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<mods:title id="24037D82EACC4C3E247559349FF8E203">Crocodylomorph and dinosaur tracks from the lowermost Jurassic of Le Veillon (western France): ichnotaxonomic revision of the type material (Lapparent collection)</mods:title>
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<mods:namePart id="9BCB8D64DD9C67ADA6971CD983D6B08E">Vullo, Romain</mods:namePart>
<mods:affiliation id="80A88D092CE15F237E7A2F8DE90F0F9B">Univ Rennes, CNRS, Géosciences Rennes, UMR 6118, 263 avenue du Général Leclerc, F- 35000 Rennes (France)</mods:affiliation>
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<mods:namePart id="0428511FEBD3BD1E613DCADAE5404F1D">Thomas, Jérôme</mods:namePart>
<mods:affiliation id="09A743B3502B2856E9CB4B9640FBA07A">Biogéosciences, UMR 6282 CNRS, Université Bourgogne, 6 boulevard Gabriel, 21000 Dijon (France)</mods:affiliation>
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<mods:namePart id="128B07FB3C2164A90871D31C8BDA2DE4">Gand, Georges</mods:namePart>
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<mods:date id="20C810FCF1302B140027E7399ED3E028">2024</mods:date>
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<emphasis id="3BEEEACEDE037C01FF7692CADBF2FC78" bold="true" box="[184,391,917,944]" italics="true" pageId="17" pageNumber="358">Grallator maximus</emphasis>
<bibRefCitation id="6D0B4B2DDE037C01FE5492C9D8ABFC78" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[410,734,917,943]" pageId="17" pageNumber="358" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat, 1967</bibRefCitation>
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(
<figureCitation id="91A12A59DE037C01FE6E92E9DB9EFC18" box="[416,491,949,975]" captionStart="FIG" captionStartId="18.[132,143,1395,1412]" captionTargetBox="[173,1432,185,1369]" captionTargetId="figure-247@18.[150,996,234,1387]" captionTargetPageId="18" captionText="FIG. 10. — Grallator minusculus (Hitchcock, 1858), Demathieu, Gand, Sciau &amp; Freytet, 2002: A-C, track ULB-04C13_B (plaster cast of the holotype of “G. maximus” that is here invalidated), photograph (A), DEM and false-colour depth map (B) and interpretative sketch (C); D-F, track ULB-04C10_B, photograph (D), DEM and false-colour depth map (E) and interpretative sketch (F). Scale bars: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.11501878" httpUri="https://zenodo.org/record/11501878/files/figure.png" pageId="17" pageNumber="358">Fig. 10</figureCitation>
)
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<treatmentCitation id="883B10CDDE037C01FF4A92B2D8B5FBD1" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[132,704,1006,1030]" page="22" pageId="17" pageNumber="358" year="1967">
<taxonomicName id="CE9A4D5FDE037C01FF4A92B2D8B5FBD1" authority="Lapparent &amp; Montenat, 1967: 20 - 22" authorityName="&quot; Lapparent &amp; Montenat" authorityPageNumber="20 - 22" authorityYear="1967" box="[132,704,1006,1030]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="17" pageNumber="358" phylum="Chordata" rank="species" species="maximus">
<emphasis id="3BEEEACEDE037C01FF4A92B2DB4EFBD2" box="[132,315,1006,1030]" italics="true" pageId="17" pageNumber="358">Grallator maximus</emphasis>
<bibRefCitation id="6D0B4B2DDE037C01FE8D92B2D8B5FBD1" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[323,704,1006,1030]" pageId="17" pageNumber="358" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat, 1967: 20-22</bibRefCitation>
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, pls X, XI.1, fig. 11.
