="figure@5.[170,1414,796,1836]" captionTargetPageId="5" captionText="FIGURES 5 – 7. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Mouthparts in ventral view (5); details of anterior part of labium in ventral view (6); left maxillary palpus in ventral view (7). Abbreviations: gal, galea; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere I – III; md, mandible; mn, mentum; mst, mesostipes; mxp 1 – 4, maxillary palpomere I – IV." httpUri="https://zenodo.org/record/285695/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FCCFF979FC56F965" box="[876,936,1687,1711]" italics="true" pageId="3" pageNumber="4">mxp3</emphasis>
|
||
) only 3–
|
||
<date id="667110304F103029FBB3F979FBE0F965" box="[1040,1054,1687,1711]" pageId="3" pageNumber="4">4</date>
|
||
x as long as broad, palpomere IV (
|
||
<figureCitation id="8AF42A754F103029FF3CF954FF1AF918" box="[159,228,1722,1747]" captionStart="FIGURES 5 – 7" captionStartId="5.[151,264,1876,1899]" captionTargetBox="[170,1413,796,1836]" captionTargetId="figure@5.[170,1414,796,1836]" captionTargetPageId="5" captionText="FIGURES 5 – 7. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Mouthparts in ventral view (5); details of anterior part of labium in ventral view (6); left maxillary palpus in ventral view (7). Abbreviations: gal, galea; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere I – III; md, mandible; mn, mentum; mst, mesostipes; mxp 1 – 4, maxillary palpomere I – IV." httpUri="https://zenodo.org/record/285695/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FF53F955FED3F919" box="[240,301,1723,1747]" italics="true" pageId="3" pageNumber="4">mxp4</emphasis>
|
||
) strongly elongate and slender, with broadened short and setose basal part and distinctly demarcated, abruptly narrowing and nearly rod-like pointed distal part; prementum with short subtriangular or subtrapezoidal ligula (
|
||
<figureCitation id="8AF42A754F103029FF44F8ECFEAAF8D1" box="[231,340,1794,1819]" captionStart="FIGURES 5 – 7" captionStartId="5.[151,264,1876,1899]" captionTargetBox="[170,1413,796,1836]" captionTargetId="figure@5.[170,1414,796,1836]" captionTargetPageId="5" captionText="FIGURES 5 – 7. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Mouthparts in ventral view (5); details of anterior part of labium in ventral view (6); left maxillary palpus in ventral view (7). Abbreviations: gal, galea; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere I – III; md, mandible; mn, mentum; mst, mesostipes; mxp 1 – 4, maxillary palpomere I – IV." httpUri="https://zenodo.org/record/285695/files/figure.png" pageId="3" pageNumber="4">Figs. 5–6</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FEC3F8EDFE88F8D1" box="[352,374,1795,1819]" italics="true" pageId="3" pageNumber="4">lg</emphasis>
|
||
) bearing two or three thick setae (in two specimens of
|
||
<taxonomicName id="D5CF4D734F103029FC45F8EDFBB1F8D0" box="[998,1103,1795,1818]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nanus">
|
||
<emphasis id="20BBEAE24F103029FC45F8EDFBB1F8D0" box="[998,1103,1795,1818]" italics="true" pageId="3" pageNumber="4">M. nanus</emphasis>
|
||
</taxonomicName>
|
||
studied by SEM one has two setae, the other one, illustrated here, three setae; two setae were also found in
|
||
<taxonomicName id="D5CF4D734F103029FBE9F8C9FB2DF8F4" ID-CoL="435SS" box="[1098,1235,1831,1854]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="minimus">
|
||
<emphasis id="20BBEAE24F103029FBE9F8C9FB2DF8F4" box="[1098,1235,1831,1854]" italics="true" pageId="3" pageNumber="4">M. minimus</emphasis>
|
||
</taxonomicName>
|
||
); prothorax with indistinct, weakly carinate prosternal intercoxal process (
|
||
<figureCitation id="8AF42A754F103029FC86F8A4FC92F8A9" box="[805,876,1866,1891]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FCDFF8A5FC5DF8A8" box="[892,931,1867,1890]" italics="true" pageId="3" pageNumber="4">psp</emphasis>
