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<document id="8F4FF75ACAA2A748F4903E45F3BD15C3" ID-DOI="10.11646/zootaxa.3529.1.1" ID-ISSN="1175­5334" ID-ZooBank="A16A2C8E-8074-4B5C-B097-4C365DBB77C2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629819740303" checkinUser="felipe" docAuthor="Larson, Helen K. &amp; Buckle, Duncan J." docDate="2012" docId="7B14879FFF92E224FF40FE78FC9C071C" docLanguage="en" docName="zt03529p069.pdf" docOrigin="Zootaxa 3529 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3529.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Gnatholepis gymnocara , Randall and Greenfield 2001" docType="treatment" docVersion="2" lastPageNumber="39" masterDocId="872DFFE7FFB4E202FFD7FFE9FF930259" masterDocTitle="A revision of the goby genus Gnatholepis Bleeker (Teleostei, Gobiidae, Gobionellinae), with description of a new species" masterLastPageNumber="69" masterPageNumber="1" pageNumber="39" updateTime="1699116362524" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7726E3D7D6C5BF2DCF39A459452260C3">A revision of the goby genus Gnatholepis Bleeker (Teleostei, Gobiidae, Gobionellinae), with description of a new species</mods:title>
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<subSubSection id="BBA76502FF92E224FF40FE78FF6F0395" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="F3023689FF92E224FF40FE78FCBD03F3" blockId="38.[151,814,400,460]" box="[151,814,400,427]" pageId="38" pageNumber="39">
<heading id="A84A81E5FF92E224FF40FE78FCBD03F3" bold="true" box="[151,814,400,427]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<taxonomicName id="34BD4D0AFF92E224FF40FE78FCBD03F3" ID-CoL="3GLNZ" authority="Randall and Greenfield, 2001" authorityName=", Randall and Greenfield" authorityYear="2001" box="[151,814,400,427]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="gymnocara">
<emphasis id="C1C9EA9BFF92E224FF40FE78FCBD03F3" bold="true" box="[151,814,400,427]" pageId="38" pageNumber="39">
<emphasis id="C1C9EA9BFF92E224FF40FE78FE2B03F2" bold="true" box="[151,440,401,427]" italics="true" pageId="38" pageNumber="39">Gnatholepis gymnocara</emphasis>
<bibRefCitation id="972C4B78FF92E224FE68FE79FCBD03F3" author="Randall and Greenfield" box="[447,814,400,426]" firstAuthor="Randall" pageId="38" pageNumber="39" pagination="1 - 17" refId="ref45808" refString="Randall, J. E. &amp; Greenfield, D. W. (2001) A preliminary review of the Indo-Pacific gobiid fishes of the genus Gnatholepis. Ichthyological Bulletin, 69, 1 - 17." type="journal article" year="2001">Randall and Greenfield, 2001</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F3023689FF92E224FF40FE5DFF6F0395" blockId="38.[151,814,400,460]" box="[151,252,436,460]" pageId="38" pageNumber="39">
(
<tableCitation id="BE3F0332FF92E224FF49FE5DFF600395" box="[158,243,436,460]" captionStart="TABLE 5" captionStartId="13.[151,239,152,176]" captionText="TABLE 5. Frequency distribution of fin ray counts in Gnatholepis species (G. yoshinoi not included). Data for G. gymnocara is from Randall and Greenfield (2001) and pectoral ray data for G. pascuensis is combined with that in Randall (2009); for both these species, the pectoral ray counts are halved (as these authors included both sides in their tables). Data for G. cauerensis is split as per Randall (2009) species for comparison purposes." pageId="38" pageNumber="39">Table 5</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BBA76502FF92E224FF40FE12FA0F0069" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="F3023689FF92E224FF40FE12FD050048" blockId="38.[151,1436,507,560]" box="[151,662,507,530]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="D3AD11A7FF92E224FF40FE12FD050048" box="[151,662,507,530]" pageId="38" pageNumber="39">
?
<treatmentCitation id="721C1098FF92E224FF75FE15FD000048" author="De Vis, C. W." box="[162,659,507,530]" page="687" pageId="38" pageNumber="39" year="1884">
<taxonomicName id="34BD4D0AFF92E224FF75FE15FD000048" authority="de Vis, 1884: 687 (Moreton Bay)" authorityName="De Vis" authorityYear="1884" box="[162,659,507,530]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="pauper">
<emphasis id="C1C9EA9BFF92E224FF75FE15FEA90048" box="[162,314,508,530]" italics="true" pageId="38" pageNumber="39">Gobius pauper</emphasis>
<bibRefCitation id="972C4B78FF92E224FEE8FE12FE610048" author="de Vis" box="[319,498,507,529]" firstAuthor="de Vis" page="687" pageId="38" pageNumber="39" pagination="685 - 698" refId="ref42401" refString="De Vis, C. W. (1884) New Australian fishes in the Queensland Museum. No. 4. Proceedings of the Linnean Society of New South Wales, 9 (3), 685 - 698." type="journal article" year="1884">de Vis, 1884: 687</bibRefCitation>
(Moreton Bay)
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F3023689FF92E224FF40FDF3FA0F0069" blockId="38.[151,1436,507,560]" box="[151,1436,538,560]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="D3AD11A7FF92E224FF40FDF3FA0F0069" box="[151,1436,538,560]" pageId="38" pageNumber="39">
<treatmentCitation id="721C1098FF92E224FF40FDF3FD700069" author="Randall, J. E. &amp; Greenfield, D. W." box="[151,739,538,560]" page="14" pageId="38" pageNumber="39" year="2001">
<taxonomicName id="34BD4D0AFF92E224FF40FDF3FD700069" ID-CoL="3GLNZ" authority="Randall and Greenfield, 2001: 14" authorityName=", Randall and Greenfield" authorityPageNumber="14" authorityYear="2001" box="[151,739,538,560]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="gymnocara">
<emphasis id="C1C9EA9BFF92E224FF40FDF3FE180069" box="[151,395,538,560]" italics="true" pageId="38" pageNumber="39">Gnatholepis gymnocara</emphasis>
<materialsCitation id="43D53CD4FF92E224FE46FDF3FD700069" box="[401,739,538,560]" collectorName="Townshend I." latitude="-22.204" location="Supply Bay" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Queensland">
<bibRefCitation id="972C4B78FF92E224FE46FDF3FD700069" author="Randall and Greenfield" box="[401,739,538,560]" firstAuthor="Randall" page="14" pageId="38" pageNumber="39" pagination="1 - 17" refId="ref45808" refString="Randall, J. E. &amp; Greenfield, D. W. (2001) A preliminary review of the Indo-Pacific gobiid fishes of the genus Gnatholepis. Ichthyological Bulletin, 69, 1 - 17." type="journal article" year="2001">Randall and Greenfield, 2001: 14</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(
<location id="F6626052FF92E224FD27FDF3FCF90069" LSID="urn:lsid:plazi:treatment:7B14879FFF92E224FF40FE78FC9C071C:F6626052FF92E224FD27FDF3FCF90069" box="[752,874,538,560]" latitude="-22.204" name="Supply Bay" pageId="38" pageNumber="39" stateProvince="Queensland">Supply Bay</location>
,
<geoCoordinate id="9689504EFF92E224FCA4FDF3FC770069" box="[883,996,538,560]" degrees="22" direction="south" minutes="12.24" orientation="latitude" pageId="38" pageNumber="39" precision="9" value="-22.204">22°12.24'S</geoCoordinate>
, 150°28.53&quot;E,
<collectorName id="5E48535FFF92E224FBA9FDF3FA9A0069" box="[1150,1289,538,560]" pageId="38" pageNumber="39">Townshend I.</collectorName>
,
<collectingRegion id="3179F86BFF92E224FAC2FDF3FA020069" box="[1301,1425,538,560]" country="Australia" name="Queensland" pageId="38" pageNumber="39">Queensland</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBA76502FF92E224FF40FDB4FC660128" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="F3023689FF92E224FF40FDB4FC660128" blockId="38.[151,1436,605,1350]" pageId="38" pageNumber="39">
<emphasis id="C1C9EA9BFF92E224FF40FDB4FE82002F" bold="true" box="[151,273,605,630]" inLineHeading="true" pageId="38" pageNumber="39" reason="1">Diagnosis.</emphasis>
(modified from
<bibRefCitation id="972C4B78FF92E224FE1DFDB4FD6F002C" author="Randall &amp; Greenfield" box="[458,764,605,629]" firstAuthor="Randall" pageId="38" pageNumber="39" pagination="1 - 17" refId="ref45808" refString="Randall, J. E. &amp; Greenfield, D. W. (2001) A preliminary review of the Indo-Pacific gobiid fishes of the genus Gnatholepis. Ichthyological Bulletin, 69, 1 - 17." type="journal article" year="2001">Randall &amp; Greenfield 2001</bibRefCitation>
) A small
<taxonomicName id="34BD4D0AFF92E224FCBDFDB4FC60002C" box="[874,1011,605,629]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="genus">
<emphasis id="C1C9EA9BFF92E224FCBDFDB4FC60002C" box="[874,1011,605,629]" italics="true" pageId="38" pageNumber="39">Gnatholepis</emphasis>
</taxonomicName>
(up to
<quantity id="34459B6CFF92E224FB95FDB4FB36002C" box="[1090,1189,605,629]" metricMagnitude="-2" metricUnit="m" metricValue="3.54" pageId="38" pageNumber="39" unit="mm" value="35.4">35.4 mm</quantity>
SL) lacking scales on side of cheek, opercle and predorsal midline, few cycloid scales present on pectoral fin base and on breast; no distinct flap on end of lower lip (low fold may be present); no distinct canine or enlarged teeth; second dorsal fin rays I,11; anal fin rays I,1112 (usually I,12); pectoral rays 1519 (usually 17); lateral scales 2628, ctenoid; predorsal scales absent from nape midline, scales on side of head reaching to nearly behind eyes; body pale with dusky speckling and mid-lateral row of six dusky blotches and horizontal row of darker spots and short streaks, an irregular cross-shaped dark mark on cheek extending down from eye, pupil-sized brown spot just above mid-point of caudal fin base, males with small black spot on rear part of first dorsal fin.
</paragraph>
</subSubSection>
<subSubSection id="BBA76502FF92E224FF10FC94FC5E01E0" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="F3023689FF92E224FF10FC94FC5E01E0" blockId="38.[151,1436,605,1350]" pageId="38" pageNumber="39">
<emphasis id="C1C9EA9BFF92E224FF10FC94FE2D01CF" bold="true" box="[199,446,893,918]" pageId="38" pageNumber="39">Material Examined.</emphasis>
<materialsCitation id="43D53CD4FF92E224FE1BFC94FC5A01E0" collectingDate="1998-09-16" collectionCode="AMS" collectorName="S. Reader" location="Townshend Island" municipality="Supply Bay" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="95ACAE4CFF92E224FE1BFC94FD9B01CC" box="[460,520,893,917]" pageId="38" pageNumber="39">AMS</collectionCode>
I.34318-032,
<typeStatus id="2C06882BFF92E224FD60FC97FCB101CF" box="[695,802,894,918]" pageId="38" pageNumber="39" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="34BD4D0AFF92E224FC81FC94FBFF01CC" authorityName=", Randall and Greenfield" authorityYear="2001" box="[854,1132,893,917]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="gymnocara">
<emphasis id="C1C9EA9BFF92E224FC81FC94FBFF01CC" box="[854,1132,893,917]" italics="true" pageId="38" pageNumber="39">Gnatholepis gymnocara</emphasis>
</taxonomicName>
, 32(1733),
<collectingMunicipality id="1366ACF3FF92E224FADBFC94FA0B01CF" box="[1292,1432,893,918]" pageId="38" pageNumber="39">Supply Bay</collectingMunicipality>
,
<location id="F6626052FF92E224FF40FC48FEF601E0" LSID="urn:lsid:plazi:treatment:7B14879FFF92E224FF40FE78FC9C071C:F6626052FF92E224FF40FC48FEF601E0" box="[151,357,929,953]" municipality="Supply Bay" name="Townshend Island" pageId="38" pageNumber="39" stateProvince="Queensland">Townshend Island</location>
,
<collectingRegion id="3179F86BFF92E224FEA6FC48FE6B01E0" box="[369,504,929,953]" country="Australia" name="Queensland" pageId="38" pageNumber="39">Queensland</collectingRegion>
,
<collectorName id="5E48535FFF92E224FDD4FC48FDE301E0" box="[515,624,929,953]" pageId="38" pageNumber="39">S. Reader</collectorName>
and party,
<date id="87031049FF92E224FD3BFC48FC5A01E0" box="[748,969,929,953]" pageId="38" pageNumber="39" value="1998-09-16">
<collectingDate id="9747E9A1FF92E224FD3BFC48FC5A01E0" box="[748,969,929,953]" pageId="38" pageNumber="39" value="1998-09-16">16 September 1998</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BBA76502FF92E224FF10FC2CFC9C071C" pageId="38" pageNumber="39" type="discussion">
<paragraph id="F3023689FF92E224FF10FC2CFBFB0634" blockId="38.[151,1436,605,1350]" pageId="38" pageNumber="39">
<emphasis id="C1C9EA9BFF92E224FF10FC2CFEA80184" bold="true" box="[199,315,965,989]" inLineHeading="true" pageId="38" pageNumber="39" reason="1">Remarks.</emphasis>
In their description of
<taxonomicName id="34BD4D0AFF92E224FDEEFC2CFC750184" authority=", Randall and Greenfield" authorityName=", Randall and Greenfield" authorityYear="2001" box="[569,998,965,989]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="gymnocara">
<emphasis id="C1C9EA9BFF92E224FDEEFC2CFD460184" box="[569,725,965,989]" italics="true" pageId="38" pageNumber="39">G. gymnocara</emphasis>
, Randall and Greenfield
</taxonomicName>
included
<specimenCount id="E5BBFD00FF92E224FB84FC2FFACD0184" box="[1107,1374,965,990]" pageId="38" pageNumber="39" type="generic" typeStatus="paratype">two paratype specimens</specimenCount>
from the
<collectingRegion id="3179F86BFF92E224FF16FC00FE030658" box="[193,400,1001,1025]" country="Australia" name="Northern Territory" pageId="38" pageNumber="39">Northern Territory</collectingRegion>
(AMS I.23930-011). As no other specimens of
<taxonomicName id="34BD4D0AFF92E224FC79FC00FBD80658" authorityName=", Randall and Greenfield" authorityYear="2001" box="[942,1099,1001,1025]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="gymnocara">
<emphasis id="C1C9EA9BFF92E224FC79FC00FBD80658" box="[942,1099,1001,1025]" italics="true" pageId="38" pageNumber="39">G. gymnocara</emphasis>
</taxonomicName>
have so far been found in the Territory, these
<typeStatus id="2C06882BFF92E224FE9FFBE7FE20067F" box="[328,435,1038,1062]" pageId="38" pageNumber="39" type="paratype">paratypes</typeStatus>
were examined and were found to be female
<taxonomicName id="34BD4D0AFF92E224FC60FBE4FB85067C" authorityName="Larson and Buckle" authorityYear="2005" box="[951,1046,1037,1061]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="argus">
<emphasis id="C1C9EA9BFF92E224FC60FBE4FB85067C" box="[951,1046,1037,1061]" italics="true" pageId="38" pageNumber="39">G. argus</emphasis>
</taxonomicName>
(the larger specimen had a second dorsal fin count of I,11, but anal ray count of I,11, while the smaller specimen had I,10 second dorsal rays and I,11 anal rays; both have scaleless pectoral fin bases and typical female
<taxonomicName id="34BD4D0AFF92E224FC5FFBBCFC750634" authorityName="Larson and Buckle" authorityYear="2005" box="[904,998,1109,1133]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="argus">
<emphasis id="C1C9EA9BFF92E224FC5FFBBCFC750634" box="[904,998,1109,1133]" italics="true" pageId="38" pageNumber="39">G. argus</emphasis>
</taxonomicName>
colouring).
</paragraph>
<paragraph id="F3023689FF92E224FF10FB90FC9C071C" blockId="38.[151,1436,605,1350]" pageId="38" pageNumber="39">
The species described as
<taxonomicName id="34BD4D0AFF92E224FE3BFB90FCA906C8" authority="De Vis, 1884" authorityName="De Vis" authorityYear="1884" box="[492,826,1145,1169]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="pauper">
<emphasis id="C1C9EA9BFF92E224FE3BFB90FD0606C8" box="[492,661,1145,1169]" italics="true" pageId="38" pageNumber="39">Gobius pauper</emphasis>
<bibRefCitation id="972C4B78FF92E224FD48FB93FCA906C8" author="De Vis" box="[671,826,1145,1169]" firstAuthor="De Vis" pageId="38" pageNumber="39" pagination="685 - 698" refId="ref42401" refString="De Vis, C. W. (1884) New Australian fishes in the Queensland Museum. No. 4. Proceedings of the Linnean Society of New South Wales, 9 (3), 685 - 698." type="journal article" year="1884">De Vis, 1884</bibRefCitation>
</taxonomicName>
, from Moreton Bay,
<collectingRegion id="3179F86BFF92E224FBFAFB90FB2706C8" box="[1069,1204,1145,1169]" country="Australia" name="Queensland" pageId="38" pageNumber="39">Queensland</collectingRegion>
, has no known
<typeStatus id="2C06882BFF92E224FABBFB93FA0F06CB" box="[1388,1436,1146,1170]" pageId="38" pageNumber="39">type</typeStatus>
specimen. It is possible that it may have been deposited in the
<collectingRegion id="3179F86BFF92E224FC80FB74FC4806EC" box="[855,987,1181,1205]" country="Australia" name="Queensland" pageId="38" pageNumber="39">Queensland</collectingRegion>
Museum, but it is not there now and its whereabouts are unknown. From De Vis original description, it appears to be a species of
<taxonomicName id="34BD4D0AFF92E224FB50FB28FA830680" box="[1159,1296,1217,1241]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="genus">
<emphasis id="C1C9EA9BFF92E224FB50FB28FA830680" box="[1159,1296,1217,1241]" italics="true" pageId="38" pageNumber="39">Gnatholepis</emphasis>
</taxonomicName>
, as noted by
<bibRefCitation id="972C4B78FF92E224FF40FB0FFE2706A4" author="Hoese and Larson" box="[151,436,1253,1277]" firstAuthor="Hoese" pageId="38" pageNumber="39" pagination="1612 - 1697" refId="ref43381" refString="Hoese, D. F. &amp; Larson, H. K. (2006) Gobiidae. Gobies. In: Hoese, D. F., Bray, D. J., Paxton, J. R. &amp; Allen, G. R. (Eds), Zoological Catalogue of Australia. Volume 35. Fishes. Part 3 (of 3). ABRS and CSIRO Publishing, Canberra, pp 1612 - 1697." type="book chapter" year="2006">Hoese and Larson (2006)</bibRefCitation>
. De Vis described his species as being without canines, which implies
<taxonomicName id="34BD4D0AFF92E224FB10FB0CFAF006A4" authorityName=", Randall and Greenfield" authorityYear="2001" box="[1223,1379,1253,1277]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="gymnocara">
<emphasis id="C1C9EA9BFF92E224FB10FB0CFAF006A4" box="[1223,1379,1253,1277]" italics="true" pageId="38" pageNumber="39">G. gymnocara</emphasis>
</taxonomicName>
(and not
<taxonomicName id="34BD4D0AFF92E224FF14FAE0FECA0778" box="[195,345,1289,1313]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="cauerensis">
<emphasis id="C1C9EA9BFF92E224FF14FAE0FECA0778" box="[195,345,1289,1313]" italics="true" pageId="38" pageNumber="39">G. cauerensis</emphasis>
</taxonomicName>
), but we are reluctant to synonymise
<taxonomicName id="34BD4D0AFF92E224FCD2FAE0FCE70778" authorityName="De Vis" authorityYear="1884" box="[773,884,1289,1313]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="pauper">
<emphasis id="C1C9EA9BFF92E224FCD2FAE0FCE70778" box="[773,884,1289,1313]" italics="true" pageId="38" pageNumber="39">G. pauper</emphasis>
</taxonomicName>
with
<taxonomicName id="34BD4D0AFF92E224FC61FAE0FBC70778" authorityName=", Randall and Greenfield" authorityYear="2001" box="[950,1108,1289,1313]" class="Actinopterygii" family="Gobiidae" genus="Gnatholepis" kingdom="Animalia" order="Perciformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="gymnocara">
<emphasis id="C1C9EA9BFF92E224FC61FAE0FBC70778" box="[950,1108,1289,1313]" italics="true" pageId="38" pageNumber="39">G. gymnocara</emphasis>
</taxonomicName>
or any other species without more certainty as to the identity of the
<typeStatus id="2C06882BFF92E224FD9EFAC7FDEA071F" box="[585,633,1326,1350]" pageId="38" pageNumber="39">type</typeStatus>
specimen(s).
</paragraph>
</subSubSection>
</treatment>
</document>