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. — Subjective junior synonym of
<taxonomicName id="CE9A4D5FDE037C01FDF49564D8A8FBBD" authority="(Hitchcock, 1858) Demathieu, Gand, Sciau &amp; Freytet, 2002" authorityName="Demathieu, Gand, Sciau &amp; Freytet" authorityYear="2002" baseAuthorityName="Hitchcock" baseAuthorityYear="1858" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="17" pageNumber="358" phylum="Chordata" rank="species" species="minusculus">
<emphasis id="3BEEEACEDE037C01FDF49564D97CFB87" box="[570,777,1080,1104]" italics="true" pageId="17" pageNumber="358">Grallator minusculus</emphasis>
(
<bibRefCitation id="6D0B4B2DDE037C01FF43950EDB40FBBD" author="HITCHCOCK E." box="[141,309,1106,1130]" pageId="17" pageNumber="358" refId="ref17969" refString="HITCHCOCK E. 1858. - Ichnology of New England: a Report on the Sandstone of the Connecticut Valley Especially its Fossil Footmarks, Made to the Government of the Commonwealth of Massachusetts. William White, Boston, 220 p." type="book" year="1858">Hitchcock, 1858</bibRefCitation>
)
<bibRefCitation id="6D0B4B2DDE037C01FE8C950ED8A8FBBD" author="DEMATHIEU G. &amp; GAND G. &amp; SCIAU J. &amp; FREYTET P." box="[322,733,1106,1131]" pageId="17" pageNumber="358" pagination="1 - 143" refId="ref17203" refString="DEMATHIEU G., GAND G., SCIAU J. &amp; FREYTET P. 2002. - Les traces de pas de dinosaures et autres archosaures du Lias inferieur des Grands Causses, Sud de la France. Palaeovertebrata 31: 1 - 143. https: // doi. org / 10.18563 / pv. 31.1 - 4.1 - 143" type="journal article" year="2002">Demathieu, Gand, Sciau &amp; Freytet, 2002</bibRefCitation>
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.
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. —
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<typeStatus id="D621887EDE037C01FEBF95DEDBBBFB4D" box="[369,462,1154,1178]" pageId="17" pageNumber="358" type="holotype">Holotype</typeStatus>
:
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(plaster cast).
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Other specimens:
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;
<specimenCode id="593C9EA7DE037C01FE1B95C0D81DFB63" box="[469,616,1180,1204]" pageId="17" pageNumber="358">ULB-04C06_A</specimenCode>
;
<specimenCode id="593C9EA7DE037C01FDA195C0D977FB63" box="[623,770,1180,1204]" pageId="17" pageNumber="358">ULB-04C10_B</specimenCode>
.
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</paragraph>
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The tracks are tridactyl, longer than wide (L/W = 1.5-(1.6)- 1.7), 26.0-(26.8)-
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long and 15.5-(16.6)-
<quantity id="CE629B39DE037C01FDB7944FD8A5FAF9" box="[633,720,1299,1326]" metricMagnitude="-1" metricUnit="m" metricValue="1.77" pageId="17" pageNumber="358" unit="cm" value="17.7">17.7 cm</quantity>
wide (
<figureCitation id="91A12A59DE037C01FF43946FDAADFA9A" box="[141,216,1331,1357]" captionStart="FIG" captionStartId="18.[132,143,1395,1412]" captionTargetBox="[173,1432,185,1369]" captionTargetId="figure-247@18.[150,996,234,1387]" captionTargetPageId="18" captionText="FIG. 10. — Grallator minusculus (Hitchcock, 1858), Demathieu, Gand, Sciau &amp; Freytet, 2002: A-C, track ULB-04C13_B (plaster cast of the holotype of “G. maximus” that is here invalidated), photograph (A), DEM and false-colour depth map (B) and interpretative sketch (C); D-F, track ULB-04C10_B, photograph (D), DEM and false-colour depth map (E) and interpretative sketch (F). Scale bars: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.11501878" httpUri="https://zenodo.org/record/11501878/files/figure.png" pageId="17" pageNumber="358">Fig. 10</figureCitation>
A-F;
<tableCitation id="44180367DE037C01FEC0946FDB29FA9A" box="[270,348,1331,1357]" captionStart="TABLE" captionStartId="7.[132,143,220,237]" captionTargetPageId="7" captionText="TABLE 2. — Measured values on Hettangian tridactyl theropod tracks from Le Veillon. Abbreviations: L, length of the trace; W, width of the trace; D, length of the free part of digit III; II-IV, divarication angle between digit II and digit IV (all in cm except II-IV in degrees)." httpUri="http://table.plazi.org/id/5DE56654DE157C17FF4A9180D999FED4" pageId="17" pageNumber="358" tableUuid="5DE56654DE157C17FF4A9180D999FED4">Table 2</tableCitation>
). D is quite long (L/D = 2.9-(3.0)-3.2). Impressions of digits are large, separated, well defined and elongated. The traces of digits II and III are the smallest and the longest, respectively. The angle between digits II and IV is 30°-(33°)-37° (
<tableCitation id="44180367DE037C01FE8B94EFDBEDFA1A" box="[325,408,1459,1485]" captionStart="TABLE" captionStartId="7.[132,143,220,237]" captionTargetPageId="7" captionText="TABLE 2. — Measured values on Hettangian tridactyl theropod tracks from Le Veillon. Abbreviations: L, length of the trace; W, width of the trace; D, length of the free part of digit III; II-IV, divarication angle between digit II and digit IV (all in cm except II-IV in degrees)." httpUri="http://table.plazi.org/id/5DE56654DE157C17FF4A9180D999FED4" pageId="17" pageNumber="358" tableUuid="5DE56654DE157C17FF4A9180D999FED4">Table 2</tableCitation>
). Impressions of digital pads are well preserved and are circular to oval. The position of the digito-metatarsal pad of digit IV is more proximal than that of digit II (
<figureCitation id="91A12A59DE037C01FF33974EDB3DF9FA" box="[253,328,1554,1581]" captionStart="FIG" captionStartId="18.[132,143,1395,1412]" captionTargetBox="[173,1432,185,1369]" captionTargetId="figure-247@18.[150,996,234,1387]" captionTargetPageId="18" captionText="FIG. 10. — Grallator minusculus (Hitchcock, 1858), Demathieu, Gand, Sciau &amp; Freytet, 2002: A-C, track ULB-04C13_B (plaster cast of the holotype of “G. maximus” that is here invalidated), photograph (A), DEM and false-colour depth map (B) and interpretative sketch (C); D-F, track ULB-04C10_B, photograph (D), DEM and false-colour depth map (E) and interpretative sketch (F). Scale bars: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.11501878" httpUri="https://zenodo.org/record/11501878/files/figure.png" pageId="17" pageNumber="358">Fig. 10</figureCitation>
A-F).
</paragraph>
</subSubSection>
<subSubSection id="41806557DE037C01FF4A970FDF3DFB79" pageId="17" pageNumber="358" type="discussion">
<paragraph id="092536DCDE037C01FF4A970FDA9DF9BC" blockId="17.[130,777,1619,2028]" box="[132,232,1619,1645]" pageId="17" pageNumber="358">
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</paragraph>
<paragraph id="092536DCDE037C01FF4A972EDF3DFB79" blockId="17.[130,777,1619,2028]" lastBlockId="17.[812,1457,853,1198]" pageId="17" pageNumber="358">
Based on this material,
<bibRefCitation id="6D0B4B2DDE037C01FEBC972FD8C3F95B" author="LAPPARENT A. F. DE &amp; MONTENAT C." box="[370,694,1650,1677]" pageId="17" pageNumber="358" pagination="1 - 43" refId="ref18124" refString="LAPPARENT A. F. DE &amp; MONTENAT C. 1967. - Les empreintes de pas de reptiles de l'Infralias du Veillon (Vendee). Memoires de la Societe geologique de France 46: 1 - 43." type="journal article" year="1967">Lapparent &amp; Montenat (1967)</bibRefCitation>
erected (without diagnosis) the ichnospecies “
<taxonomicName id="CE9A4D5FDE037C01FDE597CED881F97B" authorityName="&quot; Lapparent &amp; Montenat" authorityYear="1967" box="[555,756,1682,1708]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="17" pageNumber="358" phylum="Chordata" rank="species" species="maximus">
<emphasis id="3BEEEACEDE037C01FDE597CED881F97B" box="[555,756,1682,1708]" italics="true" pageId="17" pageNumber="358">Grallator maximus</emphasis>
</taxonomicName>
”. These tracks being similar to
<taxonomicName id="CE9A4D5FDE037C01FE7597EED83AF91B" authorityName="Demathieu, Gand, Sciau &amp; Freytet" authorityYear="2002" baseAuthorityName="Hitchcock" baseAuthorityYear="1858" box="[443,591,1714,1740]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="17" pageNumber="358" phylum="Chordata" rank="species" species="minusculus">
<emphasis id="3BEEEACEDE037C01FE7597EED83AF91B" box="[443,591,1714,1740]" italics="true" pageId="17" pageNumber="358">G. minusculus</emphasis>
</taxonomicName>
(
<figureCitation id="91A12A59DE037C01FD9197EED8E9F91B" box="[607,668,1714,1740]" captionStart="FIG" captionStartId="10.[132,143,1963,1980]" captionTargetBox="[237,1380,197,1889]" captionTargetId="graphics-18@10.[311,1344,1074,1820]" captionTargetPageId="10" captionText="FIG. 4. — Footprints from Le Veillon compared to tridactyl tracks from the Early Jurassic of the Causses Basin (based on data from Demathieu et al. 2002; Moreau et al. 2021) and the Early Jurassic of the eastern United States (based on data from Weems 1992, 2019; Gand et al. 2018): A, bivariate diagram L vs (LD)/D; B, bivariate diagram L vs L/W. L in metres." figureDoi="http://doi.org/10.5281/zenodo.11501858" httpUri="https://zenodo.org/record/11501858/files/figure.png" pageId="17" pageNumber="358">Fig. 4</figureCitation>
), we consider “
<taxonomicName id="CE9A4D5FDE037C01FF04978EDB2DF93B" authorityName="Lapparent &amp; Montenat" authorityYear="1967" box="[202,344,1746,1772]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="17" pageNumber="358" phylum="Chordata" rank="species" species="maximum">
<emphasis id="3BEEEACEDE037C01FF04978EDB2DF93B" box="[202,344,1746,1772]" italics="true" pageId="17" pageNumber="358">G. maximum</emphasis>
</taxonomicName>
” as a subjective junior synonym of this ichnospecies. Based on footprints from the Jurassic of the
<collectingCountry id="718D764CDE037C01FF48964EDB69F8FB" box="[134,284,1810,1836]" name="United States of America" pageId="17" pageNumber="358">United States</collectingCountry>
,
<bibRefCitation id="6D0B4B2DDE037C01FEE4964EDB9AF8FB" author="HITCHCOCK E." box="[298,495,1809,1836]" pageId="17" pageNumber="358" refId="ref17969" refString="HITCHCOCK E. 1858. - Ichnology of New England: a Report on the Sandstone of the Connecticut Valley Especially its Fossil Footmarks, Made to the Government of the Commonwealth of Massachusetts. William White, Boston, 220 p." type="book" year="1858">Hitchcock (1858)</bibRefCitation>
first described such large tridactyl tracks under the name
<taxonomicName id="CE9A4D5FDE037C01FE39966EDB35F8BB" authority="Hitchcock, 1858" authorityName="Hitchcock" authorityYear="1858" family="Grallatoridae" genus="Brontozoum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="358" phylum="Chordata" rank="species" species="minusculum">
<emphasis id="3BEEEACEDE037C01FE39966ED97CF89C" box="[503,777,1841,1867]" italics="true" pageId="17" pageNumber="358">Brontozoum minusculum</emphasis>
<bibRefCitation id="6D0B4B2DDE037C01FF4A960DDB35F8BB" author="HITCHCOCK E." box="[132,320,1873,1900]" pageId="17" pageNumber="358" refId="ref17969" refString="HITCHCOCK E. 1858. - Ichnology of New England: a Report on the Sandstone of the Connecticut Valley Especially its Fossil Footmarks, Made to the Government of the Commonwealth of Massachusetts. William White, Boston, 220 p." type="book" year="1858">Hitchcock, 1858</bibRefCitation>
</taxonomicName>
that were renamed
<taxonomicName id="CE9A4D5FDE037C01FDEC960DDB45F85C" authority="Lull, 1904" authorityName="Lull" authorityYear="1904" class="Reptilia" family="Grallatoridae" genus="Anchisauripus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="17" pageNumber="358" phylum="Chordata" rank="species" species="minusculus">
<emphasis id="3BEEEACEDE037C01FDEC960DDAC2F85C" italics="true" pageId="17" pageNumber="358">Anchisauripus minusculus</emphasis>
<bibRefCitation id="6D0B4B2DDE037C01FF71962DDB45F85C" author="LULL R. S." box="[191,304,1905,1931]" pageId="17" pageNumber="358" pagination="461 - 557" refId="ref18537" refString="LULL R. S. 1904. - Fossil footprints of the Jura-Trias of North America. Memoirs of the Boston Society of Natural History 5: 461 - 557." type="journal article" year="1904">Lull, 1904</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="6D0B4B2DDE037C01FE96962DDBA1F85C" author="LULL R. S." box="[344,468,1905,1931]" pageId="17" pageNumber="358" pagination="461 - 557" refId="ref18537" refString="LULL R. S. 1904. - Fossil footprints of the Jura-Trias of North America. Memoirs of the Boston Society of Natural History 5: 461 - 557." type="journal article" year="1904">Lull (1904)</bibRefCitation>
. Based on material from the Hettangian-Sinemurian of the Causses Basin,
<bibRefCitation id="6D0B4B2DDE037C01FD4296CDDABBF81C" author="DEMATHIEU G." pageId="17" pageNumber="358" pagination="229 - 252" refId="ref16998" refString="DEMATHIEU G. 1993. - Empreintes de pas de dinosaures dans les Causses (France). Zubia 5: 229 - 252." type="journal article" year="1993">Demathieu (1993)</bibRefCitation>
identified tracks that show similarities with the
<typeStatus id="D621887EDE037C01FD1796EED972F81B" box="[729,775,1970,1996]" pageId="17" pageNumber="358">type</typeStatus>
material of
<taxonomicName id="CE9A4D5FDE037C01FECC968DD878F83C" authorityName="Lull" authorityYear="1904" box="[258,525,2001,2027]" class="Reptilia" family="Grallatoridae" genus="Anchisauripus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="17" pageNumber="358" phylum="Chordata" rank="species" species="minusculus">
<emphasis id="3BEEEACEDE037C01FECC968DD878F83C" box="[258,525,2001,2027]" italics="true" pageId="17" pageNumber="358">Anchisauripus minusculus</emphasis>
</taxonomicName>
. Since
<taxonomicName id="CE9A4D5FDE037C01FD90968DD885F83C" baseAuthorityName="Lull" baseAuthorityYear="1904" box="[606,752,2001,2027]" class="Reptilia" family="Grallatoridae" genus="Anchisauripus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="17" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="3BEEEACEDE037C01FD90968DD885F83C" box="[606,752,2001,2027]" italics="true" pageId="17" pageNumber="358">Anchisauripus</emphasis>
</taxonomicName>
is characterised by the occasional presence of a hallux trace (
<bibRefCitation id="6D0B4B2DDE037C01FA4A9209D913FC58" author="LULL R. S." pageId="17" pageNumber="358" pagination="461 - 557" refId="ref18537" refString="LULL R. S. 1904. - Fossil footprints of the Jura-Trias of North America. Memoirs of the Boston Society of Natural History 5: 461 - 557." type="journal article" year="1904">Lull 1904</bibRefCitation>
), which is always absent in the material from
<collectingCountry id="718D764CDE037C01FAAF9229DFD9FC58" box="[1377,1452,885,911]" name="France" pageId="17" pageNumber="358">France</collectingCountry>
,
<bibRefCitation id="6D0B4B2DDE037C01FCE092C9D98CFC78" author="DEMATHIEU G." box="[814,1017,917,943]" pageId="17" pageNumber="358" pagination="229 - 252" refId="ref16998" refString="DEMATHIEU G. 1993. - Empreintes de pas de dinosaures dans les Causses (France). Zubia 5: 229 - 252." type="journal article" year="1993">Demathieu (1993)</bibRefCitation>
,
<bibRefCitation id="6D0B4B2DDE037C01FBC692C9DF43FC78" author="DEMATHIEU G. &amp; SCIAU J." box="[1032,1334,917,943]" pageId="17" pageNumber="358" pagination="1561 - 1566" refId="ref17084" refString="DEMATHIEU G. &amp; SCIAU J. 1992. - Des pistes de dinosaures et de crocodiliens dans les dolomies de l'Hettangien du Causse du Larzac. Comptes rendus hebdomadaires des seances de l'Academie des sciences 315: 1561 - 1566. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 6296920 w / f 663. item" type="journal article" year="1992">Demathieu &amp; Sciau (1992)</bibRefCitation>
and
<bibRefCitation id="6D0B4B2DDE037C01FABF92C9DE70FC18" author="DEMATHIEU G. &amp; GAND G. &amp; SCIAU J. &amp; FREYTET P." pageId="17" pageNumber="358" pagination="1 - 143" refId="ref17203" refString="DEMATHIEU G., GAND G., SCIAU J. &amp; FREYTET P. 2002. - Les traces de pas de dinosaures et autres archosaures du Lias inferieur des Grands Causses, Sud de la France. Palaeovertebrata 31: 1 - 143. https: // doi. org / 10.18563 / pv. 31.1 - 4.1 - 143" type="journal article" year="2002">
Demathieu
<emphasis id="3BEEEACEDE037C01FCB492E9D9C4FC18" box="[890,945,949,975]" italics="true" pageId="17" pageNumber="358">et al.</emphasis>
(2002)
</bibRefCitation>
used
<taxonomicName id="CE9A4D5FDE037C01FB8592E9DF59FC18" authorityName="Demathieu, Gand, Sciau &amp; Freytet" authorityYear="2002" baseAuthorityName="Hitchcock" baseAuthorityYear="1858" box="[1099,1324,949,975]" class="Reptilia" family="Grallatoridae" genus="Grallator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="17" pageNumber="358" phylum="Chordata" rank="species" species="minusculus">
<emphasis id="3BEEEACEDE037C01FB8592E9DF59FC18" box="[1099,1324,949,975]" italics="true" pageId="17" pageNumber="358">Grallator minusculus</emphasis>
</taxonomicName>
rather than
<taxonomicName id="CE9A4D5FDE037C01FCE19288DE42FC39" authorityName="Lull" authorityYear="1904" box="[815,1079,980,1006]" class="Reptilia" family="Grallatoridae" genus="Anchisauripus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="17" pageNumber="358" phylum="Chordata" rank="species" species="minusculus">
<emphasis id="3BEEEACEDE037C01FCE19288DE42FC39" box="[815,1079,980,1006]" italics="true" pageId="17" pageNumber="358">Anchisauripus minusculus</emphasis>
</taxonomicName>
.
<bibRefCitation id="6D0B4B2DDE037C01FB8A9288DF32FC38" author="DEMATHIEU G. &amp; GAND G. &amp; SCIAU J. &amp; FREYTET P." box="[1092,1351,980,1007]" pageId="17" pageNumber="358" pagination="1 - 143" refId="ref17203" refString="DEMATHIEU G., GAND G., SCIAU J. &amp; FREYTET P. 2002. - Les traces de pas de dinosaures et autres archosaures du Lias inferieur des Grands Causses, Sud de la France. Palaeovertebrata 31: 1 - 143. https: // doi. org / 10.18563 / pv. 31.1 - 4.1 - 143" type="journal article" year="2002">
Demathieu
<emphasis id="3BEEEACEDE037C01FB0A9289DE82FC39" box="[1220,1271,980,1006]" italics="true" pageId="17" pageNumber="358">et al.</emphasis>
(2002)
</bibRefCitation>
emended the diagnosis of
<taxonomicName id="CE9A4D5FDE037C01FC2792A9DEF6FBD9" authorityName="Lull" authorityYear="1904" box="[1001,1155,1012,1039]" class="Reptilia" family="Grallatoridae" genus="Anchisauripus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="17" pageNumber="358" phylum="Chordata" rank="species" species="minusculus">
<emphasis id="3BEEEACEDE037C01FC2792A9DEF6FBD9" box="[1001,1155,1012,1039]" italics="true" pageId="17" pageNumber="358">G. minusculus</emphasis>
</taxonomicName>
as follows: “
<emphasis id="3BEEEACEDE037C01FADD92A9DE50FB79" italics="true" pageId="17" pageNumber="358">
Large tridactyl tracks II-IV of bipeds with L × l = ca 300 ×
<quantity id="CE629B39DE037C01FADB9548DF01FBF9" box="[1301,1396,1044,1070]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="358" unit="mm" value="200.0">200 mm</quantity>
, with large digits and well-marked pads. Claws are weakly developed and the digito-metatarsal pad of digit IV is often marked. The angle II-IV is 39° in average. The projection of III is low with a III/
<taxonomicName id="CE9A4D5FDE037C01FC9C95C8D999FB79" box="[850,1004,1172,1198]" family="Chirotheriidae" genus="Dahutherium" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="358" phylum="Chordata" rank="subSpecies" species="ratio" subSpecies="around">D ratio around</taxonomicName>
1.93
</emphasis>
” (translated from French).
</paragraph>
</subSubSection>
</treatment>
</document>