127 lines
12 KiB
XML
127 lines
12 KiB
XML
<document id="81311EE8E2AB51A50406A54EC118358A" ID-CLB-Dataset="33981" ID-DOI="http://dx.doi.org/10.3897/zookeys.541.6058" ID-GBIF-Dataset="fa40454d-8f0e-4387-86ee-8dfc421d0e05" ID-PMC="PMC4714381" ID-Pensoft-Pub="1313-2970-541-109" ID-PubMed="26798284" ID-ZooBank="C336A3C4DBCB49C5898C8FA38BDFF0C0" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-541-109" ModsDocOrigin="ZooKeys 541" ModsDocTitle="Systematics of Nothopsini (Serpentes, Dipsadidae), with a new species of Synophis from the Pacific Andean slopes of southwestern Ecuador" checkinTime="1451243656936" checkinUser="pensoft" docAuthor="Pyron, R. Alexander, Guayasamin, Juan M., Penafiel, Nicolas, Bustamante, Lucas & Arteaga, Alejandro" docDate="2015" docId="8500E45155720D04BB8CDAA83FA73E0C" docLanguage="en" docName="ZooKeys 541: 109-147" docOrigin="ZooKeys 541" docSource="http://dx.doi.org/10.3897/zookeys.541.6058" docTitle="Synophis plectovertebralis Sheil & Grant 2001" docType="treatment" docVersion="6" lastPageNumber="129" masterDocId="FF8E84462424FFD3A52779666640FF82" masterDocTitle="Systematics of Nothopsini (Serpentes, Dipsadidae), with a new species of Synophis from the Pacific Andean slopes of southwestern Ecuador" masterLastPageNumber="147" masterPageNumber="109" pageNumber="128" updateTime="1701381387359" updateUser="ExternalLinkService">
|
|
<mods:mods id="54B8A7DA7BDBE348361F301577CADD5F" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="75739A4BEC6C621DD9CB47C0EDF87DB5">
|
|
<mods:title id="2C93A88F0407228A8D71443020CBDA83">Systematics of Nothopsini (Serpentes, Dipsadidae), with a new species of Synophis from the Pacific Andean slopes of southwestern Ecuador</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="166F0DF4021EA63201BDE48F4F20552D" type="personal">
|
|
<mods:role id="24E420A2381707E74E9B495B35BEEB55">
|
|
<mods:roleTerm id="4849969A5EB67B6E5345B95AE8376814">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="A0174C39F0496188A3C75A5AEE2EB38E">Pyron, R. Alexander</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="48C419A6C75C61C0BE7D79F1DED8F218" type="personal">
|
|
<mods:role id="587FAC14B7EFD4EB44639D984ED95178">
|
|
<mods:roleTerm id="60B666CDD7768C6CB74B54DE86EBF7E2">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="D6AC075AAE27AB9BD143B38823DC85CA">Guayasamin, Juan M.</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="197C27549A4ED9DBE0C56CDE72DF00A0" type="personal">
|
|
<mods:role id="6846BD9D44B68D4EB583C47D173CCA19">
|
|
<mods:roleTerm id="FF797012235F8F3191D4EC5B48CA7C9A">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="C068DE0CAE082441452CF2F6C4DA22D9">Penafiel, Nicolas</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="3A970B1A164B74C20C1A17C65D6A45E2" type="personal">
|
|
<mods:role id="AA84B8E40B55E8FC12A43760ECC8B9F9">
|
|
<mods:roleTerm id="85C6CE224F7030567E2103FC560DE819">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="78C7E009ABEB9259B3D3EA4225018F09">Bustamante, Lucas</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="4E85929E1E20FA4029CFD589525F9C3E" type="personal">
|
|
<mods:role id="1704CBDAEE95E6EB701F81EDAB87CA01">
|
|
<mods:roleTerm id="0571A487C1810A4E4811ADF15F13259E">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="EB1E9939962FECB7901D489039E17D8C">Arteaga, Alejandro</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource id="E8CFBCE0190C598782DFABE2327CFCCE">text</mods:typeOfResource>
|
|
<mods:relatedItem id="EC0095C3A258B438480E0FCCFB05E5FB" type="host">
|
|
<mods:titleInfo id="D226DCE8B9BA95C21E6FA35C740175BE">
|
|
<mods:title id="980C8F3EFD4C3BC4761D26B604A071B4">ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="CFCF9F02D8C6640B2ABC72D10F78CDE2">
|
|
<mods:date id="B4191B3D8FDF283DAD60174EF4BD204E">2015</mods:date>
|
|
<mods:detail id="9399D252BD1F4EE456461A3555674F65" type="volume">
|
|
<mods:number id="FBEC4BF68C9D501546A1F06994E0052C">541</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="FF5F760FA87BFD563B1FF905E925F67B" unit="page">
|
|
<mods:start id="2D41874C030E55AD6FE82113A372FAB8">109</mods:start>
|
|
<mods:end id="58B66983690A8FD23F34D83CBDE145C6">147</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location id="1647750E0C1ECE0273C72A60249FD331">
|
|
<mods:url id="EC8C65C804D43FED60DFA545A39036C4">http://dx.doi.org/10.3897/zookeys.541.6058</mods:url>
|
|
</mods:location>
|
|
<mods:classification id="90B4396620ABB0B0B170ACB7DB939407">journal article</mods:classification>
|
|
<mods:identifier id="2BD716332FB39FE988BB02DA24B97360" type="DOI">http://dx.doi.org/10.3897/zookeys.541.6058</mods:identifier>
|
|
<mods:identifier id="924770856B300E47FE8B9586CD1988D1" type="Pensoft-Pub">1313-2970-541-109</mods:identifier>
|
|
<mods:identifier id="8A4764707CF8171E39BD14DBB627458B" type="ZooBank">C336A3C4DBCB49C5898C8FA38BDFF0C0</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="8500E45155720D04BB8CDAA83FA73E0C" ID-GBIF-Taxon="127891632" LSID="urn:lsid:plazi:treatment:8500E45155720D04BB8CDAA83FA73E0C" httpUri="http://treatment.plazi.org/id/8500E45155720D04BB8CDAA83FA73E0C" lastPageId="20" lastPageNumber="129" pageId="19" pageNumber="128">
|
|
<subSubSection id="2818774212CEF08C2B874E5221626A1B" pageId="19" pageNumber="128" type="multiple">
|
|
<paragraph id="49960DAE7EB120C06C8A99DB8517C7F7" pageId="19" pageNumber="128">Taxon classification Animalia Squamata Colubridae</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C1B380634BD8D232E76727577C4A2ACB" pageId="19" pageNumber="128" type="nomenclature">
|
|
<paragraph id="E5377C257A3BAAAEEF7D461C9FD91044" pageId="19" pageNumber="128">
|
|
<taxonomicName id="A4D211994C6B806EB5C1D37506E847F0" ID-CoL="53YC8" authority="Sheil & Grant, 2001" authorityName="Sheil & Grant" authorityYear="2001" class="Squamata" family="Colubridae" genus="Synophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synophis plectovertebralis" pageId="19" pageNumber="128" phylum="Chordata" rank="species" species="plectovertebralis">Synophis plectovertebralis Sheil & Grant, 2001</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D026602F48D19CF50F70F59098E7CBB4" pageId="19" pageNumber="128" type="holotype">
|
|
<paragraph id="03B9EA24E4EB9ACBE226435D88FB2B12" pageId="19" pageNumber="128">Holotype.</paragraph>
|
|
<paragraph id="2F39CA98DC09ED757D458B937C92115B" pageId="19" pageNumber="128">UVC 11858, from Hacienda San Pedro, about 6 km south El Queremal, Municipio Dagua, Departamento del Valle del Cauca, Colombia.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="244C6643F1BD63EEB8493C86650A0CB8" pageId="19" pageNumber="128" type="paratype">
|
|
<paragraph id="D3402E43CD08E26160A648E1BDC5E88B" pageId="19" pageNumber="128">Paratype.</paragraph>
|
|
<paragraph id="F7F5093737E3E60AFC7511543B91E905" pageId="19" pageNumber="128">UVC 11580, from type locality.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="28997DC298CAFCB5E374582A729C4AD3" pageId="19" pageNumber="128" type="etymology">
|
|
<paragraph id="3F24BF712F5860E6C73E7FBE1BED57FE" pageId="19" pageNumber="128">Etymology.</paragraph>
|
|
<paragraph id="568FFF00DCD72BA103CA94D8E356A2E0" pageId="19" pageNumber="128">
|
|
From the Latin plecto- for
|
|
<normalizedToken id="9EA7C2778D7C13B81FE9808B15B5EEBC" originalValue="“braided”">"braided"</normalizedToken>
|
|
or
|
|
<normalizedToken id="3F284B652D34E27FA239159C8B06C1E4" originalValue="“woven”">"woven"</normalizedToken>
|
|
and veretbralis for
|
|
<normalizedToken id="F26B30D310342DE33F001FC2871603EA" originalValue="“vertebrae,”">"vertebrae,"</normalizedToken>
|
|
referring to the appearance of the interlocking zygapophyses viewed from above (
|
|
<bibRefCitation id="D4E09F0D81DFCCDE5135E5EAE6F9384A" author="Sheil, CA" journalOrPublisher="Journal of Herpetology" pageId="26" pageNumber="135" pagination="204 - 209" title="A new species of colubrid snake (Synophis) from western Colombia." url="10.2307/1566109" volume="35" year="2001">Sheil and Grant 2001</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="3AE82E2339F7BB91AFC828325145D9C1" lastPageId="20" lastPageNumber="129" pageId="19" pageNumber="128" type="description">
|
|
<paragraph id="B2D34733FE358E69A7416BC5B67E90EF" pageId="19" pageNumber="128">Description.</paragraph>
|
|
<paragraph id="9676B96C77E8EE27326B5F771EC52D27" lastPageId="20" lastPageNumber="129" pageId="19" pageNumber="128">
|
|
Relatively small (~200mm SVL) dipsadine snakes of the Pacific versant of the Andean Highlands of W Colombia, diagnosable by 24 maxillary teeth, 7 or 8 infralabials, 7 or 8 supralabials, fused prefrontals, internasals in contact, loreal present, 1 postocular, nuchal collar present, 144-147 ventrals, 79-91 subcaudals, dorsal scales in 19-19-17 weakly keeled rows, neural spines expanded and flattened, and later
|
|
<pageBreakToken id="666098760C42129BEFEBA5ACE5A67C1A" pageId="20" pageNumber="129" start="start">ally</pageBreakToken>
|
|
expanded zygapophyses forming a partially interlocking complex. The type locality is a middle elevation (~1800m) cloud forest. Both known specimens were collected in moist leaf litter; one was active at night. The stomach of the holotype contained a
|
|
<taxonomicName id="EAB016196A0ABB45715D3B09D82A4EA8" class="Squamata" family="Alopoglossidae" genus="Ptychoglossus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptychoglossus stenolepis" pageId="20" pageNumber="129" phylum="Chordata" rank="species" species="stenolepis">Ptychoglossus stenolepis</taxonomicName>
|
|
(
|
|
<taxonomicName id="3BD3C113BB2A4C4D0A5BC402B00E91ED" lsidName="Sauria" pageId="20" pageNumber="129" rank="suborder" suborder="Sauria">Sauria</taxonomicName>
|
|
:
|
|
<taxonomicName id="A6B5FA284D72458E074B450162840F71" family="Gymnophthalmidae" lsidName="" pageId="20" pageNumber="129" rank="family">Gymnophthalmidae</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7A4D7F96C3B120F8CECC476F1AECD9B8" pageId="20" pageNumber="129" type="notes">
|
|
<paragraph id="F150D9E2B28BED4FF91D9C70713E6432" pageId="20" pageNumber="129">Notes.</paragraph>
|
|
<paragraph id="75A371D1F5977D44EA6A904AFCFD66FD" pageId="20" pageNumber="129">
|
|
Known only from the holotype and paratype (apparently juveniles), though other material has apparently been collected in Colombia, near the type locality (T. Grant and E. Meneses-Pelayo, pers. comm.). The hemipenes have not been examined. A more detailed description of the two specimens is provided by
|
|
<bibRefCitation id="C09B64CAAA85240D45A3619FEC46465E" author="Sheil, CA" journalOrPublisher="Journal of Herpetology" pageId="26" pageNumber="135" pagination="204 - 209" title="A new species of colubrid snake (Synophis) from western Colombia." url="10.2307/1566109" volume="35" year="2001">Sheil and Grant (2001)</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="EEDED205A22DC9D382420DCCD57D7E2F" pageId="20" pageNumber="129">
|
|
Given our restriction of the name, we also provide the following re-description of the re-delimited
|
|
<taxonomicName id="F46AB2DB446B1DFDC495DC4FC18D6662" lsidName="" pageId="20" pageNumber="129" rank="tribe" tribe="Nothopsini">Nothopsini</taxonomicName>
|
|
. Note that we have not performed a comparative examination of a large series of preserved material, and these data are summarized from the literature (
|
|
<bibRefCitation id="A28E2700FC975EDCED504E6F71B0004C" author="Dunn, ER" journalOrPublisher="Copeia" pageId="24" pageNumber="133" pagination="255 - 261" title="The neotropical snake genus Nothopsis Cope." url="10.2307/1439148" volume="1957" year="1957">Dunn and Dowling 1957</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="60CB89F65072D28CDAC3509227C5FD04" author="Savage, JM" journalOrPublisher="University of Chicago Press" pageId="25" pageNumber="134" title="The Amphibians and Reptiles of Costa Rica: A Herpetofauna Between Two Continents, Between Two Seas" year="2002">Savage 2002</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="72061633A498ECB8246E2A1B92D60223" author="Kohler, G" journalOrPublisher="Herpeton Verlag Elke Kohler, Offenbach" pageId="24" pageNumber="133" title="Reptiles of Central America." year="2008">Kohler 2008</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="9B23E71D2EAB655078551C191856AAC2" author="McCranie, JR" journalOrPublisher="SSAR, Salt Lake City" pageId="25" pageNumber="134" title="The snakes of Honduras." year="2011">McCranie 2011</bibRefCitation>
|
|
) to provide a basis for future revisions.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |