treatments-xml/data/86/32/84/8632842AFF9D1D1DA624FE82FBE3F895.xml

<document id="D361A9A89E4FF530907CE263FA6E8997" ID-CLB-Dataset="51374" ID-DOI="10.5281/zenodo.156789" ID-GBIF-Dataset="49eb0094-f3f4-4741-9a8d-88af88ca478a" ID-ISSN="1175­5326" ID-Zenodo-Dep="156789" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459583253783" checkinUser="plazi" docAuthor="Solís, Ángel &amp; Kohlmann, Bert" docDate="2003" docId="8632842AFF9D1D1DA624FE82FBE3F895" docLanguage="en" docName="zt00139.pdf" docOrigin="Zootaxa 139" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Copris tridentatus Solis &amp; Kohlmann, sp. nov." docType="treatment" docVersion="10" lastPageNumber="2" masterDocId="7A0BFC52FF9C1D1CA72CFF9EFFF7FFE5" masterDocTitle="New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama" masterLastPageNumber="14" masterPageNumber="1" pageNumber="2" updateTime="1698202942371" updateUser="ExternalLinkService">
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<mods:title id="71E2D495E269EADDB8FE5CC27B0CD111">New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama</mods:title>
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<mods:namePart id="EDB7F9B877B442FB6AEE007297059051">Solís, Ángel</mods:namePart>
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<mods:namePart id="E4C0BFDE6C0D3356AA69123C600F3DA5">Kohlmann, Bert</mods:namePart>
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<mods:date id="63AF8485CC5DF426ECB7147E677F07D2">2003</mods:date>
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<treatment id="8632842AFF9D1D1DA624FE82FBE3F895" ID-DOI="http://doi.org/10.5281/zenodo.6274456" ID-GBIF-Taxon="119330984" ID-Zenodo-Dep="6274456" LSID="urn:lsid:plazi:treatment:8632842AFF9D1D1DA624FE82FBE3F895" httpUri="http://treatment.plazi.org/id/8632842AFF9D1D1DA624FE82FBE3F895" lastPageNumber="2" pageId="1" pageNumber="2">
<subSubSection id="468166B7FF9D1D1DA624FE82FC69FED0" box="[264,926,283,310]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="0E24353CFF9D1D1DA624FE82FC69FED0" blockId="1.[264,926,283,310]" box="[264,926,283,310]" pageId="1" pageNumber="2">
<heading id="556C8250FF9D1D1DA624FE82FC69FED0" bold="true" box="[264,926,283,310]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="3CEFE92EFF9D1D1DA624FE82FC69FED0" bold="true" box="[264,926,283,310]" pageId="1" pageNumber="2">
<taxonomicName id="C99B4EBFFF9D1D1DA624FE82FD25FED0" ID-CoL="7ZMN8" authority="Solis &amp; Kohlmann" authorityName="Solis &amp; Kohlmann" box="[264,722,283,310]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tridentatus" status="sp. nov.">
<emphasis id="3CEFE92EFF9D1D1DA624FE82FE16FED3" bold="true" box="[264,481,284,310]" italics="true" pageId="1" pageNumber="2">Copris tridentatus</emphasis>
Solís &amp; Kohlmann
</taxonomicName>
, 
<taxonomicNameLabel id="27DC5455FF9D1D1DA5F3FE85FCCAFED0" box="[735,829,283,309]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="96A029B9FF9D1D1DA460FE85FC63FED0" box="[844,916,283,309]" captionStart="FIGURE 1" captionStartId="4.[264,368,1805,1829]" captionTargetBox="[300,1291,334,1765]" captionTargetId="figure@4.[288,1300,329,1779]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal habitus of Copris tridentatus Solís &amp; Kohlmann, sp. nov. (holotype)." httpUri="https://zenodo.org/record/168690/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF9D1D1DA624FEF8FDF2FE1C" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="0E24353CFF9D1D1DA624FEF8FDF2FE1C" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">
<emphasis id="3CEFE92EFF9D1D1DA624FEF8FE7FFE65" bold="true" box="[264,392,358,384]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
This species is distinguished from other 
<taxonomicName id="C99B4EBFFF9D1D1DA443FEF9FC48FE65" box="[879,959,359,384]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3CEFE92EFF9D1D1DA443FEF9FC48FE65" box="[879,959,359,384]" italics="true" pageId="1" pageNumber="2">Copris</emphasis>
</taxonomicName>
species by the following combination of characters: head with a tridentate clypeus; head and thorax are unarmed, the head having only a small gibbosity; head and pronotum covered by large, flat and shallow umbilicate punctures.
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF9D1D1DA614FD98FDC9FAFD" pageId="1" pageNumber="2" type="description">
<paragraph id="0E24353CFF9D1D1DA614FD98FD99FDAC" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">
<emphasis id="3CEFE92EFF9D1D1DA614FD98FD68FDC5" bold="true" box="[312,671,518,544]" pageId="1" pageNumber="2">
Description. 
<typeStatus id="D1208B9EFF9D1D1DA6FBFD98FDBCFDC5" box="[471,587,518,544]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
. Male:
</emphasis>
Length 
<quantity id="C96398D9FF9D1D1DA42CFD99FC9BFDC4" box="[768,876,519,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="1" pageNumber="2" unit="mm" value="13.2">13.2 mm</quantity>
. Humeral width 
<quantity id="C96398D9FF9D1D1DA302FD99FB7CFDC4" box="[1070,1163,519,545]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="2" unit="mm" value="7.1">7.1 mm</quantity>
. Body oblong and convex, completely black.
</paragraph>
<paragraph id="0E24353CFF9D1D1DA614FDC9FC3EFD0C" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">Head unarmed; clypeus tridentate, teeth not prominent; clypeus with faint simple punctation; genae and base of head with coarse umbilical punctation; frons with a smooth gibbosity; eyes small, maximum ocular width about two thirds of ocular length; genal suture slightly arched; posterior angles of head subquadrate.</paragraph>
<paragraph id="0E24353CFF9D1D1DA614FD69FD35FC6C" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">Pronotum unarmed; anterior angles acute, the margin behind them sinuate; lateral carina acute; coarse umbilicate punctures all around pronotal margin, and on anterior angles, lateral fossae, median longitudinal sulcus and to both sides of the pronotal disc; median longitudinal sulcus deeply impressed.</paragraph>
<paragraph id="0E24353CFF9D1D1DA614FC09FDBCFBE4" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">
Elytra with eighth and tenth striae complete, ninth stria arising at about one third of elytron; striae crenate­punctate, separated by 1­ 
<date id="7A2513FCFF9D1D1DA41BFC21FCBEFC3C" box="[823,841,959,985]" pageId="1" pageNumber="2">2</date>
x their widths on disc; intervals convex, sparsely minutely punctate.
</paragraph>
<paragraph id="0E24353CFF9D1D1DA614FB90FBFFFBCD" blockId="1.[264,1325,358,1904]" box="[312,1032,1038,1064]" pageId="1" pageNumber="2">Pygidium coarsely and uniformly punctate, margin complete.</paragraph>
<paragraph id="0E24353CFF9D1D1DA614FBA8FBDEFB45" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">Protibial spurs curved slightly inward and tapering to an acute point, pro­ meso­ and metafemora covered ventrally with coarse umbilicate punctures, as well as a great part of the lower surfaces with the exception of the disc of the metasternum</paragraph>
<paragraph id="0E24353CFF9D1D1DA614FB30FDD9FB2D" blockId="1.[264,1325,358,1904]" box="[312,558,1198,1224]" pageId="1" pageNumber="2">
<emphasis id="3CEFE92EFF9D1D1DA614FB30FE64FB2D" bold="true" box="[312,403,1198,1224]" pageId="1" pageNumber="2">Female</emphasis>
. ­ Unknown.
</paragraph>
<paragraph id="0E24353CFF9D1D1DA614FB48FDC9FAFD" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">
<emphasis id="3CEFE92EFF9D1D1DA614FB48FE45FB15" bold="true" box="[312,434,1238,1264]" pageId="1" pageNumber="2">Variation.</emphasis>
Length 
<quantity id="C96398D9FF9D1D1DA533FB48FD3BFB15" box="[543,716,1238,1264]" metricMagnitude="-2" metricUnit="m" metricValue="1.285" metricValueMax="1.3199999999999998" metricValueMin="1.25" pageId="1" pageNumber="2" unit="mm" value="12.85" valueMax="13.2" valueMin="12.5">12.5­13.2 mm</quantity>
. Humeral width 
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. Frons gibbosity can also be umbilico­punctate.
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF9D1D1DA614FAB8FD58FA5D" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="0E24353CFF9D1D1DA614FAB8FB16FA8D" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">
<emphasis id="3CEFE92EFF9D1D1DA614FAB8FDDFFAA5" bold="true" box="[312,552,1318,1344]" pageId="1" pageNumber="2">Examined material</emphasis>
(
<specimenCount id="189DFEB5FF9D1D1DA51BFAB8FD39FAA5" box="[567,718,1318,1344]" pageId="1" pageNumber="2" type="generic">2 specimens</specimenCount>
). 
<emphasis id="3CEFE92EFF9D1D1DA5CCFAB8FCA5FAA5" bold="true" box="[736,850,1318,1344]" pageId="1" pageNumber="2">
<typeStatus id="D1208B9EFF9D1D1DA5CCFAB8FCA5FAA5" box="[736,850,1318,1344]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
</emphasis>
, male: 
<collectingCountry id="768C75ACFF9D1D1DA484FAB8FBA0FAA5" box="[936,1111,1318,1344]" name="Costa Rica" pageId="1" pageNumber="2">COSTA RICA</collectingCountry>
. San José: Ciudad Colón, El Rodeo, 
<quantity id="C96398D9FF9D1D1DA6F7FAD0FDC2FA8D" box="[475,565,1358,1384]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="2" unit="m" value="1000.0">1000 m</quantity>
, 
<date id="7A2513FCFF9D1D1DA56EFAD0FD22FA8D" box="[578,725,1358,1384]" pageId="1" pageNumber="2" value="1988-10-17">17 Oct 1988</date>
, Col.: A. Solís. 
<emphasis id="3CEFE92EFF9D1D1DA4BCFAD0FBFEFA8D" bold="true" box="[912,1033,1358,1384]" pageId="1" pageNumber="2">
<typeStatus id="D1208B9EFF9D1D1DA4BCFAD0FBF3FA8D" box="[912,1028,1358,1384]" pageId="1" pageNumber="2" type="paratype">Paratype</typeStatus>
.
</emphasis>
<emphasis id="3CEFE92EFF9D1D1DA33DFAD1FBC9FA8D" box="[1041,1086,1359,1384]" italics="true" pageId="1" pageNumber="2">ibid</emphasis>
., 
<specimenCount id="189DFEB5FF9D1D1DA378FAD0FB2BFA8D" box="[1108,1244,1358,1384]" pageId="1" pageNumber="2" type="generic">1 specimen</specimenCount>
.
</paragraph>
<paragraph id="0E24353CFF9D1D1DA614FAE8FD58FA5D" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">
<emphasis id="3CEFE92EFF9D1D1DA614FAE8FE68FA75" bold="true" box="[312,415,1398,1424]" pageId="1" pageNumber="2">Habitat.</emphasis>
The specimens were collected using traps baited with human excrement inside a tropical seasonal dry forest.
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF9D1D1DA614FA58FC07F8AD" pageId="1" pageNumber="2" type="description">
<paragraph id="0E24353CFF9D1D1DA614FA58FC18F9ED" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">
<emphasis id="3CEFE92EFF9D1D1DA614FA58FD70FA05" bold="true" box="[312,647,1478,1504]" pageId="1" pageNumber="2">Geographical distribution.</emphasis>
This species is only known so far from the “El Rodeo” Forest Reseve, near San José, at the base of the Escazú heights.
</paragraph>
<paragraph id="0E24353CFF9D1D1DA614F988FADDF9BD" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">
<emphasis id="3CEFE92EFF9D1D1DA614F988FDA1F9D5" bold="true" box="[312,598,1558,1584]" pageId="1" pageNumber="2">Chorological affinities.</emphasis>
The distribution of this new species is embedded within the distribution of its closest relative, 
<taxonomicName id="C99B4EBFFF9D1D1DA5BBF9A1FC5FF9BD" authority="Gahan" authorityName="Gahan" box="[663,936,1598,1624]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="3CEFE92EFF9D1D1DA5BBF9A1FCA4F9BD" box="[663,851,1599,1624]" italics="true" pageId="1" pageNumber="2">C. costaricensis</emphasis>
Gahan
</taxonomicName>
, found from Chiapas to 
<collectingCountry id="768C75ACFF9D1D1DA3E9F9A0FAD1F9BD" box="[1221,1318,1598,1624]" name="Panama" pageId="1" pageNumber="2">Panama</collectingCountry>
.
</paragraph>
<paragraph id="0E24353CFF9D1D1DA614F9F8FC07F8AD" blockId="1.[264,1325,358,1904]" pageId="1" pageNumber="2">
<emphasis id="3CEFE92EFF9D1D1DA614F9F8FD87F965" bold="true" box="[312,624,1638,1664]" pageId="1" pageNumber="2">Taxonomic relationships.</emphasis>
<taxonomicName id="C99B4EBFFF9D1D1DA552F9F9FCA2F965" box="[638,853,1639,1664]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tridentatus">
<emphasis id="3CEFE92EFF9D1D1DA552F9F9FCA2F965" box="[638,853,1639,1664]" italics="true" pageId="1" pageNumber="2">Copris tridentatus</emphasis>
</taxonomicName>
is in the remotus complex as defined by 
<bibRefCitation id="6A0A48CDFF9D1D1DA624F910FE21F94D" author="Matthews" box="[264,470,1678,1704]" pageId="1" pageNumber="14" refString="Matthews, E. (1961) A revision of the genus Copris Muller of the Western Hemisphere (Coleoptera, Scarabaeidae). Entomologica Americana, 41, 1 - 137." type="journal article" year="1961">Matthews (1961)</bibRefCitation>
, and is keyed there to 
<taxonomicName id="C99B4EBFFF9D1D1DA5C0F911FC17F94D" box="[748,992,1679,1704]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="costaricencis">
<emphasis id="3CEFE92EFF9D1D1DA5C0F911FC17F94D" box="[748,992,1679,1704]" italics="true" pageId="1" pageNumber="2">Copris costaricencis</emphasis>
</taxonomicName>
. It is easily separated from this species by the presence of a tridentate clypeus, anterior pronotal margin without a central tooth, a complete pygidial margin and the lack of a male horn. In general the species has characters intermediate between 
<taxonomicName id="C99B4EBFFF9D1D1DA593F899FC8BF8C5" box="[703,892,1799,1824]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="3CEFE92EFF9D1D1DA593F899FC8BF8C5" box="[703,892,1799,1824]" italics="true" pageId="1" pageNumber="2">C. costaricensis</emphasis>
</taxonomicName>
and 
<emphasis id="3CEFE92EFF9D1D1DA494F899FBD3F8C5" box="[952,1060,1799,1824]" italics="true" pageId="1" pageNumber="2">
<taxonomicName id="C99B4EBFFF9D1D1DA494F899FBE9F8C5" box="[952,1054,1799,1824]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="sallei">C. sallei</taxonomicName>
,
</emphasis>
although the pronotal punctation has basically the same pattern as in 
<taxonomicName id="C99B4EBFFF9D1D1DA402F8B1FC1EF8AD" box="[814,1001,1839,1864]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="3CEFE92EFF9D1D1DA402F8B1FC1EF8AD" box="[814,1001,1839,1864]" italics="true" pageId="1" pageNumber="2">C. costaricensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF9D1D1DA614F8C8FBE3F895" box="[312,1044,1878,1904]" pageId="1" pageNumber="2" type="etymology">
<paragraph id="0E24353CFF9D1D1DA614F8C8FBE3F895" blockId="1.[264,1325,358,1904]" box="[312,1044,1878,1904]" pageId="1" pageNumber="2">
<emphasis id="3CEFE92EFF9D1D1DA614F8C8FE32F895" bold="true" box="[312,453,1878,1904]" pageId="1" pageNumber="2">Etymology.</emphasis>
The name refers to the clypeus having three teeth.
</paragraph>
</subSubSection>
</treatment>
</document>