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<mods:affiliation id="2EE931FD2879ABE7E64931EC9B9B5351">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal &amp; Departamento de Biologia, Faculdade de Ciências, Universidade do Porto, 4099 - 002 Porto, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Portugal acrottini @ cibio. up. pt; https: // orcid. org / 0000 - 0002 - 8505 - 3050</mods:affiliation>
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<emphasis id="35D6531A1326BF37FF28F9DCFE2F4446" bold="true" box="[151,444,1626,1653]" italics="true" pageId="12" pageNumber="13">Lygodactylus expectatus</emphasis>
<bibRefCitation id="6333F2F91326BF37FE7BF9DDFD4F4447" author="Pasteur, G. &amp; Blanc, C. P." box="[452,732,1626,1653]" pageId="12" pageNumber="13" pagination="583 - 597" refId="ref39811" refString="Pasteur, G. &amp; Blanc, C. P. (1967) Les lezards du sous-genre malgache de lygodactyles Domerguella (Gekkonides). Bulletin de la Societe Zoologique de France, 92, 583 - 597." type="journal article" year="1967">Pasteur &amp; Blanc, 1967</bibRefCitation>
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<taxonomicName id="C0A2F48B1326BF37FF28F919FC844484" authority="Pasteur &amp; Blanc, 1967" authorityName="Pasteur &amp; Blanc" authorityYear="1967" box="[151,791,1695,1719]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="expectatus" subGenus="Domerguella">
<emphasis id="35D6531A1326BF37FF28F919FDB84484" box="[151,555,1695,1719]" italics="true" pageId="12" pageNumber="13">Lygodactylus (Domerguella) expectatus</emphasis>
<bibRefCitation id="6333F2F91326BF37FD8EF926FC844484" author="Pasteur, G. &amp; Blanc, C. P." box="[561,791,1695,1719]" pageId="12" pageNumber="13" pagination="583 - 597" refId="ref39811" refString="Pasteur, G. &amp; Blanc, C. P. (1967) Les lezards du sous-genre malgache de lygodactyles Domerguella (Gekkonides). Bulletin de la Societe Zoologique de France, 92, 583 - 597." type="journal article" year="1967">Pasteur &amp; Blanc, 1967</bibRefCitation>
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<paragraph id="071D8F081326BF37FF28F95AFE8D44C7" blockId="12.[151,1436,1756,1872]" box="[151,286,1756,1780]" pageId="12" pageNumber="13">Chresonyms:</paragraph>
<paragraph id="071D8F081326BF37FF28F97DFED04501" blockId="12.[151,1436,1756,1872]" pageId="12" pageNumber="13">
<taxonomicName id="C0A2F48B1326BF37FF28F97DFDB84520" authority=": Kluge (1991)" authorityName="Kluge" authorityYear="1991" box="[151,555,1787,1811]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="expectatus">
<emphasis id="35D6531A1326BF37FF28F97DFE0B4520" box="[151,408,1787,1811]" italics="true" pageId="12" pageNumber="13">Lygodactylus expectatus:</emphasis>
<bibRefCitation id="6333F2F91326BF37FE20F97DFDB84520" author="Kluge, A. G." box="[415,555,1787,1811]" pageId="12" pageNumber="13" refId="ref38931" refString="Kluge, A. G. (1991) Checklist of Gekkonoid Lizards. Smithsonian Herpetological Information Service 85, 36 pp. https: // doi. org / 10.5479 / si. 23317515.85.1" type="book" year="1991">Kluge (1991)</bibRefCitation>
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;
<bibRefCitation id="6333F2F91326BF37FD89F97DFC8F4520" author="Glaw, F. &amp; Vences, M." box="[566,796,1787,1811]" pageId="12" pageNumber="13" refId="ref38204" refString="Glaw, F. &amp; Vences, M. (1992) A Fieldguide to the Amphibians and Reptiles of Madagascar. Vences &amp; Glaw Verlags GbR, Cologne, Germany, 331 pp. [First Edition.]" type="book" year="1992">Glaw &amp; Vences (1992</bibRefCitation>
,
<bibRefCitation id="6333F2F91326BF37FC98F97DFCCF4520" author="Glaw, F. &amp; Vences, M." box="[807,860,1787,1811]" pageId="12" pageNumber="13" refId="ref38244" refString="Glaw, F. &amp; Vences, M. (1994) A Fieldguide to the Amphibians and Reptiles of Madagascar. Vences &amp; Glaw Verlags GbR, Cologne, Germany, 480 pp. [Second Edition.]" type="book" year="1994">1994</bibRefCitation>
, 2007);
