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<document id="07FDB3518EED6DDDC22E22824A79FD5B" ID-CLB-Dataset="51374" ID-DOI="10.5281/zenodo.156789" ID-GBIF-Dataset="49eb0094-f3f4-4741-9a8d-88af88ca478a" ID-ISSN="1175­5326" ID-Zenodo-Dep="156789" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459583253783" checkinUser="plazi" docAuthor="Solís, Ángel &amp; Kohlmann, Bert" docDate="2003" docId="8632842AFF951D11A624FD4AFE00FEBB" docLanguage="en" docName="zt00139.pdf" docOrigin="Zootaxa 139" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Trichillum (Eutrichillum) arcus Solis &amp; Kohlmann, sp. nov." docType="treatment" docVersion="10" lastPageNumber="14" masterDocId="7A0BFC52FF9C1D1CA72CFF9EFFF7FFE5" masterDocTitle="New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama" masterLastPageNumber="14" masterPageNumber="1" pageNumber="10" updateTime="1698202942371" updateUser="ExternalLinkService">
<mods:mods id="AB833A8AB5CB0B256C84A5B2E75063B2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="09ABB60024A5726EAE8936DA77B070FA">New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama</mods:title>
</mods:titleInfo>
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<mods:role id="0864AB7EC664AC2B8A630FC865C24702">
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<mods:namePart id="8961DAB65629EBB44BDCEE75675FAD9F">Solís, Ángel</mods:namePart>
</mods:name>
<mods:name id="0800047C73D2B71901B81E7E221280FF" type="personal">
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<mods:namePart id="35DB5FE228F596BD1263D2811A7ABAEE">Kohlmann, Bert</mods:namePart>
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<mods:typeOfResource id="28D57CAFF0700ABC1C8F23C1A98FDF5C">text</mods:typeOfResource>
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<mods:title id="FD58ED59ED95865CF41828DB135863D4">Zootaxa</mods:title>
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<mods:date id="5DD378D309F3DBB3F544849A4B934FBD">2003</mods:date>
<mods:detail id="03E578551E2E7408EAC921F84BBD658B" type="volume">
<mods:number id="8A6E219310ABA591C6171AE4DF175F84">139</mods:number>
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<mods:classification id="ACA47737061A946C58DA22324963DA82">journal article</mods:classification>
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<mods:identifier id="53B92297E64FC62D54C66999BBF45D80" type="DOI">10.5281/zenodo.156789</mods:identifier>
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<mods:identifier id="16095BAF112E9DEA3573B63F3D41A89D" type="ISSN">1175­5326</mods:identifier>
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<treatment id="8632842AFF951D11A624FD4AFE00FEBB" ID-DOI="http://doi.org/10.5281/zenodo.6274464" ID-GBIF-Taxon="119330986" ID-Zenodo-Dep="6274464" LSID="urn:lsid:plazi:treatment:8632842AFF951D11A624FD4AFE00FEBB" httpUri="http://treatment.plazi.org/id/8632842AFF951D11A624FD4AFE00FEBB" lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10">
<subSubSection id="468166B7FF951D15A624FD4AFBB1FD08" box="[264,1094,723,750]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="0E24353CFF951D15A624FD4AFBB1FD08" blockId="9.[264,1094,723,750]" box="[264,1094,723,750]" pageId="9" pageNumber="10">
<heading id="556C8250FF951D15A624FD4AFBB1FD08" bold="true" box="[264,1094,723,750]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="3CEFE92EFF951D15A624FD4AFBB1FD08" bold="true" box="[264,1094,723,750]" pageId="9" pageNumber="10">
<taxonomicName id="C99B4EBFFF951D15A624FD4AFC8CFD08" ID-CoL="5892Y" authority="Solis &amp; Kohlmann" authorityName="Solis &amp; Kohlmann" box="[264,891,723,750]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="arcus" status="sp. nov." subGenus="Eutrichillum">
<emphasis id="3CEFE92EFF951D15A624FD4AFD7DFD0B" bold="true" box="[264,650,724,750]" italics="true" pageId="9" pageNumber="10">Trichillum (Eutrichillum) arcus</emphasis>
Solís &amp; Kohlmann
</taxonomicName>
,
<taxonomicNameLabel id="27DC5455FF951D15A4A4FD4DFC12FD08" box="[904,997,723,749]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="96A029B9FF951D15A4D9FD4DFBCAFD08" box="[1013,1085,723,749]" captionStart="FIGURE 6" captionStartId="11.[264,368,1730,1754]" captionTargetBox="[289,1291,381,1690]" captionTargetId="figure@11.[284,1304,369,1703]" captionTargetPageId="11" captionText="FIGURE 6. Dorsal habitus of Trichillum arcus Solís &amp; Kohlmann, sp. nov. (holotype)." httpUri="https://zenodo.org/record/168694/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF951D15A624FCBDFD68FC3B" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="0E24353CFF951D15A624FCBDFD68FC3B" blockId="9.[264,1324,803,1790]" pageId="9" pageNumber="10">
