285 lines
36 KiB
XML
285 lines
36 KiB
XML
<document id="4EEFF5308C1503457FFAB7224FD63962" ID-DOI="10.11646/zootaxa.3694.4.1" ID-GBIF-Dataset="26650535-f621-48d3-9b71-cb68e50edeb5" ID-ISSN="1175-5326" ID-Zenodo-Dep="217543" ID-ZooBank="6F2F58C1-3927-4E66-B09C-AFAF4B4A2C71" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460658430822" checkinUser="plazi" docAuthor="David, Patrick, Vogel, Gernot & Rooijen, Johan Van" docDate="2013" docId="855087F67E01FFA1FF5F73F7B04366BD" docLanguage="en" docName="zt03694p335.pdf" docOrigin="Zootaxa 3694 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphiesma kerinciense David & Das 2003" docType="treatment" docVersion="6" lastPageNumber="324" masterDocId="7969FF8E7E15FFB6FFC8770CB5416710" masterDocTitle="On some taxonomically confused species of the genus Amphiesma Duméril, Bibron & Duméril, 1854 related to Amphiesma khasiense (Boulenger, 1890) (Squamata, Natricidae)" masterLastPageNumber="335" masterPageNumber="301" pageNumber="321" updateTime="1698347242376" updateUser="plazi">
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<mods:mods id="63423FE53CE91752C05997CC8559F0A9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A536A805E59029A0BE1FDAE7A9F0DEA8">
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<mods:title id="89D6679AA1BB1A95EF1BC5743958EB56">On some taxonomically confused species of the genus Amphiesma Duméril, Bibron & Duméril, 1854 related to Amphiesma khasiense (Boulenger, 1890) (Squamata, Natricidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="98358BDFDA208CA99144AA861C9927A2" type="personal">
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<mods:role id="03E522AFD97EF6631DD228DD60D1F22F">
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<mods:roleTerm id="8CEB346E223D204ADA28157191CC7D6A">Author</mods:roleTerm>
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<mods:namePart id="E9F8B62CA5B1E8C92468F157A7981458">David, Patrick</mods:namePart>
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<mods:roleTerm id="3C901717381AB3545ACDB46B3460D370">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4354DE8990A746C2CD66EDDD91758287">Vogel, Gernot</mods:namePart>
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</mods:name>
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<mods:name id="BFCA49F8E88E3C84525C0DE1F974AA08" type="personal">
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<mods:roleTerm id="3B7FF23948E78AD889011DBA9249695C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="164634BDEB63089BFABC0F154E45DA35">Rooijen, Johan Van</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="177E1C486A323124AD0194BDD40E2653">text</mods:typeOfResource>
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<mods:title id="5F5D7AA0C528442470D458BB288A3B6A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CA1CC2F22451413B292240EF948E8DED">
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<mods:date id="35D1062A377587A875D03E1B74AAB5FC">2013</mods:date>
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<mods:detail id="4449FA15110C5AF11476609B4C93E53F" type="volume">
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<mods:number id="CF4F1DD719FE27052CF052E9A6511F9F">3694</mods:number>
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</mods:detail>
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<mods:detail id="E5318D38DEDD16BEE0F08B3851D47893" type="issue">
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<mods:number id="E2CD9823CDA11F5B9ABD1F9BD0A69B8D">4</mods:number>
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<mods:start id="E3EE42E081F1E1CFDB3E7BBBAA989C77">301</mods:start>
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<mods:end id="6BC6FEFF7C03DF9E97B4624FD8EC0DA8">335</mods:end>
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<mods:classification id="391909C23BACB30F79D5AD054ABF0B1A">journal article</mods:classification>
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<mods:identifier id="7B46A4E612B6F549D908D2D4128646FF" type="DOI">10.11646/zootaxa.3694.4.1</mods:identifier>
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<mods:identifier id="62141D20A956281DBDE4D601EFAD8B78" type="GBIF-Dataset">26650535-f621-48d3-9b71-cb68e50edeb5</mods:identifier>
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<mods:identifier id="5C96601A76B1F71FB11C24CE51B9A926" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="38208DEF96F8BD7DA533790DB5A0AD93" type="Zenodo-Dep">217543</mods:identifier>
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<mods:identifier id="EEB145B788CC443A0F74A4B0812DB4D7" type="ZooBank">6F2F58C1-3927-4E66-B09C-AFAF4B4A2C71</mods:identifier>
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<treatment id="855087F67E01FFA1FF5F73F7B04366BD" ID-DOI="http://doi.org/10.5281/zenodo.6165428" ID-GBIF-Taxon="119574062" ID-Zenodo-Dep="6165428" LSID="urn:lsid:plazi:treatment:855087F67E01FFA1FF5F73F7B04366BD" httpUri="http://treatment.plazi.org/id/855087F67E01FFA1FF5F73F7B04366BD" lastPageId="23" lastPageNumber="324" pageId="20" pageNumber="321">
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<subSubSection id="45E3656B7E01FFA2FF5F73F7B7BA628B" pageId="20" pageNumber="321" type="nomenclature">
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<paragraph id="0D4636E07E01FFA2FF5F73F7B7EB6206" blockId="20.[151,682,1275,1336]" box="[151,682,1275,1302]" pageId="20" pageNumber="321">
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<heading id="560E818C7E01FFA2FF5F73F7B7EB6206" bold="true" box="[151,682,1275,1302]" fontSize="11" level="1" pageId="20" pageNumber="321" reason="1">
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<taxonomicName id="CAF94D637E01FFA2FF5F73F7B7EB6206" authority="David & Das, 2003" authorityName="David & Das" authorityYear="2003" box="[151,682,1275,1302]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="kerinciense">
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<emphasis id="3F8DEAF27E01FFA2FF5F73F7B7EB6206" bold="true" box="[151,682,1275,1302]" pageId="20" pageNumber="321">
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<emphasis id="3F8DEAF27E01FFA2FF5F73F7B4F66205" bold="true" box="[151,439,1275,1301]" italics="true" pageId="20" pageNumber="321">Amphiesma kerinciense</emphasis>
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David & Das, 2003
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="0D4636E07E01FFA2FF5F7213B5AA6227" blockId="20.[151,682,1275,1336]" box="[151,235,1311,1336]" pageId="20" pageNumber="321">
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(
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<figureCitation id="95C22A657E01FFA2FF577213B5A26227" box="[159,227,1311,1336]" captionStart="FIGURE 7" captionStartId="21.[151,250,1352,1374]" captionTargetBox="[226,1360,481,1330]" captionTargetId="figure@21.[226,1360,481,1331]" captionTargetPageId="21" captionText="FIGURE 7. Head portrait of Amphiesma kerinciense David & Das, 2003, ZRC 2.3521, holotype, adult female. Photo Patrick David." httpUri="https://zenodo.org/record/217550/files/figure.png" pageId="20" pageNumber="321">Fig. 7</figureCitation>
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)
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</paragraph>
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<paragraph id="0D4636E07E01FFA2FF5F726BB7BA628B" blockId="20.[151,1436,1383,1589]" pageId="20" pageNumber="321">
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<taxonomicName id="CAF94D637E01FFA2FF5F726BB403626D" box="[151,322,1383,1405]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="inas">
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<emphasis id="3F8DEAF27E01FFA2FF5F726BB403626D" box="[151,322,1383,1405]" italics="true" pageId="20" pageNumber="321">Amphiesma inas</emphasis>
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</taxonomicName>
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(nec
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<taxonomicName id="CAF94D637E01FFA2FE4B726BB7A1626D" authority="Laidlaw, 1901" authorityName="Laidlaw" authorityYear="1901" box="[387,736,1383,1405]" class="Squamata" family="Colubridae" genus="Tropidonotus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="inas">
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<emphasis id="3F8DEAF27E01FFA2FE4B726BB77C626D" box="[387,573,1383,1405]" italics="true" pageId="20" pageNumber="321">Tropidonotus inas</emphasis>
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Laidlaw, 1901
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</taxonomicName>
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, a valid species).—Mumpuni 2001: 38—David & Das 2003: 417 (citation of the specimen described by Mumpuni 2001).
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</paragraph>
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</subSubSection>
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<subSubSection id="45E3656B7E01FFA2FF5F72A8B7016125" pageId="20" pageNumber="321" type="description">
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<paragraph id="0D4636E07E01FFA2FF5F72A8B4416106" blockId="20.[151,1436,1383,1589]" pageId="20" pageNumber="321">
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<taxonomicName id="CAF94D637E01FFA2FF5F72A8B7C962AA" authority="David & Das, 2003: 414" authorityName="David & Das" authorityPageNumber="414" authorityYear="2003" box="[151,648,1444,1466]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="kerinciense">
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<emphasis id="3F8DEAF27E01FFA2FF5F72A8B4C762AA" box="[151,390,1444,1466]" italics="true" pageId="20" pageNumber="321">Amphiesma kerinciense</emphasis>
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David & Das, 2003: 414
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</taxonomicName>
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, 415:
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<figureCitation id="95C22A657E01FFA2FD0D72A8B65762AA" box="[709,790,1444,1466]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="5.[151,250,1149,1171]" captionStartId-1="6.[151,250,1561,1583]" captionStartId-2="10.[151,250,1970,1992]" captionStartId-3="11.[151,250,1268,1290]" captionTargetBox-0="[184,1369,201,1148]" captionTargetBox-1="[228,1358,587,1480]" captionTargetBox-2="[279,1307,193,1948]" captionTargetBox-3="[151,1436,193,1246]" captionTargetId-0="figure@5.[184,1404,146,1155]" captionTargetId-1="figure@6.[151,1436,504,1540]" captionTargetId-2="figure@10.[279,1307,193,1949]" captionTargetId-3="figure@11.[151,1436,193,1247]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="FIGURE 1. Object scores of first principal component along the geographic coordinates: circles: A. khasiense, males; triangles pointed upwards: A. khasiense, females; triangles pointed downwards: “ A. cf. khasiense ” of the Indochinese Peninsula, males; triangles pointed to the right: “ A. cf. khasiense ” of the Indochinese Peninsula, females; triangles pointed to the left: A. boulengeri, males; pentagons: A. boulengeri, females; squares: A. inas, males; diamonds: A. inas, females." captionText-1="FIGURE 2. Object scores of first principal component along the geographic coordinates: circles: A. boulengeri, Cambodia, males; triangles pointed upwards: A. boulengeri, Cambodia, females; triangles pointed downwards: A. boulengeri, males; triangles pointed to the right: A. boulengeri, females." captionText-2="FIGURE 3. One of the syntypes of Tropidonotus khasiensis, BMNH 1946.1. 12.81, Khasi Hills, India. Photos Gernot Vogel." captionText-3="FIGURE 4. Comparison of the pattern of the upper labials of three species of the Amphiesma khasiense group. A. A. boulengeri, BMNH 1969.1710, Kampot Province, Cambodia. B. A. boulengeri, FMNH 180153, Nakhon Ratchasima Province, Thailand. C. A. modestum, BMNH 1925.9. 17.2, Kachin Hills, Myanmar. D. A. khasiense, BMNH 1946.1. 12.82, syntype, State of Meghalaya, India. E. A. khasiense, BMNH 1946.1. 13.63, holotype of Natrix gilhodesi, Kachin State, Myanmar. F. A. khasiense, BMNH 1925.4. 2.13, Kachin State, Myanmar. Photos Gernot Vogel." httpUri-0="https://zenodo.org/record/217544/files/figure.png" httpUri-1="https://zenodo.org/record/217545/files/figure.png" httpUri-2="https://zenodo.org/record/217546/files/figure.png" httpUri-3="https://zenodo.org/record/217547/files/figure.png" pageId="20" pageNumber="321">Fig. 1–4</figureCitation>
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.—
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<typeStatus id="D24288427E01FFA2FCFD72A8B62D62AA" box="[821,876,1444,1466]" pageId="20" pageNumber="321">Type</typeStatus>
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locality. Sungai Jalnei Dalam, at base of Gunung
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<collectingCountry id="75EE76707E01FFA2FAAF72A9B0DD62AA" box="[1383,1436,1445,1466]" name="Togo" pageId="20" pageNumber="321">Tugu</collectingCountry>
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(or Tujuh) [= Mt.
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<collectingCountry id="75EE76707E01FFA2FEB372C8B4EE62C9" box="[379,431,1476,1497]" name="Togo" pageId="20" pageNumber="321">Tugu</collectingCountry>
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or Tujuh] (
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<geoCoordinate id="68CD50277E01FFA2FDEB72CFB7F762C9" box="[547,694,1475,1497]" direction="south" orientation="latitude" pageId="20" pageNumber="321" precision="1" value="-1.716389">01°42'59.0"S-</geoCoordinate>
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<geoCoordinate id="68CD50277E01FFA2FD7E72CFB60862C9" box="[694,841,1475,1497]" direction="east" orientation="longitude" pageId="20" pageNumber="321" precision="1" value="101.36197">101°21'43.1"E</geoCoordinate>
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), Gunung Kerinci [Mt. Kerinci], Sumatera Barat Province, Sumatra Island,
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<collectingCountry id="75EE76707E01FFA2FEB872EDB49462E7" box="[368,469,1505,1527]" name="Indonesia" pageId="20" pageNumber="321">Indonesia</collectingCountry>
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.—
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<typeStatus id="D24288427E01FFA2FE3B72EDB71562E7" box="[499,596,1505,1527]" pageId="20" pageNumber="321" type="holotype">Holotype</typeStatus>
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. ZRC 2.3521, adult female; collected by Darren Yeo and Heok Hui Tan,
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<date id="794710207E01FFA2FA8472EDB5BC6106" pageId="20" pageNumber="321" value="1996-06-12">12 June 1996</date>
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.
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</paragraph>
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<paragraph id="0D4636E07E01FFA2FF5F7113B7016125" blockId="20.[151,1436,1383,1589]" box="[151,576,1567,1589]" pageId="20" pageNumber="321">
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<taxonomicName id="CAF94D637E01FFA2FF5F7113B4C76125" box="[151,390,1567,1589]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="kerinciense">
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<emphasis id="3F8DEAF27E01FFA2FF5F7113B4C76125" box="[151,390,1567,1589]" italics="true" pageId="20" pageNumber="321">Amphiesma kerinciense</emphasis>
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</taxonomicName>
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.—Das 2010: 332.
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</paragraph>
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</subSubSection>
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<subSubSection id="45E3656B7E01FFA2FF5F716DB0DC6042" pageId="20" pageNumber="321" type="materials_examined">
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<paragraph id="0D4636E07E01FFA2FF5F716DB1DC61D2" blockId="20.[151,1437,1633,2018]" pageId="20" pageNumber="321">
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<emphasis id="3F8DEAF27E01FFA2FF5F716DB76C616A" bold="true" box="[151,557,1633,1658]" pageId="20" pageNumber="321">
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Material examined (
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<specimenCount id="1BFFFD697E01FFA2FE58716DB767616A" box="[400,550,1633,1658]" pageId="20" pageNumber="321" type="generic">2 specimens</specimenCount>
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)
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</emphasis>
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.
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<emphasis id="3F8DEAF27E01FFA2FDF7716DB7FA616A" bold="true" box="[575,699,1633,1658]" pageId="20" pageNumber="321">
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<collectingCountry id="75EE76707E01FFA2FDF7716DB7F7616A" box="[575,694,1633,1658]" name="Indonesia" pageId="20" pageNumber="321">Indonesia</collectingCountry>
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.
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</emphasis>
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<emphasis id="3F8DEAF27E01FFA2FD0E716EB6646169" box="[710,805,1634,1657]" italics="true" pageId="20" pageNumber="321">Sumatra</emphasis>
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. Lampung Province. MZB 2186, Kubu Perahu, Bukit Barisan Selatan National Park, near Lake Ranau. Sumatera Barat Province. ZRC 2.3521 (
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<typeStatus id="D24288427E01FFA2FB417189B1B0618E" box="[1161,1265,1669,1694]" pageId="20" pageNumber="321" type="holotype">holotype</typeStatus>
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), Sungai Jalnei Dalam, at base of Gunung
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<collectingCountry id="75EE76707E01FFA2FE0B71A6B4BC61D2" box="[451,509,1706,1730]" name="Togo" pageId="20" pageNumber="321">Tugu</collectingCountry>
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(or Tujuh) (
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<geoCoordinate id="68CD50277E01FFA2FD4E71A5B66861D2" box="[646,809,1705,1730]" direction="south" orientation="latitude" pageId="20" pageNumber="321" precision="1" value="-1.716389">01°42'59.0"S-</geoCoordinate>
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<geoCoordinate id="68CD50277E01FFA2FCE071A5B68D61D2" box="[808,972,1705,1730]" direction="east" orientation="longitude" pageId="20" pageNumber="321" precision="1" value="101.36197">101°21'43.1"E</geoCoordinate>
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), Gunung Kerinci.
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</paragraph>
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<paragraph id="0D4636E07E01FFA2FF0F71C1B0DC6042" blockId="20.[151,1437,1633,2018]" pageId="20" pageNumber="321">
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<emphasis id="3F8DEAF27E01FFA2FF0F71C1B49761F6" bold="true" box="[199,470,1741,1766]" pageId="20" pageNumber="321">Taxonomic comments.</emphasis>
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<taxonomicName id="CAF94D637E01FFA2FE2A71C1B7AE61F6" box="[482,751,1741,1766]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="kerinciense">
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<emphasis id="3F8DEAF27E01FFA2FE2A71C1B7AE61F6" box="[482,751,1741,1766]" italics="true" pageId="20" pageNumber="321">Amphiesma kerinciense</emphasis>
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</taxonomicName>
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was described from a single, female specimen. Mumpuni (2001) briefly described a specimen of
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<taxonomicName id="CAF94D637E01FFA2FDA271FDB7AC601A" box="[618,749,1777,1802]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="genus">
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<emphasis id="3F8DEAF27E01FFA2FDA271FDB7AC601A" box="[618,749,1777,1802]" italics="true" pageId="20" pageNumber="321">Amphiesma</emphasis>
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</taxonomicName>
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from the southern Barisan Range (MZB 2186) which she identified as
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<taxonomicName id="CAF94D637E01FFA2FEE2701AB438603D" box="[298,377,1814,1837]" class="Squamata" family="Colubridae" genus="Tropidonotus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="inas">
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<emphasis id="3F8DEAF27E01FFA2FEE2701AB438603D" box="[298,377,1814,1837]" italics="true" pageId="20" pageNumber="321">A. inas</emphasis>
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</taxonomicName>
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. After examination, this specimen proved to be referable to
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<taxonomicName id="CAF94D637E01FFA2FBEE7019B071603E" box="[1062,1328,1813,1838]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="kerinciense">
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<emphasis id="3F8DEAF27E01FFA2FBEE7019B071603E" box="[1062,1328,1813,1838]" italics="true" pageId="20" pageNumber="321">Amphiesma kerinciense</emphasis>
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</taxonomicName>
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. We here slightly expand the description of this species on the basis of the second known specimen and the first known male.
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</paragraph>
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</subSubSection>
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<subSubSection id="45E3656B7E01FFA2FF0F7051B7E6608A" pageId="20" pageNumber="321" type="description">
|
||
<paragraph id="0D4636E07E01FFA2FF0F7051B7E6608A" blockId="20.[151,1437,1633,2018]" pageId="20" pageNumber="321">
|
||
<taxonomicName id="CAF94D637E01FFA2FF0F7051B4906066" box="[199,465,1885,1910]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="kerinciense">
|
||
<emphasis id="3F8DEAF27E01FFA2FF0F7051B4906066" box="[199,465,1885,1910]" italics="true" pageId="20" pageNumber="321">Amphiesma kerinciense</emphasis>
|
||
</taxonomicName>
|
||
seems to be allied to the Indo-Malayan
|
||
<taxonomicName id="CAF94D637E01FFA2FC517051B1126065" box="[921,1107,1885,1910]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="inas">
|
||
<emphasis id="3F8DEAF27E01FFA2FC517051B1126065" box="[921,1107,1885,1910]" italics="true" pageId="20" pageNumber="321">Amphiesma inas</emphasis>
|
||
</taxonomicName>
|
||
but its affinities may also lie with
|
||
<taxonomicName id="CAF94D637E01FFA2FF07708DB7E2608A" authority="Smedley, 1932" authorityName="Smedley" authorityYear="1932" box="[207,675,1921,1946]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="species" species="sanguineum">
|
||
<emphasis id="3F8DEAF27E01FFA2FF07708DB4A1608A" box="[207,480,1921,1946]" italics="true" pageId="20" pageNumber="321">Amphiesma sanguineum</emphasis>
|
||
(Smedley, 1932)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="45E3656B7E01FFA3FF0F70A9B62766BD" lastPageId="21" lastPageNumber="322" pageId="20" pageNumber="321" type="diagnosis">
|
||
<paragraph id="0D4636E07E01FFA3FF0F70A9B62766BD" blockId="20.[151,1437,1633,2018]" lastBlockId="21.[151,1436,151,429]" lastPageId="21" lastPageNumber="322" pageId="20" pageNumber="321">
|
||
<emphasis id="3F8DEAF27E01FFA2FF0F70A9B40060AE" bold="true" box="[199,321,1957,1982]" pageId="20" pageNumber="321">Diagnosis.</emphasis>
|
||
A species of the genus
|
||
<taxonomicName id="CAF94D637E01FFA2FD8F70A9B78B60AE" box="[583,714,1957,1982]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="321" phylum="Chordata" rank="genus">
|
||
<emphasis id="3F8DEAF27E01FFA2FD8F70A9B78B60AE" box="[583,714,1957,1982]" italics="true" pageId="20" pageNumber="321">Amphiesma</emphasis>
|
||
</taxonomicName>
|
||
characterized by the combination of (1) a stout body; (2) nostrils directed laterally; (3) 20–21 maxillary teeth, gradually enlarged, the last 2 teeth distinctly enlarged; (4) 19 dorsal scale rows at midbody, strongly keeled on upper rows, many of which are distinctly notched posteriorly; scales of first dorsal row enlarged, feebly keeled; (5) dorsal pattern made of a broad, darker vertebral band, with or without irregularly placed dark scales producing faint and irregular, discontinuous crossbars, edged on each side by a faint pale ochre brown dorsolateral stripe, widening to produce three or four irregular blotches on the neck; (6) posterior supralabial slightly and irregularly paler, without definite blotches; (7) neck uniform or with a narrow pale streak; (8) venter pale with a row of well defined dark brown blotches at 3/4 of their width; (9) eye large, about 2.0 times the distance between the lower margins of eye and of lip; (10) 138–140 VEN; (11) internasals broadly truncated anteriorly, (12) 8 or 9 supralabials; and (13) 1 anterior temporal.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="598666687E00FFA3FF5F7244B59D626D" httpUri="https://zenodo.org/record/217550/files/figure.png" pageId="21" pageNumber="322" targetBox="[226,1360,481,1330]" targetPageId="21">
|
||
<paragraph id="0D4636E07E00FFA3FF5F7244B59D626D" blockId="21.[151,1436,1352,1405]" pageId="21" pageNumber="322">
|
||
<emphasis id="3F8DEAF27E00FFA3FF5F7244B455624E" bold="true" box="[151,276,1352,1374]" pageId="21" pageNumber="322">FIGURE 7.</emphasis>
|
||
Head portrait of
|
||
<taxonomicName id="CAF94D637E00FFA3FE0E7244B6C9624E" authority="David & Das, 2003" authorityName="David & Das" authorityYear="2003" box="[454,904,1352,1374]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="322" phylum="Chordata" rank="species" species="kerinciense">
|
||
<emphasis id="3F8DEAF27E00FFA3FE0E7244B7F7624E" box="[454,694,1352,1374]" italics="true" pageId="21" pageNumber="322">Amphiesma kerinciense</emphasis>
|
||
David & Das, 2003
|
||
</taxonomicName>
|
||
, ZRC 2.3521, holotype, adult female. Photo Patrick David.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="45E3656B7E00FFA3FF0F72A5B1E561F2" pageId="21" pageNumber="322" type="materials_examined">
|
||
<paragraph id="0D4636E07E00FFA3FF0F72A5B0666142" blockId="21.[151,1437,1449,2020]" pageId="21" pageNumber="322">
|
||
<emphasis id="3F8DEAF27E00FFA3FF0F72A5B71062D2" bold="true" box="[199,593,1449,1474]" pageId="21" pageNumber="322">
|
||
Variation (based on
|
||
<specimenCount id="1BFFFD697E00FFA3FE7272A5B70B62D2" box="[442,586,1449,1474]" pageId="21" pageNumber="322" type="generic">2 specimens</specimenCount>
|
||
)
|
||
</emphasis>
|
||
. Body stout, cylindrical, elongate; head rather elongate, oval, depressed in front of the eye, barely distinct from neck; snout long, blunt when viewed from above, rectangular in lateral view, amounting for about 26.0 % of HL or 1.6 times as long as diameter of eye in two specimens; nostrils large, crescentic, directed laterally and piercing in the middle of the nasal; eye large, about 2.0 times the distance between the lower margin of eye and the lower edge of lip, with a round pupil; tail long, cylindrical and tapering.
|
||
</paragraph>
|
||
<paragraph id="0D4636E07E00FFA3FF0F7151B7DE61AE" blockId="21.[151,1437,1449,2020]" pageId="21" pageNumber="322">
|
||
The maximal total length known is
|
||
<quantity id="CA019B057E00FFA3FDA07151B7886166" box="[616,713,1629,1654]" metricMagnitude="-1" metricUnit="m" metricValue="5.16" pageId="21" pageNumber="322" unit="mm" value="516.0">516 mm</quantity>
|
||
(SVL
|
||
<quantity id="CA019B057E00FFA3FCD37151B63F6166" box="[795,894,1629,1654]" metricMagnitude="-1" metricUnit="m" metricValue="3.5799999999999996" pageId="21" pageNumber="322" unit="mm" value="358.0">358 mm</quantity>
|
||
; TaL
|
||
<quantity id="CA019B057E00FFA3FC0B7151B1666166" box="[963,1063,1629,1654]" metricMagnitude="-1" metricUnit="m" metricValue="1.58" pageId="21" pageNumber="322" unit="mm" value="158.0">158 mm</quantity>
|
||
; specimen ZRC 2.3521; female,
|
||
<typeStatus id="D24288427E00FFA3FF5F718DB5BF618A" box="[151,254,1665,1690]" pageId="21" pageNumber="322" type="holotype">holotype</typeStatus>
|
||
). The SVL of the second available specimen, a juvenile male with an incomplete tail, is only
|
||
<quantity id="CA019B057E00FFA3FAFE718EB0D96189" box="[1334,1432,1666,1690]" metricMagnitude="-1" metricUnit="m" metricValue="2.77" pageId="21" pageNumber="322" unit="mm" value="277.0">277 mm</quantity>
|
||
. Ratio TaL / TL:
|
||
<quantity id="CA019B057E00FFA3FE8671AAB4EB61AE" box="[334,426,1701,1726]" metricMagnitude="-3" metricUnit="m" metricValue="7.8232" pageId="21" pageNumber="322" unit="in" value="0.308">0.308 in</quantity>
|
||
the female specimen.
|
||
</paragraph>
|
||
<paragraph id="0D4636E07E00FFA3FF0F71C7B1E561F2" blockId="21.[151,1437,1449,2020]" box="[199,1188,1737,1762]" pageId="21" pageNumber="322">
|
||
<emphasis id="3F8DEAF27E00FFA3FF0F71C7B47061F2" box="[199,305,1739,1762]" italics="true" pageId="21" pageNumber="322">Dentition</emphasis>
|
||
. Maxillary teeth 20–21, gradually enlarged, the last 2 teeth distinctly enlarged.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="45E3656B7E00FFA1FF0F71E1B76C67C5" lastPageId="23" lastPageNumber="324" pageId="21" pageNumber="322" type="description">
|
||
<paragraph id="0D4636E07E00FFA3FF0F71E1B6F36041" blockId="21.[151,1437,1449,2020]" pageId="21" pageNumber="322">
|
||
<emphasis id="3F8DEAF27E00FFA3FF0F71E1B4326016" box="[199,371,1773,1798]" italics="true" pageId="21" pageNumber="322">Body scalation</emphasis>
|
||
. DSR: 19–19–17 scale rows; dorsal scales rhomboedric, usually distinctly notched at their posterior extremity, strongly keeled with a narrow, sharp keel; dorsal scales, more strongly in the posterior half of the body on all rows; scales of 1st DSR feebly keeled or nearly smooth.
|
||
</paragraph>
|
||
<paragraph id="0D4636E07E00FFA3FF0F7051B5B26089" blockId="21.[151,1437,1449,2020]" pageId="21" pageNumber="322">
|
||
VEN: 138–140 (plus 0 or 2 preventrals); SC:
|
||
<quantity id="CA019B057E00FFA3FD097051B7BB6065" box="[705,762,1884,1909]" metricMagnitude="0" metricUnit="m" metricValue="2.2606" pageId="21" pageNumber="322" unit="in" value="89.0">89 in</quantity>
|
||
the adult female, all paired; ratio VEN / SC: 1.56; anal plate divided.
|
||
</paragraph>
|
||
<paragraph id="0D4636E07E00FFA3FF0F70ABB10760F4" blockId="21.[151,1437,1449,2020]" pageId="21" pageNumber="322">Position of the reduction to 6 scale rows around the tail at the level of 8th SC. Ratio length of the portion of tail with 4 dorsal scale rows / length of the portion of tail with 6 dorsal scale rows: 1.45.</paragraph>
|
||
<paragraph id="0D4636E07E03FFA0FF0F779BB6A8659C" blockId="22.[151,1436,151,2027]" pageId="22" pageNumber="323">
|
||
<emphasis id="3F8DEAF27E03FFA0FF0F779BB43667A0" box="[199,375,151,176]" italics="true" pageId="22" pageNumber="323">Head scalation</emphasis>
|
||
. Rostral trapezoidal wider than high, visible from above; nasals subrectangular, elongate, vertically divided on their lower half, with the posterior part larger and higher than anterior one; internasals subtriangular, rather small, in broad contact, about 1.0 to 1.2 times as long as wide, abruptly truncated anteriorly with anterior margin about 0.5-0.6 times the width of the posterior margin; 2 wide, subrectangular prefrontals, 1.6– 1.8 times as long as internasals; frontal hexagonal, shield-like with apex directed posteriorly, rather small, 1.4 times longer than wide, 1.9–2.0 times as long as prefrontal; 1 supraocular on each side, longer than wide, about 0.3 times as wide as frontal, narrower than internasals; two large parietals, 1.2–1.4 times longer than frontal, in contact for a length 1.1 times as great as the frontal length; 1 small, rectangular loreal scale, 0.8 times as high as long, in broad contact with the nasal; 8 or 9 SL, 2nd–8th or 9th longer than high; 1st and 2nd SL small and short, in contact with nasal; 2nd and 3rd SL in contact with the loreal; 4th–5th or 4th–6th SL entering orbit; 6th and 7th SL or 7th and 8th SL largest; 1 relatively large or 2 small preoculars on each side; 3 postoculars, the upper one largest; 0 or 1 subocular; 1 anterior temporal, narrow and elongate, with complete temporal formulas as 1 + 1 + 2 or 1 + 2 + 2; 10 or 11 infralabials, first pair in contact each with other, 1st–4th or 1st–5th
|
||
<collectingCountry id="75EE76707E03FFA0FD2E755DB6406578" box="[742,769,593,616]" name="Iceland" pageId="22" pageNumber="323">IL</collectingCountry>
|
||
in contact with anterior chin shields, 6th
|
||
<collectingCountry id="75EE76707E03FFA0FB1A755DB1AD6578" box="[1234,1260,593,616]" name="Iceland" pageId="22" pageNumber="323">IL</collectingCountry>
|
||
largest; mental scale small; posterior chin shields about 1.2 times longer than anterior ones.
|
||
</paragraph>
|
||
<paragraph id="0D4636E07E03FFA0FF0F7597B4AC6367" blockId="22.[151,1436,151,2027]" pageId="22" pageNumber="323">
|
||
<emphasis id="3F8DEAF27E03FFA0FF0F7597B70D65A4" box="[199,588,667,692]" italics="true" pageId="22" pageNumber="323">Coloration and pattern in alcohol</emphasis>
|
||
. Background colour of sides, up to 4th dorsal scale row row, dark reddish grey-brown, due to an intricate speckling of dark reddish-brown pigments on a pale greyish-ochre background, with 1st–2nd lower scale rows rather purple greyish-brown, slightly iridescent, with ventroposterior extremities of scales of the 1st row dark brown; widely scattered small blackish-brown spots on 2nd, 3rd and 4th dorsal rows; these dark brown spots produce faint and irregular, discontinuous crossbars, more visible on the posterior part of the body. A broad blackish-brown vertebral band on 5th–9th scale rows and the vertebral row, with indistinct irregular black crossbars. A dorsolateral stripe extending from the neck to the tail on top of 5th and the whole of 6th scale row, well defined and greyish-pink on the neck and foremost part of the body (up to 6th VEN), becoming discontinuous and broken into 4 or 5 irregular and diffuse blotches more or less connected, then changing into a diffuse, ill defined pale ochre-brown stripe. The tail is blackish-brown above, greyish-ochre on the upper part of the side in the extension of the dorsolateral band of the body, with a blackish-brown band on its lower side and extremities of subcaudals, divided into two equal parts by a narrow greyish-purple line in the extension of the colour present on 1st and 2nd scale rows of the body.
|
||
</paragraph>
|
||
<paragraph id="0D4636E07E03FFA0FF0F738EB60D61B1" blockId="22.[151,1436,151,2027]" pageId="22" pageNumber="323">Head dark reddish grey-brown above with ochre vermiculations, turning into blackish-brown on its posterior part; a fine speckling with purple greyish-brown on temporals; two small faint yellowish-white spots on the parietals and a very faint whitish-brown sagittal line just behind the parietals. Anterior supralabials are ochre brown, with brown speckling and edged with dark brown on their posterior part. Posterior supralabials light ivory cream; posterior part of 6th SL blackish-brown; 7th SL with a curved dark brown marking in the posterior part of its centre and with its posterior upper corner dark brown; 8th SL divided into a lower anterior part ivory cream and an upper posterior part dark reddish grey-brown, these two parts being separated by an oblique blackish-brown streak connecting the upper tip of the 7th SL to the lip edge at the junction of 8th and 9th SL; 9th SL dark reddish greybrown, with its lower part blackish-brown and its upper part ochre brown, narrowly bordered with blackish-brown above. An ill defined and narrow postocular streak, pale ochre brown narrowly edged below with blackish-brown, extending from the upper preocular to 9th SL through the anterior temporal and the upper tip of 8th SL. Behind the 9th SL, a narrow, irregular pale postocular streak, bent upwards; on the side of the neck, a very thin, greyish-pink stripe, connected both to the postocular stripe and the dorsolateral stripe. Throat, chin and ventral part of the neck ivory cream, uniform in their middle, with some dark brown speckling on the outer parts of the throat and lower side of the neck; posterior margins of infralabials dark brown.</paragraph>
|
||
<paragraph id="0D4636E07E03FFA0FF0F71BCB44B60FB" blockId="22.[151,1436,151,2027]" pageId="22" pageNumber="323">Venter uniformly ivory cream, with about 1/4th of the outer part of each scale purple greyish-brown as dorsal rows 1–2, becoming blackish-brown on the scale outer tip; a subrectangular, irregular blackish-brown spot between the purple greyish-brown and ivory parts; these spots become progressively larger backwards and are connected to the purple greyish-brown colour of the flank from about the 20th VEN onwards; before this point, the blackishbrown ventral spots are separated from the purple greyish-brown colour of the lower flank by a narrow ivory cream line, producing a thin and conspicuous but discontinuous series of dark brown spots at the bottom of the flanks; the ivory line is progressively replaced by the purple greyish-brown hue. Below, the tail is as the venter, bordered on each side by a blackish-brown stripe described above, with a middle line made of irregular and faint dark brown speckling.</paragraph>
|
||
<paragraph id="0D4636E07E02FFA1FF0F779BB48A67A0" blockId="23.[151,1437,151,429]" box="[199,459,151,176]" pageId="23" pageNumber="324">
|
||
<emphasis id="3F8DEAF27E02FFA1FF0F779BB40667A0" bold="true" box="[199,327,151,176]" pageId="23" pageNumber="324">Hemipenis</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph id="0D4636E07E02FFA1FF0F77B0B76C67C5" blockId="23.[151,1437,151,429]" box="[199,557,188,213]" pageId="23" pageNumber="324">
|
||
<emphasis id="3F8DEAF27E02FFA1FF0F77B0B4F667C5" bold="true" box="[199,439,188,213]" pageId="23" pageNumber="324">Sexual dimorphism.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="45E3656B7E02FFA1FF0F77D3B6A5660C" pageId="23" pageNumber="324" type="distribution">
|
||
<paragraph id="0D4636E07E02FFA1FF0F77D3B6A5660C" blockId="23.[151,1437,151,429]" pageId="23" pageNumber="324">
|
||
<emphasis id="3F8DEAF27E02FFA1FF0F77D3B41867E8" bold="true" box="[199,345,223,248]" pageId="23" pageNumber="324">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="75EE76707E02FFA1FEAF77D3B49D67E8" box="[359,476,223,248]" name="Indonesia" pageId="23" pageNumber="324">
|
||
<emphasis id="3F8DEAF27E02FFA1FEAF77D3B49D67E8" bold="true" box="[359,476,223,248]" pageId="23" pageNumber="324">Indonesia</emphasis>
|
||
</collectingCountry>
|
||
.
|
||
<emphasis id="3F8DEAF27E02FFA1FE2177ECB70967E7" box="[489,584,224,247]" italics="true" pageId="23" pageNumber="324">Sumatra</emphasis>
|
||
. Endemic to this island. Province of Sumatera Barat. Massif of Mt. Kerinci. Lampung Province. Bukit Barisan Selatan National Park, near Lake Ranau.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="45E3656B7E02FFA1FF0F762BB04366BD" pageId="23" pageNumber="324" type="biology_ecology">
|
||
<paragraph id="0D4636E07E02FFA1FF0F762BB04366BD" blockId="23.[151,1437,151,429]" pageId="23" pageNumber="324">
|
||
<emphasis id="3F8DEAF27E02FFA1FF0F762BB45E6650" bold="true" box="[199,287,295,320]" pageId="23" pageNumber="324">Biology</emphasis>
|
||
. The
|
||
<typeStatus id="D24288427E02FFA1FEAE762BB4896650" box="[358,456,295,320]" pageId="23" pageNumber="324" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="CAF94D637E02FFA1FE3D7624B7D76650" box="[501,662,295,320]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="324" phylum="Chordata" rank="species" species="kerinciense">
|
||
<emphasis id="3F8DEAF27E02FFA1FE3D7624B7D76650" box="[501,662,295,320]" italics="true" pageId="23" pageNumber="324">A. kerinciense</emphasis>
|
||
</taxonomicName>
|
||
was collected in a shallow hill stream, less than
|
||
<quantity id="CA019B057E02FFA1FB127624B062662F" box="[1242,1315,296,320]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="324" unit="cm" value="30.0">30 cm</quantity>
|
||
deep, fast flowing in parts through an open, grassy area. The elevation was unfortunately not recorded. The clear water flowed over a substratum of stone and gravel. The snake was collected while it was feeding on tadpoles, possibly of the genus
|
||
<taxonomicName id="CAF94D637E02FFA1FEC07699B40166BC" box="[264,320,405,428]" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="23" pageNumber="324" phylum="Chordata" rank="genus">
|
||
<emphasis id="3F8DEAF27E02FFA1FEC07699B40166BC" box="[264,320,405,428]" italics="true" pageId="23" pageNumber="324">Huia</emphasis>
|
||
</taxonomicName>
|
||
, which it later regurgitated. Nothing else is known on the biology of this rare species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |