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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.531.6097" ID-GBIF-Dataset="70702787-a6a8-4af7-9702-738f344a5cbe" ID-PMC="PMC4668911" ID-Pensoft-Pub="1313-2970-531-1" ID-PubMed="26692803" ID-ZBK="C845838EC9124BD8AB4E07980F91959E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-531-1" ModsDocOrigin="ZooKeys 531" ModsDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" checkinTime="1451243834596" checkinUser="pensoft" docAuthor="Vermeulen, Jaap J., Liew, Thor-Seng &amp; Schilthuizen, Menno" docDate="2015" docId="8F555DB010B1B5A7A57B978A2BE63A3B" docLanguage="en" docName="ZooKeys 531: 1-139" docOrigin="ZooKeys 531" docSource="http://dx.doi.org/10.3897/zookeys.531.6097" docTitle="Ditropopsis trachychilus Vermeulen, Liew &amp; Schilthuizen, sp. n." docType="treatment" docUuid="DBF5C5C5-B845-4479-B2FE-44D0940DCD7D" docUuidSource="ZooBank" docVersion="10" lastPageNumber="13" masterDocId="1527DF5A6645C540FFF1FF85AF4AE358" masterDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" masterLastPageNumber="139" masterPageNumber="1" pageNumber="13" updateTime="1668160884494" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liew, Thor-Seng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schilthuizen, Menno</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>531</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>139</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.531.6097</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.531.6097</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-531-1</mods:identifier>
<mods:identifier type="ZBK">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
<mods:identifier type="ZooBank">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="144057491" LSID="urn:lsid:zoobank.org:act:DBF5C5C5-B845-4479-B2FE-44D0940DCD7D" httpUri="http://treatment.plazi.org/id/8F555DB010B1B5A7A57B978A2BE63A3B" lastPageNumber="13" pageId="12" pageNumber="13">
<subSubSection pageId="12" pageNumber="13" type="multiple">
<paragraph pageId="12" pageNumber="13">Taxon classification Animalia Architaenioglossa Cyclophoridae</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="nomenclature">
<paragraph pageId="12" pageNumber="13">
<taxonomicName LSID="http://zoobank.org/DBF5C5C5-B845-4479-B2FE-44D0940DCD7D" authority="Vermeulen, Liew &amp; Schilthuizen" class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis trachychilus" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="trachychilus">Ditropopsis trachychilus Vermeulen, Liew &amp; Schilthuizen</taxonomicName>
<taxonomicNameLabel pageId="12" pageNumber="13">sp. n.</taxonomicNameLabel>
Figure 10
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="holotype">
<paragraph pageId="12" pageNumber="13">
<materialsCitation collectingDate="2015-01-01" collectingDateMax="2015-12-31" collectingDateMin="2015-01-01" collectionCode="RMNH" collectorName="J. J. Vermeulen, M. Schilthuizen" country="Malaysia" httpUri="http://data.biodiversitydata.nl/naturalis/specimen/RMNH.5003918" location="West Coast Province" municipality="Sabah" specimenCode="RMNH. 5003918" specimenCount="1" typeStatus="holotype">
<typeStatus>Holotype</typeStatus>
.
<collectingCountry>Malaysia</collectingCountry>
,
<collectingMunicipality>Sabah</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:8F555DB010B1B5A7A57B978A2BE63A3B:82BA2E1E5120E13A94BFDDD472061ADA" country="Malaysia" municipality="Sabah" name="West Coast Province">West Coast Province</location>
,
<location LSID="urn:lsid:plazi:treatment:8F555DB010B1B5A7A57B978A2BE63A3B:B3D54A1F82275B7376F5FE28A7506A41" country="Malaysia" municipality="Sabah" name="Crocker Range N. P.">Crocker Range N.P.</location>
,
<location LSID="urn:lsid:plazi:treatment:8F555DB010B1B5A7A57B978A2BE63A3B:3D5ADD480C6C8783E011ACC3A2D944F5" country="Malaysia" municipality="Sabah" name="near the km 54 marker on the road Kota Kinabalu-Tambunan">near the km 54 marker on the road Kota Kinabalu-Tambunan</location>
,
<location LSID="urn:lsid:plazi:treatment:8F555DB010B1B5A7A57B978A2BE63A3B:D2C4B9EAB5F6FB360E8014F24EC290D5" country="Malaysia" municipality="Sabah" name="Gunung Mas">Gunung Mas</location>
(leg.
<collectorName>J.J. Vermeulen</collectorName>
&amp;
<collectorName>M. Schilthuizen</collectorName>
,
<specimenCode>RMNH.5003918</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="materials_examined">
<paragraph pageId="12" pageNumber="13">Examined material from Sabah.</paragraph>
<paragraph pageId="12" pageNumber="13">
West Coast Province, Kinabalu N.P., summit trail, between 2308-2775 m (leg. T.S. Liew, BOR/MOL 4121, BOR/MOL 4113); Bowen route at 3376 m (leg. T.S. Liew &amp; Jasilin, BOR/MOL 4111); Mesilau trail between 2092-2356 m (leg. T.S. Liew, BOR/MOL 4119, BOR/MOL 4114);
<normalizedToken originalValue="Kotals" pageId="12" pageNumber="13">Kotal's</normalizedToken>
route between 2924-3048 m (leg. T.S. Liew, J. Lapidin, Safrie &amp; Jasilin, BOR/MOL 4117, BOR/MOL 4115; leg. T.S. Liew &amp; Jasilin, BOR/MOL 4118); Sayap-Nunuhon trail between 2640-2944 m (leg. T.S. Liew, Dominik, J. Lapidin &amp; Jasilin, BOR/MOL 4116, BOR/MOL 4112, BOR/MOL 4120, BOR/MOL 4123, BOR/MOL 4124).
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="description">
<paragraph pageId="12" pageNumber="13">Description.</paragraph>
<paragraph pageId="12" pageNumber="13">Shell very small, rather thick, hardly translucent, pale greenish. Surface shiny. Spire rather high-conical. Radial sculpture: fine growth lines, locally grading into fine, densely placed riblets. Sculpture. Spiral sculpture on the last whorl: 4 cords: 1 supraperipheral and 1 peripheral, about equally wide, 1 basal; these all rather distinct; 1 umbilical, inconspicuous and located deep inside the umbilicus; next to these a fine spiral striation on the lower surface. This sculpture continuing up to about 0.8 mm from the peristome, to be replaced by densely and irregularly placed radial riblets with a periostracal crest. Aperture. Peristome simple, not expanded, parietal side slightly detached from the penultimate whorl, basal side rounded, hardly drawn out. Dimensions. Height c. 2.3 mm; width c. 2 mm; h/w c. 1.15; umbilicus measured over the basal spiral cord c. 25 % of the shell width; number of whorls c. 4 1/8; height aperture c. 0.8 mm; width aperture c. 0.8 mm.</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="habitat in sabah and distribution">
<paragraph pageId="12" pageNumber="13">Habitat in Sabah and distribution.</paragraph>
<paragraph pageId="12" pageNumber="13">Primary montane oak forest on sandstone bedrock, in leaf litter, sub-alpine forest on ultrabasic and granodioritic bedrock, 1300-1400 m (Crocker Range), 2000-3400 m (Mount Kinabalu) alt. Sabah: Mount Kinabalu, Crocker Range. Endemic to Sabah.</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="cross diagnosis">
<paragraph pageId="12" pageNumber="13">Cross diagnosis.</paragraph>
<paragraph pageId="12" pageNumber="13">
Similar to
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis koperbergi" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="koperbergi">Ditropopsis koperbergi</taxonomicName>
, but shell higher conical. The distinctive sculpture near the aperture develops only when fully adult, as in other species of the genus.
</paragraph>
<paragraph pageId="12" pageNumber="13">
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis gradata" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="gradata">Ditropopsis gradata</taxonomicName>
Quadras &amp; Moellendorff, 1896 and
<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Ditropopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ditropopsis pusilla" order="Architaenioglossa" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="pusilla">Ditropopsis pusilla</taxonomicName>
Quadras &amp; Moellendorff, 1895, from the Philippines, have a similarly high spire and spiral cord distribution. Both species have a more rounded profile to the whorls.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="etymology">
<paragraph pageId="12" pageNumber="13">Etymology.</paragraph>
<paragraph pageId="12" pageNumber="13">The name refers to the peristome [trachus (Gr.) rough; cheilos (Gr.) = lip].</paragraph>
</subSubSection>
</treatment>
</document>