<emphasis id="3CEFE92EFF951D15A624FCBDFE7FFCD8" bold="true" box="[264,392,803,829]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
This species is distinguished from other
<taxonomicName id="C99B4EBFFF951D15A453FCBAFC0EFCD8" box="[895,1017,804,829]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="3CEFE92EFF951D15A453FCBAFC0EFCD8" box="[895,1017,804,829]" italics="true" pageId="9" pageNumber="10">Trichillum</emphasis>
</taxonomicName>
species by the following combination of characters: metallic­colour, with head and pronotum a brilliant wine­red with green cupreous reflections, the elytra a darker wine­red with green cupreous reflections; clypeo­genal suture raised and keeled, arching and projecting towards the middle of the head, and interrupted medially.
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF951D10A614FC75FE21FCDB" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="description">
<paragraph id="0E24353CFF951D15A614FC75FC53FB9B" blockId="9.[264,1324,803,1790]" pageId="9" pageNumber="10">
<emphasis id="3CEFE92EFF951D15A614FC75FD50FBE0" bold="true" box="[312,679,1003,1029]" pageId="9" pageNumber="10">
Description.
<typeStatus id="D1208B9EFF951D15A6F6FC75FDB9FBE0" box="[474,590,1003,1029]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
. Male:
</emphasis>
Length
<quantity id="C96398D9FF951D15A422FC72FC99FBE3" box="[782,878,1004,1030]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="9" pageNumber="10" unit="mm" value="3.8">3.8 mm</quantity>
. Humeral width
<quantity id="C96398D9FF951D15A315FC72FB6EFBE3" box="[1081,1177,1004,1030]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="10" unit="mm" value="2.7">2.7 mm</quantity>
. Body short and rounded, dorsally convex. Dorsal surface brilliant wine­red on head and pronotum with green cupreous reflections; elytra darker wine red with green cupreous reflections; ventral surface reddish­brown. Dorsal surface with setae.
</paragraph>
<paragraph id="0E24353CFF951D15A614FB12FDEBFAA3" blockId="9.[264,1324,803,1790]" pageId="9" pageNumber="10">Head smooth, covered with umbilicate punctures that bear a golden seta; clypeus bidentate separated by a slightly rounded V­shaped notch, teeth well developed; genae projecting in an even arch; clypeo­genal suture raised and keeled, arching and projecting towards the middle of the head, but interrupted medially; eyes small and emarginated, 3.5 times longer than wide.</paragraph>
<paragraph id="0E24353CFF951D15A614FACAFBFEFA03" blockId="9.[264,1324,803,1790]" pageId="9" pageNumber="10">Pronotum smooth, covered haphazardly with umbilicate punctures of different sizes that bear a golden seta; postero­laterally with a small, black gibbosity; base not emarginated; pronotal lateral borders slightly angled anteriorly; an arched carina goes from behind and below the lateral gibbosity to the antero­lateral pronotal arch.</paragraph>
<paragraph id="0E24353CFF951D15A614FA6AFE0FF91B" blockId="9.[264,1324,803,1790]" pageId="9" pageNumber="10">Elytra with eight striae, including the epipleural. Intervals slightly convex and smooth. Striae slightly impressed, more so at their apex, faintly punctate. First stria with a row of umbilicate punctures bearing an erect golden seta along its inner side. Striae 2­8 with the same row of punctures along the external side, with the exception of the second stria, which also has on its apical third the seta bearing punctures. Pygidium completely margined, smooth and convex, covered with umbilicate punctures of varying sizes and bearing an erect golden seta.</paragraph>
<caption id="5AE465B4FF961D16A624F888FC03F8C8" box="[264,1012,1814,1838]" httpUri="https://zenodo.org/record/168693/files/figure.png" pageId="10" pageNumber="11" targetBox="[285,1306,382,1780]" targetPageId="10">
<paragraph id="0E24353CFF961D16A624F888FC03F8C8" blockId="10.[264,1012,1814,1838]" box="[264,1012,1814,1838]" pageId="10" pageNumber="11">
<emphasis id="3CEFE92EFF961D16A624F888FE7DF8CB" bold="true" box="[264,394,1814,1838]" pageId="10" pageNumber="11">FIGURE 5.</emphasis>
Dorsal habitus of
<taxonomicName id="C99B4EBFFF961D16A57CF888FC07F8C8" authority="Boucomont" authorityName="Boucomont" box="[592,1008,1814,1837]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bradyporum">
<emphasis id="3CEFE92EFF961D16A57CF888FCAFF8C8" box="[592,856,1814,1837]" italics="true" pageId="10" pageNumber="11">Pedaridium bradyporum</emphasis>
(Boucomont)
</taxonomicName>
.
</paragraph>
</caption>
<caption id="5AE465B4FF971D17A624F95CFB46F93C" box="[264,1201,1730,1754]" httpUri="https://zenodo.org/record/168694/files/figure.png" pageId="11" pageNumber="12" targetBox="[289,1291,381,1690]" targetPageId="11">
<paragraph id="0E24353CFF971D17A624F95CFB46F93C" blockId="11.[264,1201,1730,1754]" box="[264,1201,1730,1754]" pageId="11" pageNumber="12">
<emphasis id="3CEFE92EFF971D17A624F95CFE7DF93F" bold="true" box="[264,394,1730,1754]" pageId="11" pageNumber="12">FIGURE 6.</emphasis>
Dorsal habitus of
<taxonomicName id="C99B4EBFFF971D17A57CF95CFC3BF93C" authority="Solis &amp; Kohlmann" authorityName="Solis &amp; Kohlmann" box="[592,972,1730,1753]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="arcus" status="sp. nov.">
<emphasis id="3CEFE92EFF971D17A57CF95CFCF5F93C" box="[592,770,1730,1753]" italics="true" pageId="11" pageNumber="12">Trichillum arcus</emphasis>
Solís &amp; Kohlmann
</taxonomicName>
,
<emphasis id="3CEFE92EFF971D17A4F2F95CFBC2F93F" bold="true" box="[990,1077,1730,1754]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="27DC5455FF971D17A4F2F95CFBC2F93F" box="[990,1077,1730,1754]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype).
</paragraph>
</caption>
<paragraph id="0E24353CFF901D10A614FE82FC24FDC3" blockId="12.[264,1325,284,1830]" pageId="12" pageNumber="13">
Femora covered by umbilicate punctures bearing an erect golden seta. Pro­, mesoepimeron, metaepisternum, pro­, meso­, metasternum and abdominal sternites covered with large, flat and shallow ocellated impressions; in the case of the metasternum the impressions lie along its margin, whereas the central area is smooth. Meso­ and metafemora have a line of punctures bearing setae at the posterior ventral margin. Like all males from the subgenus
<taxonomicName id="C99B4EBFFF901D10A6DAFE7AFD7DFE18" box="[502,650,484,509]" class="Insecta" family="Scarabaeidae" genus="Eutrichillum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="3CEFE92EFF901D10A6DAFE7AFD7DFE18" box="[502,650,484,509]" italics="true" pageId="12" pageNumber="13">Eutrichillum</emphasis>
</taxonomicName>
it has the fifth protarsus distally swollen and dorsally deeply excavated for accommodating the claws while at rest.
</paragraph>
<paragraph id="0E24353CFF901D10A614FDADFCECFD93" blockId="12.[264,1325,284,1830]" pageId="12" pageNumber="13">
<emphasis id="3CEFE92EFF901D10A614FDADFDE0FDA8" bold="true" box="[312,535,563,589]" pageId="12" pageNumber="13">Allotype. Female:</emphasis>
Length
<quantity id="C96398D9FF901D10A552FDAAFD29FDAB" box="[638,734,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="13" unit="mm" value="3.5">3.5 mm</quantity>
. Humeral width
<quantity id="C96398D9FF901D10A484FDAAFBFFFDAB" box="[936,1032,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="13" unit="mm" value="2.2">2.2 mm</quantity>
. Similar to male, except that the fifth protarsus is not distally swollen.
</paragraph>
<paragraph id="0E24353CFF901D10A614FD1DFE21FCDB" blockId="12.[264,1325,284,1830]" pageId="12" pageNumber="13">
<emphasis id="3CEFE92EFF901D10A614FD1DFE45FD78" bold="true" box="[312,434,643,669]" pageId="12" pageNumber="13">Variation.</emphasis>
Length
<quantity id="C96398D9FF901D10A535FD1AFD50FD7B" box="[537,679,644,670]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.1" metricValueMin="2.9" pageId="12" pageNumber="13" unit="mm" value="3.5" valueMax="4.1" valueMin="2.9">2.9­4.1 mm</quantity>
. Humeral width
<quantity id="C96398D9FF901D10A443FD1AFC0BFD7B" box="[879,1020,644,670]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.8" metricValueMin="1.8" pageId="12" pageNumber="13" unit="mm" value="2.3" valueMax="2.8" valueMin="1.8">1.8­2.8 mm</quantity>
. The first stria has sometimes one or two isolated umbilical punctures with an erect seta along its external margin. The second stria has a row of umbilical punctures with erect setae along its inner margin going from its apical third to its apical half. Strial punctures can become more crenulating towards the apex.
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF901D10A614FCD5FCA6FA73" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="0E24353CFF901D10A614FCD5FD36FAA3" blockId="12.[264,1325,284,1830]" pageId="12" pageNumber="13">
<emphasis id="3CEFE92EFF901D10A614FCD5FDDCFC80" bold="true" box="[312,555,843,869]" pageId="12" pageNumber="13">Examined material</emphasis>
(
<specimenCount id="189DFEB5FF901D10A510FCD2FD13FC83" box="[572,740,844,870]" pageId="12" pageNumber="13" type="generic">39 specimens</specimenCount>
).
<typeStatus id="D1208B9EFF901D10A5D5FCD5FC9CFC80" box="[761,875,843,869]" pageId="12" pageNumber="13" type="holotype">
<emphasis id="3CEFE92EFF901D10A5D5FCD5FC9CFC80" bold="true" box="[761,875,843,869]" pageId="12" pageNumber="13">Holotype</emphasis>
</typeStatus>
, male:
<collectingCountry id="768C75ACFF901D10A4EBFCD2FB8EFC83" box="[967,1145,844,870]" name="Costa Rica" pageId="12" pageNumber="13">COSTA RICA</collectingCountry>
. Est. Pitilla,
<quantity id="C96398D9FF901D10A231FCD2FED9FC6B" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="13" unit="km" value="9.0">9 km</quantity>
S. Sta. Cecilia, P.N. Guanacaste, Prov. Guanacaste,
<quantity id="C96398D9FF901D10A494FCEAFC09FC6B" box="[952,1022,884,910]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="m" value="700.0">700m</quantity>
, May, 1994, C. Moraga,
<collectionCode id="688AADF9FF901D10A624FC02FECDFC53" LSID="urn:lsid:biocol.org:col:14323" box="[264,314,924,950]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="12" pageNumber="13">CRI</collectionCode>
001 832764.
<emphasis id="3CEFE92EFF901D10A6F4FC05FDB5FC50" bold="true" box="[472,578,923,949]" pageId="12" pageNumber="13">Allotype</emphasis>
, female: Prov. Guanacaste, Est. Pitilla,
<quantity id="C96398D9FF901D10A30FFC02FB95FC53" box="[1059,1122,924,950]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="13" unit="km" value="9.0">9 km</quantity>
S Santa Cecilia,
<quantity id="C96398D9FF901D10A624FC5AFEB9FC3B" box="[264,334,964,990]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="m" value="700.0">700m</quantity>
,
<date id="7A2513FCFF901D10A670FC5AFE22FC3B" box="[348,469,964,990]" pageId="12" pageNumber="13" value="1996-10">Oct. 1996</date>
, C. Moraga,
<collectionCode id="688AADF9FF901D10A55AFC5AFD50FC3B" LSID="urn:lsid:biocol.org:col:14323" box="[630,679,964,990]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14323" name="Universidade do Extremo Sul Catarinense, Bairro Universitário" pageId="12" pageNumber="13">CRI</collectionCode>
002 500213.
<emphasis id="3CEFE92EFF901D10A469FC5DFC3DFC38" bold="true" box="[837,970,963,989]" pageId="12" pageNumber="13">
<typeStatus id="D1208B9EFF901D10A469FC5DFC32FC38" box="[837,965,963,989]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Guanacaste:
<emphasis id="3CEFE92EFF901D10A341FC5AFB6DFC38" box="[1133,1178,964,989]" italics="true" pageId="12" pageNumber="13">ibid</emphasis>
. allotype,
<specimenCount id="189DFEB5FF901D10A230FC5AFE71FBE3" pageId="12" pageNumber="13" type="generic">5 specimens</specimenCount>
;
<emphasis id="3CEFE92EFF901D10A6BEFC72FE48FBE0" box="[402,447,1004,1029]" italics="true" pageId="12" pageNumber="13">ibid</emphasis>
.
<typeStatus id="D1208B9EFF901D10A6E2FC72FDCFFBE3" box="[462,568,1004,1030]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
,
<specimenCount id="189DFEB5FF901D10A56FFC72FD3CFBE3" box="[579,715,1004,1030]" pageId="12" pageNumber="13" type="generic">1 specimen</specimenCount>
;
<emphasis id="3CEFE92EFF901D10A5FBFC72FCF3FBE0" box="[727,772,1004,1029]" italics="true" pageId="12" pageNumber="13">ibid</emphasis>
.,
<date id="7A2513FCFF901D10A436FC72FC4EFBE3" box="[794,953,1004,1030]" pageId="12" pageNumber="13" value="1994-06-03">
Junio 1994,
<specimenCount id="189DFEB5FF901D10A486FC72FC4EFBE3" box="[938,953,1004,1030]" pageId="12" pageNumber="13" type="generic">3</specimenCount>
</date>
specimens;
<emphasis id="3CEFE92EFF901D10A365FC72FB81FBE0" box="[1097,1142,1004,1029]" italics="true" pageId="12" pageNumber="13">ibid</emphasis>
.,
<date id="7A2513FCFF901D10A3A0FC72FAD1FBE3" box="[1164,1318,1004,1030]" pageId="12" pageNumber="13" value="1993-08">Agosto 1993</date>
; Sector Santa María,
<quantity id="C96398D9FF901D10A6D3FB8AFDBBFBCB" box="[511,588,1044,1070]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="13" unit="km" value="25.0">25 km</quantity>
NE de
<collectingCountry id="768C75ACFF901D10A580FB8AFCF4FBCB" box="[684,771,1044,1070]" name="Liberia" pageId="12" pageNumber="13">Liberia</collectingCountry>
,
<quantity id="C96398D9FF901D10A43EFB8AFC96FBCB" box="[786,865,1044,1070]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="12" pageNumber="13" unit="m" value="790.0">790 m</quantity>
,
<date id="7A2513FCFF901D10A443FB8AFC2BFBCB" box="[879,988,1044,1070]" pageId="12" pageNumber="13" value="1927-10-09">9­27 Oct</date>
, 1996, D. Briceño,
<specimenCount id="189DFEB5FF901D10A3E7FB8AFEBEFBB3" pageId="12" pageNumber="13" type="generic">6 specimens</specimenCount>
;
<date id="7A2513FCFF901D10A672FBA2FE2AFBB3" box="[350,477,1084,1110]" pageId="12" pageNumber="13" value="1911-11-18">11­18 Nov</date>
, 1996,
<specimenCount id="189DFEB5FF901D10A51FFBA2FD30FBB3" box="[563,711,1084,1110]" pageId="12" pageNumber="13" type="generic">2 specimens</specimenCount>
. Alajuela: Sector San Ramón de Dos Ríos, 1.5 Km NO Hda.
<collectingCountry id="768C75ACFF901D10A654FBFAFDC2FB9B" box="[376,565,1124,1150]" name="New Zealand" pageId="12" pageNumber="13">Nueva Zelandia</collectingCountry>
,
<quantity id="C96398D9FF901D10A56CFBFAFD71FB9B" box="[576,646,1124,1150]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="12" pageNumber="13" unit="m" value="620.0">620m</quantity>
,
<date id="7A2513FCFF901D10A5BDFBFAFCAEFB9B" box="[657,857,1124,1150]" pageId="12" pageNumber="13" value="1996-07-12" valueMax="1996-07-21" valueMin="1996-07-12">12­21 Julio 1996</date>
, F.A. Quesada,
<specimenCount id="189DFEB5FF901D10A33DFBFAFB44FB9B" box="[1041,1203,1124,1150]" pageId="12" pageNumber="13" type="generic">14 specimens</specimenCount>
;
<date id="7A2513FCFF901D10A392FBFAFEB2FB43" pageId="12" pageNumber="13" value="1995-09-17" valueMax="1995-09-28" valueMin="1995-09-17">17­28 set 1995</date>
,
<specimenCount id="189DFEB5FF901D10A67EFB12FE2BFB43" box="[338,476,1164,1190]" pageId="12" pageNumber="13" type="generic">1 specimen</specimenCount>
; Puesto Quebradón,
<quantity id="C96398D9FF901D10A5FEFB12FCD7FB43" box="[722,800,1164,1190]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="13" unit="m" value="300.0">300 m</quantity>
,
<date id="7A2513FCFF901D10A401FB12FC52FB43" box="[813,933,1164,1190]" pageId="12" pageNumber="13" value="1997-11">Nov 1997</date>
, G. Rodríguez, trampas de intercepción, con agua de sal, Baykil y frutas en las tazas,
<specimenCount id="189DFEB5FF901D10A4A7FB2AFBE2FB2B" box="[907,1045,1204,1230]" pageId="12" pageNumber="13" type="generic">1 specimen</specimenCount>
; San Lorenzo,
<quantity id="C96398D9FF901D10A3EBFB2AFED5FB13" metricMagnitude="2" metricUnit="m" metricValue="6.1" metricValueMax="6.2" metricValueMin="6.0" pageId="12" pageNumber="13" unit="m" value="610.0" valueMax="620.0" valueMin="600.0">600­620 m</quantity>
,
<date id="7A2513FCFF901D10A603FB42FDBFFB13" box="[303,584,1244,1270]" pageId="12" pageNumber="13" value="1996-09-16" valueMax="1996-09-19" valueMin="1996-09-16">16­19 Septiembre 1996</date>
, F.A. Quesada,
<specimenCount id="189DFEB5FF901D10A429FB42FC78FB13" box="[773,911,1244,1270]" pageId="12" pageNumber="13" type="generic">1 specimen</specimenCount>
. Heredia: Est. Biológica La Selva,
<date id="7A2513FCFF901D10A624FA9AFE62FAFB" box="[264,405,1284,1310]" pageId="12" pageNumber="13" value="1998-06-21">21­VI­1998</date>
, C. Carleton &amp; A. Tishechkin,
<specimenCount id="189DFEB5FF901D10A428FA9AFC79FAFB" box="[772,910,1284,1310]" pageId="12" pageNumber="13" type="generic">1 specimen</specimenCount>
;
<quantity id="C96398D9FF901D10A4B6FA9AFBF9FAFB" box="[922,1038,1284,1310]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="12" pageNumber="13" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50­150 m</quantity>
,
<date id="7A2513FCFF901D10A337FA9AFB3CFAFB" box="[1051,1227,1284,1310]" pageId="12" pageNumber="13" value="1993-05-14">14­Mayo 1993</date>
, INBio­ OET, bosque secundario,
<specimenCount id="189DFEB5FF901D10A519FAB2FD49FAA3" box="[565,702,1324,1350]" pageId="12" pageNumber="13" type="generic">1 specimen</specimenCount>
.
</paragraph>
<paragraph id="0E24353CFF901D10A614FACDFCA6FA73" blockId="12.[264,1325,284,1830]" pageId="12" pageNumber="13">
<emphasis id="3CEFE92EFF901D10A614FACDFE68FA88" bold="true" box="[312,415,1363,1389]" pageId="12" pageNumber="13">Habitat.</emphasis>
This species has been collected using traps baited with rotting meat or with flight interception traps inside tropical rain forest.
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF901D10A614FA3DFBE5F8C3" pageId="12" pageNumber="13" type="description">
<paragraph id="0E24353CFF901D10A614FA3DFE91F9EB" blockId="12.[264,1325,284,1830]" pageId="12" pageNumber="13">
<emphasis id="3CEFE92EFF901D10A614FA3DFD72FA58" bold="true" box="[312,645,1443,1469]" pageId="12" pageNumber="13">Geographical distribution.</emphasis>
This new species of
<taxonomicName id="C99B4EBFFF901D10A451FA3AFC00FA58" box="[893,1015,1444,1469]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="3CEFE92EFF901D10A451FA3AFC00FA58" box="[893,1015,1444,1469]" italics="true" pageId="12" pageNumber="13">Trichillum</emphasis>
</taxonomicName>
occurs on both the Atlantic and Pacific slopes, along the bases of the volcanoes of the Guanacaste and Tilarán Cordilleras.
</paragraph>
<paragraph id="0E24353CFF901D10A614F985FD6FF963" blockId="12.[264,1325,284,1830]" pageId="12" pageNumber="13">
<emphasis id="3CEFE92EFF901D10A614F985FDA3F9D0" bold="true" box="[312,596,1563,1589]" pageId="12" pageNumber="13">Chorological affinities.</emphasis>
So far this is the northernmost distribution range known for a member of the genus
<taxonomicName id="C99B4EBFFF901D10A53AF9DAFD67F9B8" box="[534,656,1604,1629]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="3CEFE92EFF901D10A53AF9DAFD67F9B8" box="[534,656,1604,1629]" italics="true" pageId="12" pageNumber="13">Trichillum</emphasis>
</taxonomicName>
. This species will most probably be also found in the Atlantic rain forests of
<collectingCountry id="768C75ACFF901D10A534F9F2FD63F963" box="[536,660,1644,1670]" name="Nicaragua" pageId="12" pageNumber="13">Nicaragua</collectingCountry>
.
</paragraph>
<paragraph id="0E24353CFF901D10A614F90DFBE5F8C3" blockId="12.[264,1325,284,1830]" pageId="12" pageNumber="13">
<emphasis id="3CEFE92EFF901D10A614F90DFD77F948" bold="true" box="[312,640,1683,1709]" pageId="12" pageNumber="13">Taxonomic Relationships.</emphasis>
The species belongs to the subgenus
<taxonomicName id="C99B4EBFFF901D10A344F90AFB0BF948" box="[1128,1276,1684,1709]" class="Insecta" family="Scarabaeidae" genus="Eutrichillum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="3CEFE92EFF901D10A344F90AFB0BF948" box="[1128,1276,1684,1709]" italics="true" pageId="12" pageNumber="13">Eutrichillum</emphasis>
</taxonomicName>
, as defined by
<bibRefCitation id="6A0A48CDFF901D10A6A0F922FDB9F933" author="Martinez" box="[396,590,1724,1750]" pageId="12" pageNumber="14" refString="Martinez, A. (1967) Notas para una monografia del genero Trichillum Harold, 1868. Revista de la Sociedad Mexicana de Historia Natural, 28, 119 - 147." type="journal article" year="1967">Martínez (1967)</bibRefCitation>
. Based on external characteristics, the clypeo­genal suture, the punctation of the pronotum and the piliferous punctation and flatness of the elytral intervals, this new species seems to be related to
<taxonomicName id="C99B4EBFFF901D10A43DF892FBE5F8C3" authority="Arrow." authorityName="Arrow." box="[785,1042,1804,1830]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="boucomonti">
<emphasis id="3CEFE92EFF901D10A43DF892FC4FF8C0" box="[785,952,1804,1829]" italics="true" pageId="12" pageNumber="13">T. boucomonti</emphasis>
Arrow.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="468166B7FF911D11A614FE85FE00FEBB" pageId="13" pageNumber="14" type="etymology">
<paragraph id="0E24353CFF911D11A614FE85FE00FEBB" blockId="13.[264,1321,283,350]" pageId="13" pageNumber="14">
<emphasis id="3CEFE92EFF911D11A614FE85FE32FED0" bold="true" box="[312,453,283,309]" pageId="13" pageNumber="14">Etymology.</emphasis>
From the Latin word meaning rainbow, a reference to the iridescent coloration of this species.
</paragraph>
</subSubSection>
</treatment>
</document>