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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.185.3082" ID-GBIF-Dataset="f54227ef-6fd3-41eb-a6fc-6203e8ed2ec5" ID-PMC="PMC3345791" ID-Pensoft-Pub="1313-2970-185-1" ID-PubMed="22577310" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-185-1" ModsDocOrigin="ZooKeys 185" ModsDocTitle="Three new cavernicolous species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from southern China, with notes on a formal congener from the Philippines (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249126133" checkinUser="pensoft" docAuthor="Golovatch, Sergei I., Li, Youbang, Liu, Weixin &amp; Geoffroy, Jean-Jacques" docDate="2012" docId="8E8A37DD0AD8B3E3427B8DE9F9062757" docLanguage="en" docName="ZooKeys 185: 1-17" docOrigin="ZooKeys 185" docSource="http://dx.doi.org/10.3897/zookeys.185.3082" docTitle="Desmoxytes lui Golovatch, Li, Liu &amp; Geoffroy, 2012, sp. n." docType="treatment" docVersion="4" lastPageNumber="11" masterDocId="B12DC7150217FFBB535C5A786D46FFD9" masterDocTitle="Three new cavernicolous species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from southern China, with notes on a formal congener from the Philippines (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="9" updateTime="1668153676840" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new cavernicolous species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from southern China, with notes on a formal congener from the Philippines (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Golovatch, Sergei I.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Youbang</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Weixin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Geoffroy, Jean-Jacques</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>185</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>17</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.185.3082</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.185.3082</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-185-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152034747" LSID="urn:lsid:zoobank.org:act:A0C24C82-1F34-437A-BCFA-C28A1E3BA72F" httpUri="http://treatment.plazi.org/id/8E8A37DD0AD8B3E3427B8DE9F9062757" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection pageId="8" pageNumber="9" type="nomenclature">
<paragraph pageId="8" pageNumber="9">
<taxonomicName LSID="urn:lsid:zoobank.org:act:A0C24C82-1F34-437A-BCFA-C28A1E3BA72F" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes lui" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lui">Desmoxytes lui</taxonomicName>
<taxonomicNameLabel pageId="8" pageNumber="9">sp. n.</taxonomicNameLabel>
Figs 56
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="holotype">
<paragraph pageId="8" pageNumber="9">Holotype.</paragraph>
<paragraph pageId="8" pageNumber="9">♂(IZAS), China, Guangxi Prov., Yongfu County, Shangxiao, Cave Dachong Dong, 17.I.2012, leg. Li Youbang and Lu Shiyi (CHIgx12-LYB03).</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="paratypes">
<paragraph pageId="8" pageNumber="9">Paratypes.</paragraph>
<paragraph pageId="8" pageNumber="9">1 ♀ (GNUG), 1 ♀ juv. (MNHN JA 131), same locality, together with holotype.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="name">
<paragraph pageId="8" pageNumber="9">Name.</paragraph>
<paragraph pageId="8" pageNumber="9">To honour Mr Lu Shiyi, one of the collectors.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="diagnosis">
<paragraph pageId="8" pageNumber="9">Diagnosis.</paragraph>
<paragraph pageId="8" pageNumber="9">Differs in the paraterga being spiniform only until segment 5, combined with the gonopods being strongly condensed, ♂femora 6 humped. See also Key below.</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph pageId="8" pageNumber="9">Description.</paragraph>
<paragraph pageId="8" pageNumber="9">Length ca 36 (♂), 43 mm (♀) or 40 mm (1 ♀ juvenile with 19 segments); width of pro- and metaterga together with paraterga 2.0 and 2.2 (♂), 3.2 and 3.4 mm (♀) or 3.0 and 3.1 mm (1 ♀ juvenile with 19 segments), respectively.</paragraph>
<paragraph pageId="8" pageNumber="9">
Body strongly moniliform (Fig. 5
<normalizedToken originalValue="AC">A-C</normalizedToken>
). Coloration of alcohol material rather uniformly light pink-brownish to nearly pallid, anterior body part a little darker, pinkish (Fig. 5). Antennomere 7 dark brown (even in the juvenile). In width, segment 3 = 4 &lt;head &lt;collum = segment 2 &lt;6 = 16; thereafter body gradually tapering towards telson. Head very densely setose throughout, epicranial suture distinct. Antennae extremely long and slender, reaching back to segment 9 (♂) or 7 (♀) dorsally, antennomeres 5 and 6 each with an apicodorsal compact group of bacilliform sensilla.
</paragraph>
<paragraph pageId="8" pageNumber="9">Tegument rather smooth and poorly shining, prozona very delicately microalveolate, metaterga finely shagreened to microgranulate/microspiculate, surface below paraterga finely shagreened to microgranulate. Collum with 6+6(7) evident setigerous spinules in a row at front margin, behind it with about 4+4 smaller spinules in an irregular transverse row.</paragraph>
<paragraph lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Following metaterga with a pattern of similar, smaller, 3+3 and 3+3 spinules in two transverse rows, last row gradually growing up to 6+6 until segment 19. Collum and following metaterga 2-4 with straight, spiniform paraterga, about as long as body height in ♂, a little shorter in ♀ and juvenile (Fig. 5
<normalizedToken originalValue="AC">A-C</normalizedToken>
), directed much more dorsally than laterally and ending up clearly above dorsum; paraterga 5 only slightly shorter than preceding ones; paraterga 6-18 contrasting short, coni- to dentiform; paraterga 19 subhorizontal, about level with dorsum, directed clearly caudad and reaching behind until about midlength (♂) or front third (♀) along telson. Paraterga 1-5 with two rather evident, setigerous indentations/spinules/knobs frontally and a short tooth posteriorly at base (Fig. 5A). Stricture between pro- and metazona very broad and shallow, always smooth at bottom (Fig. 5B, C). Pore formula normal; ozopores inconspicuous, located just at base on lateral side of poriferous paraterga. Transverse sulcus usually very vague, traceable in segments 4-18. Pleurosternal carinae evident only in segments 2 and 3 (Fig. 5A). Epiproct (Fig. 5C, D) rather simple, lateral pre-apical papillae very distinct, finger-shaped. Hypoproct (Fig. 5D) subtrapeziform,
<pageBreakToken pageId="9" pageNumber="10" start="start">caudal</pageBreakToken>
margin emarginate, setigerous cones at caudal edge very large, widely separated. Axial line missing.
</paragraph>
<paragraph pageId="9" pageNumber="10">Sterna quite densely setose, cross-impressions faint. A paramedian pair of entirely separated, short, rounded, setose tubercles between ♂ coxae 4 (Fig. 6A). Legs (Fig. 5A, B) extremely long and slender, ca 6.0 (♂) or 4.5 (♀) times longer than midbody height. ♂femora 6 strongly inflated ventrally in distal quarter (Fig. 5B).</paragraph>
<paragraph pageId="9" pageNumber="10">
Gonopods (Fig. 6
<normalizedToken originalValue="BD">B-D</normalizedToken>
) simple. Coxite rather short, subcylindrical, poorly setose distodorsally, about 1/3 times as long as telopodite. Prefemoral portion (= densely setose) about half as long as acropodite. Femorite (fe) rather slender, elongate, only slightly enlarged distad, with seminal groove running entirely on mesal face, also with a distinct sulcus demarcating a short, strongly condensed solenophore (sph). The latter distinguished by a somewhat folded lamina medialis (lm) and a simpler lamina lateralis (ll). Solenomere (sl) similarly short, flagelliform, evidently separated at base from solenophore.
</paragraph>
<caption pageId="10" pageNumber="11">
<paragraph pageId="10" pageNumber="11">
<pageBreakToken pageId="10" pageNumber="11" start="start">Figure</pageBreakToken>
5.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes lui" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lui">Desmoxytes lui</taxonomicName>
sp. n., ♂ holotype. A anterior part of body, lateral view B body segments 5 to 8, lateral view C posterior part of body, dorsal view D telson, ventral view.Photographed not to scale.
</paragraph>
</caption>
<caption pageId="10" pageNumber="11">
<paragraph pageId="10" pageNumber="11">
Figure 6.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes lui" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lui">Desmoxytes lui</taxonomicName>
sp. n., ♂ holotype. A sternal cones between coxae 4, ventral view
<normalizedToken originalValue="BD">B-D</normalizedToken>
right gonopod telopodite, mesal, dorsal and sublateral views, respectively. Scale bar: 1.0 (A) and 0.5 mm (
<normalizedToken originalValue="BD">B-D</normalizedToken>
). Designations: fe femorite sph solenophore sl solenomere ll lamina lateralis lm lamina medialis
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>