|
||
); procoxal sockets (
|
||
<figureCitation id="8AF42A754F103029FB2EF8A4FB2CF8A9" box="[1165,1234,1866,1891]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
|
||
) closed by broad lateral projection of coxal part of sternum; pronotosternal sutures (
|
||
<figureCitation id="8AF42A754F103029FC03F880FC16F84D" box="[928,1000,1902,1927]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FC5BF881FBE2F84C" box="[1016,1052,1903,1926]" italics="true" pageId="3" pageNumber="4">nss</emphasis>
|
||
) visible only as short notches in antero-ventral part of prothorax; mesoscutum and mesoscutellum (
|
||
<figureCitation id="8AF42A754F103029FC77F87CFBCDF861" box="[980,1075,1938,1963]" captionStart="FIGURES 8 – 13" captionStartId="6.[151,264,1673,1696]" captionTargetBox="[151,1436,193,1652]" captionTargetId="figure@6.[151,1436,193,1652]" captionTargetPageId="6" captionText="FIGURES 8 – 13. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Prothorax in ventral view (8); base of pronotum in dorsal view (9); pterothorax and abdomen in ventral view (10); base of elytron in dorsal view (11); pterothorax n dorsal view (12); mesothorax in ventral view (13). Abbreviations: abf, ante-basal pronotal fovea; abg, ante-basal pronotal groove; aest 2, mesanepisternum; ai, asetose impression of mesoventrite; alc, alacrista; ar, anterior ridge of mesoventrite; bst, basisternal part of prosternum; hy, hypomeron; hyr, hypomeral ridge; mma, median membranous area; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; msp, mesocoxal projection; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pre, prepectus; psn 3, postmetanotum; psp, prosternal intercoxal process; s 3 – 8, abdominal sternites III – VIII; sc 2 + scl 2, mesocutum + mesoscutellum; sc 3, metascutum; si, setose impression of mesoventrite; v 3, metaventrite." httpUri="https://zenodo.org/record/285696/files/figure.png" pageId="3" pageNumber="4">Fig. 12</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F103029FBE8F87DFB4EF861" box="[1099,1200,1939,1963]" italics="true" pageId="3" pageNumber="4">sc2+scl2</emphasis>
|
||
) not separated by scutoscutellar suture, forming single subtriangular plate not visible between elytral bases in intact specimens; aedeagus (
|
||
<figureCitation id="8AF42A754F103029FEADF834FE71F839" box="[270,399,2010,2035]" captionStart="FIGURES 17 – 19" captionStartId="8.[151,264,1700,1723]" captionTargetBox="[151,1436,193,1678]" captionTargetId="figure@8.[151,1436,193,1680]" captionTargetPageId="8" captionText="FIGURES 17 – 19. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Aedeagus in dorso-lateral view (17); apex of median lobe in dorso-lateral view (18); apex of paramere (19). Abbreviations: aml, apex of median lobe; bo, basal orifice; dw, dorsal wall of median lobe; pm, paramere." httpUri="https://zenodo.org/record/285698/files/figure.png" pageId="3" pageNumber="4">Figs 17–19</figureCitation>
|
||
) drop-shaped, with thin walls of median lobe, basally or sub-basally located foramen (
|
||
<figureCitation id="8AF42A754F103029FACCF834FA62F839" box="[1391,1436,2010,2035]" captionStart="FIGURES 17 – 19" captionStartId="8.[151,264,1700,1723]" captionTargetBox="[151,1436,193,1678]" captionTargetId="figure@8.[151,1436,193,1680]" captionTargetPageId="8" captionText="FIGURES 17 – 19. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Aedeagus in dorso-lateral view (17); apex of median lobe in dorso-lateral view (18); apex of paramere (19). Abbreviations: aml, apex of median lobe; bo, basal orifice; dw, dorsal wall of median lobe; pm, paramere." httpUri="https://zenodo.org/record/285698/files/figure.png" pageId="3" pageNumber="4">Fig.</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="127036F04F17302EFF34FF76FCE0FF1F" blockId="4.[151,1436,151,429]" pageId="4" pageNumber="5">
|
||
17;
|
||
<emphasis id="20BBEAE24F17302EFF67FF76FF1EFF7A" box="[196,224,152,176]" italics="true" pageId="4" pageNumber="5">bo</emphasis>
|
||
) and free, slender parameres (
|
||
<figureCitation id="8AF42A754F17302EFD9FFF79FD6FFF7A" box="[572,657,151,176]" captionStart="FIGURES 17 – 19" captionStartId="8.[151,264,1700,1723]" captionTargetBox="[151,1436,193,1678]" captionTargetId="figure@8.[151,1436,193,1680]" captionTargetPageId="8" captionText="FIGURES 17 – 19. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Aedeagus in dorso-lateral view (17); apex of median lobe in dorso-lateral view (18); apex of paramere (19). Abbreviations: aml, apex of median lobe; bo, basal orifice; dw, dorsal wall of median lobe; pm, paramere." httpUri="https://zenodo.org/record/285698/files/figure.png" pageId="4" pageNumber="5">Fig. 17</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F17302EFD3CFF77FD3FFF7A" box="[671,705,153,176]" italics="true" pageId="4" pageNumber="5">pm</emphasis>
|
||
), typically with single robust apical seta (
|
||
<figureCitation id="8AF42A754F17302EFB07FF79FB07FF7A" box="[1188,1273,151,176]" captionStart="FIGURES 17 – 19" captionStartId="8.[151,264,1700,1723]" captionTargetBox="[151,1436,193,1678]" captionTargetId="figure@8.[151,1436,193,1680]" captionTargetPageId="8" captionText="FIGURES 17 – 19. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Aedeagus in dorso-lateral view (17); apex of median lobe in dorso-lateral view (18); apex of paramere (19). Abbreviations: aml, apex of median lobe; bo, basal orifice; dw, dorsal wall of median lobe; pm, paramere." httpUri="https://zenodo.org/record/285698/files/figure.png" pageId="4" pageNumber="5">Fig. 19</figureCitation>
|
||
); spermatheca sclerotized and globular, usually located deeply in thorax.
|
||
</paragraph>
|
||
<paragraph id="127036F04F17302EFF64FF0EFDDCFE66" blockId="4.[151,1436,151,429]" pageId="4" pageNumber="5">
|
||
In at least some species of
|
||
<taxonomicName id="D5CF4D734F17302EFDB3FF0EFD4CFF32" ID-CoL="8VRV9" box="[528,690,224,248]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="20BBEAE24F17302EFDB3FF0EFD4CFF32" box="[528,690,224,248]" italics="true" pageId="4" pageNumber="5">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3B8857994F17302EFD1DFF0EFD00FF3D" box="[702,766,224,247]" pageId="4" pageNumber="5" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
and all in the subgenus
|
||
<taxonomicName id="D5CF4D734F17302EFB8AFF0EFA90FF32" authority="Franz, 1970" authorityName="Franz" authorityYear="1970" box="[1065,1390,224,248]" class="Insecta" family="Staphylinidae" genus="Parastenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="20BBEAE24F17302EFB8AFF0EFB29FF32" box="[1065,1239,224,248]" italics="true" pageId="4" pageNumber="5">Parastenichnus</emphasis>
|
||
<bibRefCitation id="765E4B014F17302EFB40FF0EFA90FF32" author="Franz" box="[1251,1390,224,248]" pageId="4" pageNumber="5" refString="Franz, H. (1970) Zur Kenntnis der Scydmaeniden-Fauna von Singapore, Malakka und Indonesien (Coleoptera: Scydmaenidae). Beitrage zur Entomologie, 20, 535 - 578." type="journal article" year="1970">Franz, 1970</bibRefCitation>
|
||
</taxonomicName>
|
||
the protibiae of males have modified apices, bearing externally located combs (
|
||
<figureCitation id="8AF42A754F17302EFC51FEEAFB82FED7" box="[1010,1148,260,285]" captionStart="FIGURES 14 – 16" captionStartId="7.[151,264,1786,1809]" captionTargetBox="[151,1436,193,1764]" captionTargetId="figure@7.[151,1436,193,1765]" captionTargetPageId="7" captionText="FIGURES 14 – 16. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Apex of left protibia in ventral view (14); apex of right protibia in internal view (15); apex of left protibia and tarsomere I in ventral view (16). Abbreviations: cp, cuticular projection; ec, external comb; ic, internal comb; trs 1 – 2, tarsomere I – II; vs, ventral spine." httpUri="https://zenodo.org/record/285697/files/figure.png" pageId="4" pageNumber="5">Figs. 14–16</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F17302EFB2AFEEBFB5CFED6" box="[1161,1186,261,284]" italics="true" pageId="4" pageNumber="5">ec</emphasis>
|
||
) composed of several curved teeth fused together at base. Additionally the protibial apex can be armed with another, internally located comb (
|
||
<figureCitation id="8AF42A754F17302EFF47FEA2FE93FEAF" box="[228,365,332,357]" captionStart="FIGURES 14 – 16" captionStartId="7.[151,264,1786,1809]" captionTargetBox="[151,1436,193,1764]" captionTargetId="figure@7.[151,1436,193,1765]" captionTargetPageId="7" captionText="FIGURES 14 – 16. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Apex of left protibia in ventral view (14); apex of right protibia in internal view (15); apex of left protibia and tarsomere I in ventral view (16). Abbreviations: cp, cuticular projection; ec, external comb; ic, internal comb; trs 1 – 2, tarsomere I – II; vs, ventral spine." httpUri="https://zenodo.org/record/285697/files/figure.png" pageId="4" pageNumber="5">Figs. 14–15</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F17302EFEDBFEA3FE72FEAE" box="[376,396,333,356]" italics="true" pageId="4" pageNumber="5">ic</emphasis>
|
||
) of several straight cuticular projections, and with two robust ventral spines (
|
||
<figureCitation id="8AF42A754F17302EFB4DFEA2FA8EFEAF" box="[1262,1392,332,357]" captionStart="FIGURES 14 – 16" captionStartId="7.[151,264,1786,1809]" captionTargetBox="[151,1436,193,1764]" captionTargetId="figure@7.[151,1436,193,1765]" captionTargetPageId="7" captionText="FIGURES 14 – 16. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Apex of left protibia in ventral view (14); apex of right protibia in internal view (15); apex of left protibia and tarsomere I in ventral view (16). Abbreviations: cp, cuticular projection; ec, external comb; ic, internal comb; trs 1 – 2, tarsomere I – II; vs, ventral spine." httpUri="https://zenodo.org/record/285697/files/figure.png" pageId="4" pageNumber="5">Figs 14–16</figureCitation>
|
||
;
|
||
<emphasis id="20BBEAE24F17302EFAD8FEA3FA6CFEAE" box="[1403,1426,333,356]" italics="true" pageId="4" pageNumber="5">vs</emphasis>
|
||
) at base of the tarsomere I. Ventral setae on tarsomeres in the
|
||
<typeStatus id="CD7488524F17302EFCD8FE9EFC55FE42" box="[891,939,368,392]" pageId="4" pageNumber="5">type</typeStatus>
|
||
species of the genus are accompanied by cucticular projections (
|
||
<figureCitation id="8AF42A754F17302EFE3BFE7AFE15FE67" box="[408,491,404,429]" captionStart="FIGURES 14 – 16" captionStartId="7.[151,264,1786,1809]" captionTargetBox="[151,1436,193,1764]" captionTargetId="figure@7.[151,1436,193,1765]" captionTargetPageId="7" captionText="FIGURES 14 – 16. Morphology of the type species of Microscydmus Saulcy & Croissandeau, M. nanus (Schaum), male. Apex of left protibia in ventral view (14); apex of right protibia in internal view (15); apex of left protibia and tarsomere I in ventral view (16). Abbreviations: cp, cuticular projection; ec, external comb; ic, internal comb; trs 1 – 2, tarsomere I – II; vs, ventral spine." httpUri="https://zenodo.org/record/285697/files/figure.png" pageId="4" pageNumber="5">Fig. 16</figureCitation>
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;
|
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<emphasis id="20BBEAE24F17302EFE54FE7BFDEFFE66" box="[503,529,405,428]" italics="true" pageId="4" pageNumber="5">cp</emphasis>
|
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).
|
||
</paragraph>
|
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</subSubSection>
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</treatment>
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</document> |