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<document id="AC056A2F84924AD61C6CA4EFF8692703" ID-CLB-Dataset="46900" ID-DOI="10.11646/zootaxa.4067.4.1" ID-GBIF-Dataset="4d2bf7a6-415a-4fb2-8c8f-12b79f8a400a" ID-ISSN="1175-5326" ID-Zenodo-Dep="257945" ID-ZooBank="1CC5D0E7-0B60-4E62-BACD-9775931ED7F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460354321046" checkinUser="plazi" docAuthor="Souto, Javier, Berning, Björn &amp; Ostrovsky, Andrew N." docDate="2016" docId="862A8F25FF900C58FF65FB3D5BD63D8C" docLanguage="en" docName="zootaxa.4067.4.1.pdf" docOrigin="Zootaxa 4067 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Escharella lopezfei Souto, Berning &amp; Ostrovsky, 2016, n. sp." docType="treatment" docUuid="5DD405FF-C31D-4BAC-81F9-74BD1676D7B5" docUuidSource="ZooBank" docVersion="11" lastPageNumber="434" masterDocId="7A13F75DFF8F0C79FFF2FF985F173879" masterDocTitle="Systematics and diversity of deep-water Cheilostomata (Bryozoa) from Galicia Bank (NE Atlantic)" masterLastPageNumber="459" masterPageNumber="401" pageNumber="432" updateTime="1698293824137" updateUser="ExternalLinkService">
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<mods:title id="9C4FEFF87065D7B610600D288A0211B6">Systematics and diversity of deep-water Cheilostomata (Bryozoa) from Galicia Bank (NE Atlantic)</mods:title>
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<mods:namePart id="91B40FD5EDB16A85B5D5BFCA7A3B546A">Souto, Javier</mods:namePart>
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<mods:namePart id="EA230EEC2F8F18447CEF220605C8A662">Berning, Björn</mods:namePart>
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<mods:namePart id="FB90C888A235548373D81EA4F5243CE4">Ostrovsky, Andrew N.</mods:namePart>
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<treatment id="862A8F25FF900C58FF65FB3D5BD63D8C" ID-DOI="http://doi.org/10.5281/zenodo.4547345" ID-GBIF-Taxon="119408643" ID-Zenodo-Dep="4547345" LSID="urn:lsid:zoobank.org:act:5DD405FF-C31D-4BAC-81F9-74BD1676D7B5" httpUri="http://treatment.plazi.org/id/862A8F25FF900C58FF65FB3D5BD63D8C" lastPageId="33" lastPageNumber="434" pageId="31" pageNumber="432">
<subSubSection id="46996DB8FF900C66FF65FB3D5E813C99" pageId="31" pageNumber="432" type="nomenclature">
<paragraph id="0E3C3E33FF900C66FF65FB3D5ED03CC6" blockId="31.[151,455,1189,1248]" box="[151,455,1189,1215]" pageId="31" pageNumber="432">
<heading id="5574895FFF900C66FF65FB3D5ED03CC6" bold="true" box="[151,455,1189,1215]" fontSize="11" level="1" pageId="31" pageNumber="432" reason="1">
<emphasis id="3CF7E221FF900C66FF65FB3D5ED03CC6" bold="true" box="[151,455,1189,1215]" pageId="31" pageNumber="432">
<taxonomicName id="C98345B0FF900C66FF65FB3D5E6A3CC6" box="[151,381,1189,1215]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="31" pageNumber="432" phylum="Bryozoa" rank="species" species="lopezfei" status="sp. nov.">
<emphasis id="3CF7E221FF900C66FF65FB3D5E6A3CC6" bold="true" box="[151,381,1189,1215]" italics="true" pageId="31" pageNumber="432">Escharella lopezfei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27C45F5AFF900C66FE76FB3D5ED03CC6" box="[388,455,1189,1215]" pageId="31" pageNumber="432" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="0E3C3E33FF900C66FF65FB5F5E813C99" blockId="31.[151,455,1189,1248]" box="[151,406,1223,1248]" pageId="31" pageNumber="432">
(
<figureCitation id="96B822B6FF900C66FF6DFB5F5E373C99" box="[159,288,1223,1248]" captionStart="FIGURES 76 83" captionStartId="32.[151,264,1846,1868]" captionTargetBox="[150,1436,193,1824]" captionTargetId="figure@32.[150,1436,193,1825]" captionTargetPageId="32" captionText="FIGURES 76 83. Escharella lopezfei n. sp.: 76, general view of colony (SMF 40.014); 77, ancestrula and periancestrular zooids (SMF 40.015); 78, 79, zooidal orifices (SMF 40.015); 80, lateral view of autozooid with basal pore-chambers; 81, 82, autozooids with ovicells (holotype); 83, ancestrula." httpUri="https://zenodo.org/record/257961/files/figure.png" pageId="31" pageNumber="432">Figs 7683</figureCitation>
,
<tableCitation id="43010B88FF900C66FED9FB5F5E9A3C99" box="[299,397,1223,1248]" captionStart="TABLE 14" captionStartId="33.[151,239,152,175]" captionTargetBox="[159,1302,202,646]" captionTargetPageId="33" captionText="TABLE 14. Measurements (in mm) of Escharella lopezfei n. sp." httpUri="http://table.plazi.org/id/5AFC6EBBFFAE0C58FF65FF005C5E38D6" pageId="31" pageNumber="432" tableUuid="5AFC6EBBFFAE0C58FF65FF005C5E38D6">Table 14</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="46996DB8FF900C66FF65FA975DF13D34" pageId="31" pageNumber="432" type="materials_examined">
<paragraph id="0E3C3E33FF900C66FF65FA975DF13D34" blockId="31.[151,1436,1295,1968]" pageId="31" pageNumber="432">
<emphasis id="3CF7E221FF900C66FF65FA975E9F3D51" bold="true" box="[151,392,1295,1320]" pageId="31" pageNumber="432">Material examined.</emphasis>
<typeStatus id="D1388091FF900C66FE64FA975EEA3D51" box="[406,509,1295,1320]" pageId="31" pageNumber="432" type="holotype">
<emphasis id="3CF7E221FF900C66FE64FA975EEA3D51" box="[406,509,1295,1320]" italics="true" pageId="31" pageNumber="432">Holotype</emphasis>
</typeStatus>
:
<collectionCode id="6892A6F6FF900C66FDE1FA885D5C3D51" LSID="urn:lsid:biocol.org:col:34838" box="[531,587,1296,1320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="31" pageNumber="432">SMF</collectionCode>
40.013, locality VH-97-315.
<typeStatus id="D1388091FF900C66FC47FA895B313D51" box="[949,1062,1297,1320]" pageId="31" pageNumber="432" type="paratype">
<emphasis id="3CF7E221FF900C66FC47FA895B313D51" box="[949,1062,1297,1320]" italics="true" pageId="31" pageNumber="432">Paratypes</emphasis>
</typeStatus>
:
<collectionCode id="6892A6F6FF900C66FBCEFA885B633D51" LSID="urn:lsid:biocol.org:col:34838" box="[1084,1140,1296,1320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="31" pageNumber="432">SMF</collectionCode>
40.014, 40.015, 40.016, 40.017, 40.018, 40.019, 40.020, locality VH-97-315.
</paragraph>
</subSubSection>
<subSubSection id="46996DB8FF900C66FF35FACF5E323DEC" pageId="31" pageNumber="432" type="etymology">
<paragraph id="0E3C3E33FF900C66FF35FACF5E323DEC" blockId="31.[151,1436,1295,1968]" pageId="31" pageNumber="432">
<emphasis id="3CF7E221FF900C66FF35FACF5E593D09" bold="true" box="[199,334,1367,1392]" pageId="31" pageNumber="432">Etymology.</emphasis>
Honorific for Dr Carlos López-Fé for his contributions to knowledge of bryozoan diversity in the NE Atlantic.
</paragraph>
</subSubSection>
<subSubSection id="46996DB8FF900C58FF35FA075C483BE8" lastPageId="33" lastPageNumber="434" pageId="31" pageNumber="432" type="description">
<paragraph id="0E3C3E33FF900C66FF35FA075E153ECC" blockId="31.[151,1436,1295,1968]" pageId="31" pageNumber="432">
<emphasis id="3CF7E221FF900C66FF35FA075E4E3DC1" bold="true" box="[199,345,1439,1464]" pageId="31" pageNumber="432">Description.</emphasis>
Colony encrusting, unilaminar, forming an irregular patch. Autozooids oval to hexagonal, separated by well-marked grooves; frontal shield strongly, evenly convex, with finely granular surface and single row of tiny, widely spaced marginal pores; additional pores seen in some zooids near orifice. Lateral walls much reduced, with numerous elongated basal pore-chambers, 25 combined into elongated areas framed by gymnocystal calcification, 1 distal, 2 distolateral and 2 proximolateral; basal wall only marginally calcified. Orifice suboval, longer than wide; proximal border with rounded triangular lyrula; condyles thick, short and blunt; distolateral margin with 89 long, articulated, evenly spaced oral spines, the proximal-most pair level with condyles.
</paragraph>
<paragraph id="0E3C3E33FF900C66FF35F9275EAC3FF4" blockId="31.[151,1436,1295,1968]" pageId="31" pageNumber="432">Ovicells hyperstomial, not closed by operculum (acleithral). Ooecium globular, generally wider than long, formed by and continuous with frontal shield of distal autozooid. Ectooecium membranous except for a narrow band of smooth gymnocystal calcification on proximomedian margin. Calcified endooecium thick, imperforate, surface finely granular like zooidal frontal shield. Ovicelled zooids slightly dimorphic, differing from autozooids by presence of gap between distalmost pair of oral spines that laterally border ovicell opening, and by closer spacing of the oral spines.</paragraph>
<paragraph id="0E3C3E33FF900C66FF35F80F5CA13FC9" blockId="31.[151,1436,1295,1968]" box="[199,950,1943,1968]" pageId="31" pageNumber="432">Kenozooids rare, as large as autozooids, lacking orifice and spines.</paragraph>
<caption id="5AFC6EBBFFAF0C59FF65F8AE5D893FF3" httpUri="https://zenodo.org/record/257961/files/figure.png" pageId="32" pageNumber="433" targetBox="[150,1436,193,1824]" targetPageId="32">
<paragraph id="0E3C3E33FFAF0C59FF65F8AE5D893FF3" blockId="32.[151,1436,1846,1930]" pageId="32" pageNumber="433">
<emphasis id="3CF7E221FFAF0C59FF65F8AE5E403F35" bold="true" box="[151,343,1846,1868]" pageId="32" pageNumber="433">FIGURES 7683.</emphasis>
<taxonomicName id="C98345B0FFAF0C59FEADF8AE5D333F35" box="[351,548,1846,1868]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="32" pageNumber="433" phylum="Bryozoa" rank="species" species="lopezfei" status="sp. nov.">
<emphasis id="3CF7E221FFAF0C59FEADF8AE5D333F35" box="[351,548,1846,1868]" italics="true" pageId="32" pageNumber="433">Escharella lopezfei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27C45F5AFFAF0C59FDDFF8AE5D7E3F35" box="[557,617,1846,1868]" pageId="32" pageNumber="433" rank="species">
<emphasis id="3CF7E221FFAF0C59FDDFF8AE5D7E3F35" bold="true" box="[557,617,1846,1868]" pageId="32" pageNumber="433">n. sp.</emphasis>
</taxonomicNameLabel>
:
<emphasis id="3CF7E221FFAF0C59FD8BF8AE5D8E3F35" bold="true" box="[633,665,1846,1868]" pageId="32" pageNumber="433">76,</emphasis>
general view of colony (SMF 40.014);
<emphasis id="3CF7E221FFAF0C59FBCDF8AE5B483F35" bold="true" box="[1087,1119,1846,1868]" pageId="32" pageNumber="433">77,</emphasis>
ancestrula and periancestrular zooids (SMF 40.015);
<emphasis id="3CF7E221FFAF0C59FE8DF8CD5ED13F12" bold="true" box="[383,454,1877,1899]" pageId="32" pageNumber="433">78, 79,</emphasis>
zooidal orifices (SMF 40.015);
<emphasis id="3CF7E221FFAF0C59FCE0F8CD5C263F12" bold="true" box="[786,817,1877,1899]" pageId="32" pageNumber="433">80,</emphasis>
lateral view of autozooid with basal pore-chambers;
<emphasis id="3CF7E221FFAF0C59FAA7F8CD5A8B3F12" bold="true" box="[1365,1436,1877,1899]" pageId="32" pageNumber="433">81, 82,</emphasis>
autozooids with ovicells (holotype);
<emphasis id="3CF7E221FFAF0C59FDF8F8EC5D3E3FF3" bold="true" box="[522,553,1908,1930]" pageId="32" pageNumber="433">83,</emphasis>
ancestrula.
</paragraph>
</caption>
<caption id="5AFC6EBBFFAE0C58FF65FF005C5E38D6" ID-Table-UUID="5AFC6EBBFFAE0C58FF65FF005C5E38D6" box="[151,841,152,175]" httpUri="http://table.plazi.org/id/5AFC6EBBFFAE0C58FF65FF005C5E38D6" pageId="33" pageNumber="434" targetBox="[159,1302,202,646]" targetIsTable="true" targetPageId="33">
<paragraph id="0E3C3E33FFAE0C58FF65FF005C5E38D6" blockId="33.[151,841,152,175]" box="[151,841,152,175]" pageId="33" pageNumber="434">
<emphasis id="3CF7E221FFAE0C58FF65FF005E0038D6" bold="true" box="[151,279,152,175]" pageId="33" pageNumber="434">TABLE 14.</emphasis>
Measurements (in mm) of
<taxonomicName id="C98345B0FFAE0C58FDC8FF005C1138D6" box="[570,774,152,175]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="species" species="lopezfei" status="sp. nov.">
<emphasis id="3CF7E221FFAE0C58FDC8FF005C1138D6" box="[570,774,152,175]" italics="true" pageId="33" pageNumber="434">Escharella lopezfei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27C45F5AFFAE0C58FCFFFF005C5E38D6" box="[781,841,152,175]" pageId="33" pageNumber="434" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="0E3C3E33FFAE0C58FDA0FF535A1E3AFF" pageId="33" pageNumber="434">
<table id="7C83CC93FFAEF386FF6DFF525A013AFF" box="[159,1302,202,646]" gridcols="6" gridrows="11" pageId="33" pageNumber="434">
<tr id="B0B33C71FFAEF386FF6DFF525A013899" box="[159,1302,202,224]" gridrow="0" pageId="33" pageNumber="434" rowspan-0="1">
<th id="F362550DFFAEF386FDA0FF525D9B3899" box="[594,652,202,224]" gridcol="1" gridrow="0" pageId="33" pageNumber="434">Mean</th>
<th id="F362550DFFAEF386FD0EFF525C543899" box="[764,835,202,224]" gridcol="2" gridrow="0" pageId="33" pageNumber="434">SD</th>
<th id="F362550DFFAEF386FC55FF525B1B3899" box="[935,1036,202,224]" gridcol="3" gridrow="0" pageId="33" pageNumber="434">Minimum</th>
<th id="F362550DFFAEF386FBA0FF525BAC3899" box="[1106,1211,202,224]" gridcol="4" gridrow="0" pageId="33" pageNumber="434">Maximum</th>
<th id="F362550DFFAEF386FB0EFF525A013899" box="[1276,1302,202,224]" gridcol="5" gridrow="0" pageId="33" pageNumber="434">N</th>
</tr>
<tr id="B0B33C71FFAEF386FF6DFF6E5A013975" box="[159,1302,246,268]" gridrow="1" pageId="33" pageNumber="434">
<th id="F362550DFFAEF386FF6DFF6E5EB63975" box="[159,417,246,268]" gridcol="0" gridrow="1" pageId="33" pageNumber="434">Autozooid length</th>
<td id="F362550DFFAEF386FDA0FF6E5D9B3975" box="[594,652,246,268]" gridcol="1" gridrow="1" pageId="33" pageNumber="434">0.788</td>
<td id="F362550DFFAEF386FD0EFF6E5C543975" box="[764,835,246,268]" gridcol="2" gridrow="1" pageId="33" pageNumber="434">0.0533</td>
<td id="F362550DFFAEF386FC55FF6E5B1B3975" box="[935,1036,246,268]" gridcol="3" gridrow="1" pageId="33" pageNumber="434">0.717</td>
<td id="F362550DFFAEF386FBA0FF6E5BAC3975" box="[1106,1211,246,268]" gridcol="4" gridrow="1" pageId="33" pageNumber="434">0.910</td>
<td id="F362550DFFAEF386FB0EFF6E5A013975" box="[1276,1302,246,268]" gridcol="5" gridrow="1" pageId="33" pageNumber="434">18</td>
</tr>
<tr id="B0B33C71FFAEF386FF6DFEB85A01394F" box="[159,1302,288,310]" gridrow="2" pageId="33" pageNumber="434">
<th id="F362550DFFAEF386FF6DFEB85EB6394F" box="[159,417,288,310]" gridcol="0" gridrow="2" pageId="33" pageNumber="434">Autozooid width</th>
<td id="F362550DFFAEF386FDA0FEB85D9B394F" box="[594,652,288,310]" gridcol="1" gridrow="2" pageId="33" pageNumber="434">0.610</td>
<td id="F362550DFFAEF386FD0EFEB85C54394F" box="[764,835,288,310]" gridcol="2" gridrow="2" pageId="33" pageNumber="434">0.0778</td>
<td id="F362550DFFAEF386FC55FEB85B1B394F" box="[935,1036,288,310]" gridcol="3" gridrow="2" pageId="33" pageNumber="434">0.485</td>
<td id="F362550DFFAEF386FBA0FEB85BAC394F" box="[1106,1211,288,310]" gridcol="4" gridrow="2" pageId="33" pageNumber="434">0.781</td>
<td id="F362550DFFAEF386FB0EFEB85A01394F" box="[1276,1302,288,310]" gridcol="5" gridrow="2" pageId="33" pageNumber="434">18</td>
</tr>
<tr id="B0B33C71FFAEF386FF6DFED25A013919" box="[159,1302,330,352]" gridrow="3" pageId="33" pageNumber="434">
<th id="F362550DFFAEF386FF6DFED25EB63919" box="[159,417,330,352]" gridcol="0" gridrow="3" pageId="33" pageNumber="434">Orifice length</th>
<td id="F362550DFFAEF386FDA0FED25D9B3919" box="[594,652,330,352]" gridcol="1" gridrow="3" pageId="33" pageNumber="434">0.162</td>
<td id="F362550DFFAEF386FD0EFED25C543919" box="[764,835,330,352]" gridcol="2" gridrow="3" pageId="33" pageNumber="434">0.0107</td>
<td id="F362550DFFAEF386FC55FED25B1B3919" box="[935,1036,330,352]" gridcol="3" gridrow="3" pageId="33" pageNumber="434">0.142</td>
<td id="F362550DFFAEF386FBA0FED25BAC3919" box="[1106,1211,330,352]" gridcol="4" gridrow="3" pageId="33" pageNumber="434">0.183</td>
<td id="F362550DFFAEF386FB0EFED25A013919" box="[1276,1302,330,352]" gridcol="5" gridrow="3" pageId="33" pageNumber="434">16</td>
</tr>
<tr id="B0B33C71FFAEF386FF6DFEEC5A0139F3" box="[159,1302,372,394]" gridrow="4" pageId="33" pageNumber="434">
<th id="F362550DFFAEF386FF6DFEEC5EB639F3" box="[159,417,372,394]" gridcol="0" gridrow="4" pageId="33" pageNumber="434">Orifice width</th>
<td id="F362550DFFAEF386FDA0FEEC5D9B39F3" box="[594,652,372,394]" gridcol="1" gridrow="4" pageId="33" pageNumber="434">0.143</td>
<td id="F362550DFFAEF386FD0EFEEC5C5439F3" box="[764,835,372,394]" gridcol="2" gridrow="4" pageId="33" pageNumber="434">0.0087</td>
<td id="F362550DFFAEF386FC55FEEC5B1B39F3" box="[935,1036,372,394]" gridcol="3" gridrow="4" pageId="33" pageNumber="434">0.118</td>
<td id="F362550DFFAEF386FBA0FEEC5BAC39F3" box="[1106,1211,372,394]" gridcol="4" gridrow="4" pageId="33" pageNumber="434">0.153</td>
<td id="F362550DFFAEF386FB0EFEEC5A0139F3" box="[1276,1302,372,394]" gridcol="5" gridrow="4" pageId="33" pageNumber="434">16</td>
</tr>
<tr id="B0B33C71FFAEF386FF6DFE065A0139CD" box="[159,1302,414,436]" gridrow="5" pageId="33" pageNumber="434">
<th id="F362550DFFAEF386FF6DFE065EB639CD" box="[159,417,414,436]" gridcol="0" gridrow="5" pageId="33" pageNumber="434">Ooecium length</th>
<td id="F362550DFFAEF386FDA0FE065D9B39CD" box="[594,652,414,436]" gridcol="1" gridrow="5" pageId="33" pageNumber="434">0.244</td>
<td id="F362550DFFAEF386FD0EFE065C5439CD" box="[764,835,414,436]" gridcol="2" gridrow="5" pageId="33" pageNumber="434">0.0025</td>
<td id="F362550DFFAEF386FC55FE065B1B39CD" box="[935,1036,414,436]" gridcol="3" gridrow="5" pageId="33" pageNumber="434">0.241</td>
<td id="F362550DFFAEF386FBA0FE065BAC39CD" box="[1106,1211,414,436]" gridcol="4" gridrow="5" pageId="33" pageNumber="434">0.246</td>
<td id="F362550DFFAEF386FB0EFE065A0139CD" box="[1276,1302,414,436]" gridcol="5" gridrow="5" pageId="33" pageNumber="434">3</td>
</tr>
<tr id="B0B33C71FFAEF386FF6DFE505A0139A7" box="[159,1302,456,478]" gridrow="6" pageId="33" pageNumber="434">
<th id="F362550DFFAEF386FF6DFE505EB639A7" box="[159,417,456,478]" gridcol="0" gridrow="6" pageId="33" pageNumber="434">Ooecium width</th>
<td id="F362550DFFAEF386FDA0FE505D9B39A7" box="[594,652,456,478]" gridcol="1" gridrow="6" pageId="33" pageNumber="434">0.374</td>
<td id="F362550DFFAEF386FD0EFE505C5439A7" box="[764,835,456,478]" gridcol="2" gridrow="6" pageId="33" pageNumber="434">0.0201</td>
<td id="F362550DFFAEF386FC55FE505B1B39A7" box="[935,1036,456,478]" gridcol="3" gridrow="6" pageId="33" pageNumber="434">0.360</td>
<td id="F362550DFFAEF386FBA0FE505BAC39A7" box="[1106,1211,456,478]" gridcol="4" gridrow="6" pageId="33" pageNumber="434">0.397</td>
<td id="F362550DFFAEF386FB0EFE505A0139A7" box="[1276,1302,456,478]" gridcol="5" gridrow="6" pageId="33" pageNumber="434">3</td>
</tr>
<tr id="B0B33C71FFAEF386FF6DFE6A5A013A71" box="[159,1302,498,520]" gridrow="7" pageId="33" pageNumber="434">
<th id="F362550DFFAEF386FF6DFE6A5EB63A71" box="[159,417,498,520]" gridcol="0" gridrow="7" pageId="33" pageNumber="434">Ancestrula length</th>
<td id="F362550DFFAEF386FDA0FE6A5D9B3A71" box="[594,652,498,520]" gridcol="1" gridrow="7" pageId="33" pageNumber="434">0.396</td>
<td id="F362550DFFAEF386FD0EFE6A5C543A71" box="[764,835,498,520]" gridcol="2" gridrow="7" pageId="33" pageNumber="434">0.0368</td>
<td id="F362550DFFAEF386FC55FE6A5B1B3A71" box="[935,1036,498,520]" gridcol="3" gridrow="7" pageId="33" pageNumber="434">0.370</td>
<td id="F362550DFFAEF386FBA0FE6A5BAC3A71" box="[1106,1211,498,520]" gridcol="4" gridrow="7" pageId="33" pageNumber="434">0.422</td>
<td id="F362550DFFAEF386FB0EFE6A5A013A71" box="[1276,1302,498,520]" gridcol="5" gridrow="7" pageId="33" pageNumber="434">2</td>
</tr>
<tr id="B0B33C71FFAEF386FF6DFD845A013A4B" box="[159,1302,540,562]" gridrow="8" pageId="33" pageNumber="434">
<th id="F362550DFFAEF386FF6DFD845EB63A4B" box="[159,417,540,562]" gridcol="0" gridrow="8" pageId="33" pageNumber="434">Ancestrula width</th>
<td id="F362550DFFAEF386FDA0FD845D9B3A4B" box="[594,652,540,562]" gridcol="1" gridrow="8" pageId="33" pageNumber="434">0.348</td>
<td id="F362550DFFAEF386FD0EFD845C543A4B" box="[764,835,540,562]" gridcol="2" gridrow="8" pageId="33" pageNumber="434">0.0601</td>
<td id="F362550DFFAEF386FC55FD845B1B3A4B" box="[935,1036,540,562]" gridcol="3" gridrow="8" pageId="33" pageNumber="434">0.305</td>
<td id="F362550DFFAEF386FBA0FD845BAC3A4B" box="[1106,1211,540,562]" gridcol="4" gridrow="8" pageId="33" pageNumber="434">0.390</td>
<td id="F362550DFFAEF386FB0EFD845A013A4B" box="[1276,1302,540,562]" gridcol="5" gridrow="8" pageId="33" pageNumber="434">2</td>
</tr>
<tr id="B0B33C71FFAEF386FF6DFDDE5A013A25" box="[159,1302,582,604]" gridrow="9" pageId="33" pageNumber="434">
<th id="F362550DFFAEF386FF6DFDDE5EB63A25" box="[159,417,582,604]" gridcol="0" gridrow="9" pageId="33" pageNumber="434">Ancestrular opesia length</th>
<td id="F362550DFFAEF386FDA0FDDE5D9B3A25" box="[594,652,582,604]" gridcol="1" gridrow="9" pageId="33" pageNumber="434">0.132</td>
<td id="F362550DFFAEF386FD0EFDDE5C543A25" box="[764,835,582,604]" gridcol="2" gridrow="9" pageId="33" pageNumber="434">0.0148</td>
<td id="F362550DFFAEF386FC55FDDE5B1B3A25" box="[935,1036,582,604]" gridcol="3" gridrow="9" pageId="33" pageNumber="434">0.121</td>
<td id="F362550DFFAEF386FBA0FDDE5BAC3A25" box="[1106,1211,582,604]" gridcol="4" gridrow="9" pageId="33" pageNumber="434">0.142</td>
<td id="F362550DFFAEF386FB0EFDDE5A013A25" box="[1276,1302,582,604]" gridcol="5" gridrow="9" pageId="33" pageNumber="434">2</td>
</tr>
<tr id="B0B33C71FFAEF386FF6DFDE85A013AFF" box="[159,1302,624,646]" gridrow="10" pageId="33" pageNumber="434">
<th id="F362550DFFAEF386FF6DFDE85EB63AFF" box="[159,417,624,646]" gridcol="0" gridrow="10" pageId="33" pageNumber="434">Ancestrular opesia width</th>
<td id="F362550DFFAEF386FDA0FDE85D9B3AFF" box="[594,652,624,646]" gridcol="1" gridrow="10" pageId="33" pageNumber="434">0.183</td>
<td id="F362550DFFAEF386FD0EFDE85C543AFF" box="[764,835,624,646]" gridcol="2" gridrow="10" pageId="33" pageNumber="434">0.0269</td>
<td id="F362550DFFAEF386FC55FDE85B1B3AFF" box="[935,1036,624,646]" gridcol="3" gridrow="10" pageId="33" pageNumber="434">0.164</td>
<td id="F362550DFFAEF386FBA0FDE85BAC3AFF" box="[1106,1211,624,646]" gridcol="4" gridrow="10" pageId="33" pageNumber="434">0.202</td>
<td id="F362550DFFAEF386FB0EFDE85A013AFF" box="[1276,1302,624,646]" gridcol="5" gridrow="10" pageId="33" pageNumber="434">2</td>
</tr>
</table>
</paragraph>
<paragraph id="0E3C3E33FFAE0C58FF65FD325DD93AB8" blockId="33.[151,718,682,705]" box="[151,718,682,705]" pageId="33" pageNumber="434">SD, standard deviation; N, number of measurements.</paragraph>
<paragraph id="0E3C3E33FFAE0C58FF35FD705C483BE8" blockId="33.[151,1436,744,1525]" pageId="33" pageNumber="434">Ancestrula oval, smaller than autozooids, gymnocystal walls steeply sloping, well-developed proximally, narrowing distally; opesia confined to distal part of frontal area, transversely oval, proximally constricted by extensive flat cryptocystal shelf with granular surface and narrow smooth distal rim, thin mural rim distinctly raised, surrounded by 12 spines, proximalmost spine directed distally and overarching frontal surface; first daughter autozooid budded distally, second zooid distolaterally.</paragraph>
</subSubSection>
<subSubSection id="46996DB8FFAE0C58FF35FC045BD63D8C" pageId="33" pageNumber="434" type="discussion">
<paragraph id="0E3C3E33FFAE0C58FF35FC045A173DF0" blockId="33.[151,1436,744,1525]" pageId="33" pageNumber="434">
<emphasis id="3CF7E221FFAE0C58FF35FC045E2C3BCC" bold="true" box="[199,315,924,949]" pageId="33" pageNumber="434">Remarks.</emphasis>
The new species is closely related to
<taxonomicName id="C98345B0FFAE0C58FD1EFC045B6E3BCC" authority="Calvet, 1907" authorityName="Calvet" authorityYear="1907" box="[748,1145,924,949]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="species" species="praealta">
<emphasis id="3CF7E221FFAE0C58FD1EFC045CDB3BCC" box="[748,972,924,949]" italics="true" pageId="33" pageNumber="434">Escharella praealta</emphasis>
(Calvet, 1907)
</taxonomicName>
, originally recorded from more than
<quantity id="C97B93D6FFAE0C58FEFFFC595E433BA1" box="[269,340,961,985]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="33" pageNumber="434" unit="m" value="700.0">700 m</quantity>
depth in the Gulf of Cádiz and later reported from the Canary Islands (as
<taxonomicName id="C98345B0FFAE0C58FB74FC585E5B3B84" authority="Aristegui, 1986" authorityName="Aristegui" authorityYear="1986" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="species" species="hexaespinosa">
<emphasis id="3CF7E221FFAE0C58FB74FC585A8B3BA0" box="[1158,1436,960,985]" italics="true" pageId="33" pageNumber="434">Escharella hexaespinosa</emphasis>
Arístegui, 1986
</taxonomicName>
), from the Strait of
<collectingCountry id="76947EA3FFAE0C58FDC5FC7C5D8B3B84" box="[567,668,996,1021]" name="Gibraltar" pageId="33" pageNumber="434">Gibraltar</collectingCountry>
at depths as shallow as
<quantity id="C97B93D6FFAE0C58FC4FFC7D5CEE3B85" box="[957,1017,997,1021]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="434" unit="m" value="30.0">30 m</quantity>
(see López de la Cuadra &amp; García- Gómez 1993), and from
<quantity id="C97B93D6FFAE0C58FE48FB915D043C59" box="[442,531,1033,1057]" metricMagnitude="3" metricUnit="m" metricValue="2.018" pageId="33" pageNumber="434" unit="m" value="2018.0">2018 m</quantity>
off NW
<collectingCountry id="76947EA3FFAE0C58FD70FB905DD53C58" box="[642,706,1032,1057]" name="Spain" pageId="33" pageNumber="434">Spain</collectingCountry>
(Reverter-Gil &amp; Fernandez-Pulpeiro 1999b). The anvil-shaped lyrula typical of most
<taxonomicName id="C98345B0FFAE0C58FE69FBB45D043C3C" box="[411,531,1068,1093]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="genus">
<emphasis id="3CF7E221FFAE0C58FE69FBB45D043C3C" box="[411,531,1068,1093]" italics="true" pageId="33" pageNumber="434">Escharella</emphasis>
</taxonomicName>
species is reduced to a triangular denticle in both
<taxonomicName id="C98345B0FFAE0C58FB97FBB65BF03C3C" box="[1125,1255,1068,1093]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="species" species="praealta">
<emphasis id="3CF7E221FFAE0C58FB97FBB65BF03C3C" box="[1125,1255,1068,1093]" italics="true" pageId="33" pageNumber="434">E. praealta</emphasis>
</taxonomicName>
and
<taxonomicName id="C98345B0FFAE0C58FAD6FBB45FF93C10" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="species" species="lopezfei" status="sp. nov.">
<emphasis id="3CF7E221FFAE0C58FAD6FBB45FF93C10" italics="true" pageId="33" pageNumber="434">Escharella lopezfei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27C45F5AFFAE0C58FF07FBC85E223C10" box="[245,309,1104,1129]" pageId="33" pageNumber="434" rank="species">
<emphasis id="3CF7E221FFAE0C58FF07FBC85E223C10" bold="true" box="[245,309,1104,1129]" pageId="33" pageNumber="434">n. sp.</emphasis>
</taxonomicNameLabel>
, which may, however, have the same function as a typical smittinid lyrula (cf. Berning
<emphasis id="3CF7E221FFAE0C58FAE1FBC95A523C10" box="[1299,1349,1104,1129]" italics="true" pageId="33" pageNumber="434">et al</emphasis>
. 2014). The new species is distinguished by the lack of an umbo proximal to the orifice and on the proximal ooecium margins, and in having 89 oral spines, whereas
<taxonomicName id="C98345B0FFAE0C58FD30FB015C563CC8" box="[706,833,1176,1201]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="species" species="praealta">
<emphasis id="3CF7E221FFAE0C58FD30FB015C563CC8" box="[706,833,1176,1201]" italics="true" pageId="33" pageNumber="434">E. praealta</emphasis>
</taxonomicName>
has only six spines. Moreover, the marginal pores in the frontal shield are much less conspicuous in
<taxonomicName id="C98345B0FFAE0C58FD49FB265C243CAC" box="[699,819,1212,1237]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="species" species="lopezfei" status="sp. nov.">
<emphasis id="3CF7E221FFAE0C58FD49FB265C243CAC" box="[699,819,1212,1237]" italics="true" pageId="33" pageNumber="434">E. lopezfei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27C45F5AFFAE0C58FCCEFB245C693CAC" box="[828,894,1212,1237]" pageId="33" pageNumber="434" rank="species">
<emphasis id="3CF7E221FFAE0C58FCCEFB245C693CAC" bold="true" box="[828,894,1212,1237]" pageId="33" pageNumber="434">n. sp.</emphasis>
</taxonomicNameLabel>
than in the shallow-water specimen figured by López de la Cuadra &amp; García-Gómez (1993, fig. 1H). In contrast, the deep-water populations of
<taxonomicName id="C98345B0FFAE0C58FB12FB795A483C80" box="[1248,1375,1248,1273]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="species" species="praealta">
<emphasis id="3CF7E221FFAE0C58FB12FB795A483C80" box="[1248,1375,1248,1273]" italics="true" pageId="33" pageNumber="434">E. praealta</emphasis>
</taxonomicName>
have equally few pores as in
<taxonomicName id="C98345B0FFAE0C58FE56FA9E5D0C3D64" box="[420,539,1284,1309]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="species" species="lopezfei">
<emphasis id="3CF7E221FFAE0C58FE56FA9E5EA23D64" box="[420,437,1286,1309]" italics="true" pageId="33" pageNumber="434">E</emphasis>
.
<emphasis id="3CF7E221FFAE0C58FE36FA9C5D0C3D64" box="[452,539,1284,1309]" italics="true" pageId="33" pageNumber="434">lopezfei</emphasis>
</taxonomicName>
, which can be seen in the original drawing by Calvet (1907, pl. 28, fig. 6), and also in López de la Cuadra &amp; García-Gómez' (1993, fig. 2) camera-lucida drawing of the
<typeStatus id="D1388091FFAE0C58FB4DFAB05A323D38" box="[1215,1317,1320,1345]" pageId="33" pageNumber="434" type="holotype">holotype</typeStatus>
. It is thus possible that the shallow- and deep-water populations represent distinct species. The great bathymetric range from
<quantity id="C97B93D6FFAE0C58FF65FAE95FC73DF1" box="[151,208,1393,1417]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="434" unit="m" value="30.0">30 m</quantity>
to over
<quantity id="C97B93D6FFAE0C58FEDEFAE95E643DF1" box="[300,371,1393,1417]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="33" pageNumber="434" unit="m" value="700.0">700 m</quantity>
was already highlighted by López de la Cuadra &amp; García-Gómez (1993, p. 465).
</paragraph>
<paragraph id="0E3C3E33FFAE0C58FF35FA0C5B803DA9" blockId="33.[151,1436,744,1525]" pageId="33" pageNumber="434">
Anatomical studies have shown that the ovicells are acleithral in
<taxonomicName id="C98345B0FFAE0C58FC1BFA0C5E483DA8" authority="(Fleming, 1828) (Ostrovsky 2013)" authorityName="(Fleming, 1828) (Ostrovsky" authorityYear="2013" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="species" species="immersa">
<emphasis id="3CF7E221FFAE0C58FC1BFA0C5BC73DD4" box="[1001,1232,1428,1453]" italics="true" pageId="33" pageNumber="434">Escharella immersa</emphasis>
(Fleming, 1828) (Ostrovsky 2013)
</taxonomicName>
. We suggest that the same
<typeStatus id="D1388091FFAE0C58FD79FA215DAC3DA8" box="[651,699,1465,1489]" pageId="33" pageNumber="434">type</typeStatus>
of ovicell closure is present in
<taxonomicName id="C98345B0FFAE0C58FBE8FA225B873DA8" box="[1050,1168,1464,1489]" class="Gymnolaemata" family="Romancheinidae" genus="Escharella" kingdom="Animalia" order="Cheilostomatida" pageId="33" pageNumber="434" phylum="Bryozoa" rank="species" species="lopezfei">
<emphasis id="3CF7E221FFAE0C58FBE8FA225B3C3DA8" box="[1050,1067,1466,1489]" italics="true" pageId="33" pageNumber="434">E</emphasis>
.
<emphasis id="3CF7E221FFAE0C58FBCBFA205B873DA8" box="[1081,1168,1464,1489]" italics="true" pageId="33" pageNumber="434">lopezfei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0E3C3E33FFAE0C58FF35FA445BD63D8C" blockId="33.[151,1436,744,1525]" box="[199,1217,1500,1525]" pageId="33" pageNumber="434">
The species was recorded from one station (VH-07-315) at
<quantity id="C97B93D6FFAE0C58FC93FA455CBF3D8D" box="[865,936,1501,1525]" metricMagnitude="2" metricUnit="m" metricValue="8.23" pageId="33" pageNumber="434" unit="m" value="823.0">823 m</quantity>
depth, encrusting corals.
</paragraph>
</subSubSection>
</treatment>
</document>