<bibRefCitation id="6333F2F91326BF37FC0EF97AFBED4520" author="Puente, M. &amp; Glaw, F. &amp; Vieites, D. R. &amp; Vences, M." box="[945,1150,1787,1811]" pageId="12" pageNumber="13" pagination="1 - 76" refId="ref40406" refString="Puente, M., Glaw, F., Vieites, D. R. &amp; Vences, M. (2009) Review of the systematics, morphology and distribution of Malagasy dwarf geckos, genera Lygodactylus and Microscalabotes (Squamata: Gekkonidae). Zootaxa, 2103, 1 - 76. https: // doi. org / 10.11646 / zootaxa. 2103.1.1" type="journal article" year="2009">
Puente
<emphasis id="35D6531A1326BF37FC42F97AFBB94520" box="[1021,1066,1787,1811]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. (2009)
</bibRefCitation>
; Röll
<emphasis id="35D6531A1326BF37FB03F97AFB624520" box="[1212,1265,1787,1811]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(2010);
<bibRefCitation id="6333F2F91326BF37FAF6F97DFED04501" author="Gippner, S. &amp; Travers S. L. &amp; Scherz M. D. &amp; Colston T. J. &amp; Lyra M. L. &amp; Mohan A. V. &amp; Multzsch M. &amp; Nielsen S. V. &amp; Rancilhac L. &amp; Glaw F. &amp; Bauer A. M. &amp; Vences M." pageId="12" pageNumber="13" pagination="107311" refId="ref37826" refString="Gippner, S., Travers S. L., Scherz M. D., Colston T. J., Lyra M. L., Mohan A. V., Multzsch M., Nielsen S. V., Rancilhac L., Glaw F., Bauer A. M. &amp; Vences M. (2021) A comprehensive phylogeny of dwarf geckos of the genus Lygodactylus, with insights into their systematics and morphological variation. Molecular Phylogenetics and Evolution, 165, 107311. https: // doi. org / 10.1016 / j. ympev. 2021.107311" type="journal article" year="2021">
Gippner
<emphasis id="35D6531A1326BF37FF78F89DFF694501" box="[199,250,1818,1842]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(2021)
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<taxonomicName id="C0A2F48B1326BF37FF28F8BEFD414563" authority=": Rosler (2000 b)" authorityName=": Rosler" authorityYear="2000" box="[151,722,1848,1872]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="expectatus" subGenus="Domerguella">
<emphasis id="35D6531A1326BF37FF28F8BEFDA04563" box="[151,563,1848,1872]" italics="true" pageId="12" pageNumber="13">Lygodactylus (Domerguella) expectatus:</emphasis>
<bibRefCitation id="6333F2F91326BF37FD86F8BEFD414563" author="Rosler, H." box="[569,722,1848,1872]" pageId="12" pageNumber="13" pagination="28 - 153" refId="ref41549" refString="Rosler, H. (2000 b) Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota, 2, 28 - 153." type="journal article" year="2000">Rösler (2000b)</bibRefCitation>
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<materialsCitation id="B7CA85551326BF36FF28F8F9FA8142CA" ID-GBIF-Occurrence="3897988318" collectionCode="MNHN" collectorName="G. Pasteur &amp; C. P. Blanc" lastPageId="13" lastPageNumber="14" location="Ambilobe region" pageId="12" pageNumber="13" specimenCode="MNHN 1990.1, MNHN 1990.2, 3" specimenCount="1" typeStatus="holotype">
Name-bearing type: male
<typeStatus id="D81931AA1326BF37FE01F8F9FDB345AA" box="[446,544,1919,1945]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
<specimenCode id="570427731326BF37FD97F8F9FD4045AA" box="[552,723,1919,1945]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="13">MNHN 1990.1</specimenCode>
(original number BP 640).—Type locality: “Karst d’Ambilobe (Ankarana), à une douzaine de kilomètres au NNW de cette localité”, according to the original description.—Other types: according to the original description,
<specimenCount id="11A444811326BF37FD2FF841FCAA45D2" box="[656,825,1991,2017]" pageId="12" pageNumber="13" type="generic">five specimens</specimenCount>
were examined but explicitly only two of these were designated as
<typeStatus id="D81931AA1327BF36FE8BFF11FE314282" box="[308,418,151,177]" pageId="13" pageNumber="14" type="paratype">paratypes</typeStatus>
, namely
<specimenCode id="570427731327BF36FDB9FF11FD224282" box="[518,689,151,177]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="14">MNHN 1990.2</specimenCode>
–
<specimenCode id="570427731327BF36FD7FFF11FD5F4282" box="[704,716,151,177]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="14">3</specimenCode>
(BP 641, female, and BP 642, young female, according to original description).—Etymology: From Latin expectatus = expected. As explained in the original description,
<collectorName id="AA57EADE1327BF36FAF5FF3DFF5042CA" pageId="13" pageNumber="14">G. Pasteur</collectorName>
and
<collectorName id="AA57EADE1327BF36FF46FF59FEFD42CA" box="[249,366,223,249]" pageId="13" pageNumber="14">C.P. Blanc</collectorName>
were expecting to find a new species in the karstic regions of the
<location id="027DD9D31327BF36FBEFFF59FA8142CA" LSID="urn:lsid:plazi:treatment:8F0B3E1E1326BF35FF28F9DCFD1D45EA:027DD9D31327BF36FBEFFF59FA8142CA" box="[1104,1298,223,249]" name="Ambilobe region" pageId="13" pageNumber="14">Ambilobe region</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="53DDDF801327BF36FF28F883FD6B45BA" ID-DOI="http://doi.org/10.5281/zenodo.7040757" ID-Zenodo-Dep="7040757" httpUri="https://zenodo.org/record/7040757/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1797,1821]" targetBox="[151,1436,273,1774]" targetPageId="13">
<paragraph id="071D8F081327BF36FF28F883FD6B45BA" blockId="13.[151,1436,1797,1929]" pageId="13" pageNumber="14">
<emphasis id="35D6531A1327BF36FF28F883FE80452E" bold="true" box="[151,275,1797,1821]" pageId="13" pageNumber="14">FIGURE 6.</emphasis>
View of central dorsum (posterior to forelimb insertion) in seven specimens of
<taxonomicName id="C0A2F48B1327BF36FBFEF883FAA9452E" authorityName=": Rosler" authorityYear="2000" box="[1089,1338,1797,1821]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="expectatus">
<emphasis id="35D6531A1327BF36FBFEF883FAA9452E" box="[1089,1338,1797,1821]" italics="true" pageId="13" pageNumber="14">Lygodactylus expectatus</emphasis>
</taxonomicName>
, and four representative individuals of other species. In all
<emphasis id="35D6531A1327BF36FD3CF8ACFC9D4572" box="[643,782,1834,1857]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="C0A2F48B1327BF36FD3CF8ACFC994572" authorityName=": Rosler" authorityYear="2000" box="[643,778,1834,1857]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="expectatus">L. expectatus</taxonomicName>
,
</emphasis>
including the holotype, the scales in the dorsolateral area are distinctly enlarged, while the vertebral scales are much smaller. In the other species, all scales are roughly of similar size, without obviously enlarged dorsolateral scales. Pictures not to scale.
</paragraph>
</caption>
<subSubSection id="4FB8DC831327BF35FF78F831FBAA4516" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="071D8F081327BF35FF78F831FBD74052" blockId="13.[151,1436,1975,2038]" lastBlockId="14.[151,1436,151,609]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="35D6531A1327BF36FF78F831FE4845E2" bold="true" box="[199,475,1975,2002]" pageId="13" pageNumber="14">Identity and Diagnosis.</emphasis>
According to the diagnosis given by
<bibRefCitation id="6333F2F91327BF36FCC4F83EFBCA45E1" author="Puente, M. &amp; Glaw, F. &amp; Vieites, D. R. &amp; Vences, M." box="[891,1113,1975,2002]" pageId="13" pageNumber="14" pagination="1 - 76" refId="ref40406" refString="Puente, M., Glaw, F., Vieites, D. R. &amp; Vences, M. (2009) Review of the systematics, morphology and distribution of Malagasy dwarf geckos, genera Lygodactylus and Microscalabotes (Squamata: Gekkonidae). Zootaxa, 2103, 1 - 76. https: // doi. org / 10.11646 / zootaxa. 2103.1.1" type="journal article" year="2009">
Puente
<emphasis id="35D6531A1327BF36FC71F83EFB9545E2" box="[974,1030,1975,2001]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
(2009)
</bibRefCitation>
, the species differed from all species in the
<taxonomicName id="C0A2F48B1327BF36FE8CF85AFD8945C6" baseAuthorityName="Boettger" baseAuthorityYear="1881" box="[307,538,2011,2037]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="35D6531A1327BF36FE8CF85AFD8945C6" box="[307,538,2011,2037]" italics="true" pageId="13" pageNumber="14">L. madagascariensis</emphasis>
</taxonomicName>
group known at the time by its dorsolateral scales, which are enlarged relative to the dorsal and lateral scales (not distinctly enlarged in the other species), and by the presence of two dark spots in the region of the neck (not distinct in the other species). The enlarged scales in the dorsolateral region, contrasting with the very small scales in the vertebral region, indeed represent a diagnostic character of this species that we could not observe in any other species of
<taxonomicName id="C0A2F48B1324BF35FE49FE85FD1A432E" authorityName="Pasteur" authorityYear="1964" box="[502,649,259,285]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A1324BF35FE49FE85FD1A432E" box="[502,649,259,285]" italics="true" pageId="14" pageNumber="15">Domerguella</emphasis>
</taxonomicName>
(
<figureCitation id="9F99938D1324BF35FD27FE85FD4F432E" box="[664,732,259,285]" captionStart="FIGURE 6" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[151,1436,273,1774]" captionTargetId="figure-71@13.[151,1436,273,1774]" captionTargetPageId="13" captionText="FIGURE 6. View of central dorsum (posterior to forelimb insertion) in seven specimens of Lygodactylus expectatus, and four representative individuals of other species. In all L. expectatus, including the holotype, the scales in the dorsolateral area are distinctly enlarged, while the vertebral scales are much smaller. In the other species, all scales are roughly of similar size, without obviously enlarged dorsolateral scales. Pictures not to scale." figureDoi="http://doi.org/10.5281/zenodo.7040757" httpUri="https://zenodo.org/record/7040757/files/figure.png" pageId="14" pageNumber="15">Fig. 6</figureCitation>
). This typical character state of
<taxonomicName id="C0A2F48B1324BF35FBFEFE85FB47432F" authorityName=": Rosler" authorityYear="2000" box="[1089,1236,259,284]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="expectatus">
<emphasis id="35D6531A1324BF35FBFEFE85FB47432F" box="[1089,1236,259,284]" italics="true" pageId="14" pageNumber="15">L. expectatus</emphasis>
</taxonomicName>
is visible in all of the genetically characterized specimens collected, as well as the
<typeStatus id="D81931AA1324BF35FCDFFEA1FC514372" box="[864,962,295,321]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(examined in
<date id="731CA9C81324BF35FBDFFEA1FB474372" box="[1120,1236,295,321]" pageId="14" pageNumber="15" value="2021-06">June 2021</date>
, in relatively poor state of preservation). It is also reflected by a low longitudinal count of dorsal scales, of 130 scales or less if counting the enlarged scales (slightly more, with a maximum of 164, if counting the small vertebral scales, but also this value is still smaller than in all other nominal
<taxonomicName id="C0A2F48B1324BF35FD2CFE15FCB5439E" authorityName="Pasteur" authorityYear="1964" box="[659,806,403,429]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A1324BF35FD2CFE15FCB5439E" box="[659,806,403,429]" italics="true" pageId="14" pageNumber="15">Domerguella</emphasis>
</taxonomicName>
, overlapping with only one candidate species,
<emphasis id="35D6531A1324BF35FA92FE15FAD7439F" box="[1325,1348,403,428]" italics="true" pageId="14" pageNumber="15">L.</emphasis>
sp. 17). The dark spot in the region of the neck is located anterodorsal to the forelimb region, roughly in the scapular region, and we here name it the scapular semi-ocellus, considering that it is bordered by a whitish row of tubercles dorsally, giving the impression of an ocellus but lacking a ventral light lining. This semi-ocellus is typical for
<taxonomicName id="C0A2F48B1324BF35FABEFE79FA06402B" authorityName=": Rosler" authorityYear="2000" box="[1281,1429,511,536]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="expectatus">
<emphasis id="35D6531A1324BF35FABEFE79FA06402B" box="[1281,1429,511,536]" italics="true" pageId="14" pageNumber="15">L. expectatus</emphasis>
</taxonomicName>
, but sometimes weakly expressed, and in such cases easy to confuse with dark lateral markings that can also be seen in other species of
<taxonomicName id="C0A2F48B1324BF35FED4FDC1FE6D4052" authorityName="Pasteur" authorityYear="1964" box="[363,510,583,609]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A1324BF35FED4FDC1FE6D4052" box="[363,510,583,609]" italics="true" pageId="14" pageNumber="15">Domerguella</emphasis>
</taxonomicName>
, but often in slightly different positions (
<figureCitation id="9F99938D1324BF35FC75FDC1FBA54052" box="[970,1078,583,609]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="14.[151,250,1336,1360]" captionStartId-1="15.[151,250,1636,1660]" captionTargetBox-0="[155,1436,633,1313]" captionTargetBox-1="[151,1436,181,1612]" captionTargetId-0="figure-275@14.[151,1436,633,1313]" captionTargetId-1="figure-18@15.[151,1436,181,1612]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURE 7. Lateral view of head and anterior body in seven specimens of Lygodactylus expectatus, and four representative individuals of other species. In all L. expectatus, a black spot is visible in the shoulder region, with at least traces of light color along its dorsal edge; this spot, poorly expressed in some specimens, is here called the scapular semi-ocellus. Individuals of other species of Domerguella may also have dark markings in the scapular region, but these usually are not at all bordered by light color dorsally, and often are in a more anterior position, as exemplified by four representative pictures of other species. Asterisks mark photos that were mirrored. Pictures not to scale." captionText-1="FIGURE 8. Photos of specimens of Lygodactylus expectatus from Ankarana in life. A, B, Adult male ZSM 284/2004 (FGZC 543), photographed in 2004. C, D, E, Specimen photographed in 2001 by Gerardo García, not collected." figureDoi-0="http://doi.org/10.5281/zenodo.7040761" figureDoi-1="http://doi.org/10.5281/zenodo.7040763" httpUri-0="https://zenodo.org/record/7040761/files/figure.png" httpUri-1="https://zenodo.org/record/7040763/files/figure.png" pageId="14" pageNumber="15">Figs. 7–8</figureCitation>
).
</paragraph>
<caption id="53DDDF801324BF35FF28FABEFC8F4436" ID-DOI="http://doi.org/10.5281/zenodo.7040761" ID-Zenodo-Dep="7040761" httpUri="https://zenodo.org/record/7040761/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1336,1360]" targetBox="[155,1436,633,1313]" targetPageId="14">
<paragraph id="071D8F081324BF35FF28FABEFC8F4436" blockId="14.[151,1437,1336,1541]" pageId="14" pageNumber="15">
<emphasis id="35D6531A1324BF35FF28FABEFE864762" bold="true" box="[151,277,1336,1361]" pageId="14" pageNumber="15">FIGURE 7.</emphasis>
Lateral view of head and anterior body in seven specimens of
<taxonomicName id="C0A2F48B1324BF35FC18FABFFB324763" authorityName=": Rosler" authorityYear="2000" box="[935,1185,1337,1361]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="expectatus">
<emphasis id="35D6531A1324BF35FC18FABFFB324763" box="[935,1185,1337,1361]" italics="true" pageId="14" pageNumber="15">Lygodactylus expectatus</emphasis>
</taxonomicName>
, and four representative individuals of other species. In all
<emphasis id="35D6531A1324BF35FE49FADBFD114747" box="[502,642,1373,1396]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="C0A2F48B1324BF35FE49FADBFDED4747" authorityName=": Rosler" authorityYear="2000" box="[502,638,1373,1396]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="expectatus">L. expectatus</taxonomicName>
,
</emphasis>
a black spot is visible in the shoulder region, with at least traces of light color along its dorsal edge; this spot, poorly expressed in some specimens, is here called the scapular semi-ocellus. Individuals of other species of
<taxonomicName id="C0A2F48B1324BF35FE80FA23FE57478E" authorityName="Pasteur" authorityYear="1964" box="[319,452,1445,1469]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A1324BF35FE80FA23FE57478E" box="[319,452,1445,1469]" italics="true" pageId="14" pageNumber="15">Domerguella</emphasis>
</taxonomicName>
may also have dark markings in the scapular region, but these usually are not at all bordered by light color dorsally, and often are in a more anterior position, as exemplified by four representative pictures of other species. Asterisks mark photos that were mirrored. Pictures not to scale.
</paragraph>
</caption>
<paragraph id="071D8F081324BF35FF78F9B5FD7644A6" blockId="14.[151,1437,1587,2009]" pageId="14" pageNumber="15">
Given these two diagnostic character states, which both have been verified in the
<typeStatus id="D81931AA1324BF35FBF2F9B5FB3C447E" box="[1101,1199,1587,1613]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
and in the genotyped specimens, along with the provenance of all these specimens from the Ankarana Massif, there is no doubt about the correct attribution of our specimens to
<taxonomicName id="C0A2F48B1324BF35FDF4F9FAFD4D44A6" authorityName=": Rosler" authorityYear="2000" box="[587,734,1660,1685]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="expectatus">
<emphasis id="35D6531A1324BF35FDF4F9FAFD4D44A6" box="[587,734,1660,1685]" italics="true" pageId="14" pageNumber="15">L. expectatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="071D8F081324BF35FF78F919FBAA4516" blockId="14.[151,1437,1587,2009]" pageId="14" pageNumber="15">
The species is rather small sized, with adult SVL
<quantity id="C05A22ED1324BF35FD50F919FC01448A" box="[751,914,1695,1721]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="2.9699999999999998" metricValueMin="2.43" pageId="14" pageNumber="15" unit="mm" value="27.0" valueMax="29.7" valueMin="24.3">24.3–29.7 mm</quantity>
vs. a maximum size larger than
<quantity id="C05A22ED1324BF35FB43F919FAD8448A" box="[1276,1355,1695,1721]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="mm" value="30.0">30 mm</quantity>
in several other species. There are no dorsolateral tubercles and no spiny tubercles at the tail base as they are characteristic for several other
<taxonomicName id="C0A2F48B1324BF35FEE8F961FE794532" authorityName="Pasteur" authorityYear="1964" box="[343,490,1767,1793]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A1324BF35FEE8F961FE794532" box="[343,490,1767,1793]" italics="true" pageId="14" pageNumber="15">Domerguella</emphasis>
</taxonomicName>
, and no distinct, regular broad crossbands on the tail as in
<taxonomicName id="C0A2F48B1324BF35FB3EF96EFB4F4532" box="[1153,1244,1768,1793]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rarus">
<emphasis id="35D6531A1324BF35FB3EF96EFB4F4532" box="[1153,1244,1768,1793]" italics="true" pageId="14" pageNumber="15">L. rarus</emphasis>
</taxonomicName>
(see below). According to the available counts, the species has 87–98 ventral scales longitudinally.
</paragraph>
</subSubSection>
<subSubSection id="4FB8DC831324BF35FF78F8A9FD1D45EA" pageId="14" pageNumber="15" type="distribution">
<paragraph id="071D8F081324BF35FF78F8A9FD1D45EA" blockId="14.[151,1437,1587,2009]" pageId="14" pageNumber="15">
<emphasis id="35D6531A1324BF35FF78F8A9FEF2457A" bold="true" box="[199,353,1839,1865]" pageId="14" pageNumber="15">Distribution.</emphasis>
<taxonomicName id="C0A2F48B1324BF35FED9F8B6FE6A457A" authorityName=": Rosler" authorityYear="2000" box="[358,505,1840,1865]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="expectatus">
<emphasis id="35D6531A1324BF35FED9F8B6FE6A457A" box="[358,505,1840,1865]" italics="true" pageId="14" pageNumber="15">L. expectatus</emphasis>
</taxonomicName>
is only known from its
<typeStatus id="D81931AA1324BF35FD40F8A9FCBC457A" box="[767,815,1839,1865]" pageId="14" pageNumber="15">type</typeStatus>
locality, the Ankarana Massif. According to the original description (
<bibRefCitation id="6333F2F91324BF35FE9EF8D5FD88455E" author="Pasteur, G. &amp; Blanc, C. P." box="[289,539,1875,1901]" pageId="14" pageNumber="15" pagination="583 - 597" refId="ref39811" refString="Pasteur, G. &amp; Blanc, C. P. (1967) Les lezards du sous-genre malgache de lygodactyles Domerguella (Gekkonides). Bulletin de la Societe Zoologique de France, 92, 583 - 597." type="journal article" year="1967">Pasteur &amp; Blanc 1967</bibRefCitation>
), additional specimens also came from “Ambilobé” and from “Region de DiégoSuarez”, but we have not verified the identity of the respective vouchers, and the localities are not precise enough for firmly concluding they are not in the Ankarana Massif (which is geographically located inbetween the towns of Ambilobe and
<collectingRegion id="C56641EA1324BF35FE83F839FE5045EA" box="[316,451,1983,2009]" country="Madagascar" name="Antsiranana" pageId="14" pageNumber="15">Antsiranana</collectingRegion>
(=Diego-Suarez).
</paragraph>
</subSubSection>
</treatment>
</document>
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