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<mods:title id="4571D56E44DCE4E2E006AD2454D3FA49">The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)</mods:title>
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<treatment id="784D878B6005211C8FF103DCFD5A8896" ID-DOI="http://doi.org/10.5281/zenodo.5918589" ID-GBIF-Taxon="161827870" ID-Zenodo-Dep="5918589" LSID="urn:lsid:plazi:treatment:784D878B6005211C8FF103DCFD5A8896" httpUri="http://treatment.plazi.org/id/784D878B6005211C8FF103DCFD5A8896" lastPageNumber="192" pageId="9" pageNumber="192">
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<paragraph id="F05B369D6005211C8FF103DCFD8F8CEB" blockId="9.[151,573,945,1007]" box="[151,573,945,972]" pageId="9" pageNumber="192">
<heading id="AB1381F16005211C8FF103DCFD8F8CEB" bold="true" box="[151,573,945,972]" fontSize="11" level="1" pageId="9" pageNumber="192" reason="1">
<taxonomicName id="37E44D1E6005211C8FF103DCFD8F8CEB" ID-CoL="7CHTP" authority="Wagner, 1993" authorityName="Wagner" authorityYear="1993" box="[151,573,945,972]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="192" phylum="Arthropoda" rank="species" species="sieberti">
<emphasis id="C290EA8F6005211C8FF103DCFD8F8CEB" bold="true" box="[151,573,945,972]" pageId="9" pageNumber="192">
<emphasis id="C290EA8F6005211C8FF103DCFE398CEC" bold="true" box="[151,395,945,972]" italics="true" pageId="9" pageNumber="192">Tonnoiriella sieberti</emphasis>
<bibRefCitation id="94754B6C6005211C8EF503DCFD8F8CEB" author="Wagner, R." box="[403,573,945,971]" pageId="9" pageNumber="192" pagination="401 - 416" refId="ref20909" refString="Wagner, R. (1993) Einige neue europaische Psychodiden. Entomofauna, 14 (24), 401 - 416." type="journal article" year="1993">Wagner, 1993</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="B8FE65166005211C8FF103B8FF4B8CCF" box="[151,249,980,1007]" pageId="9" pageNumber="192" type="description">
<paragraph id="F05B369D6005211C8FF103B8FF4B8CCF" blockId="9.[151,573,945,1007]" box="[151,249,980,1007]" pageId="9" pageNumber="192">
(
<figureCitation id="68DF2A186005211C8FC603B8FF438CCF" box="[160,241,980,1007]" captionStart="FIGURES 1525" captionStartId="10.[151,264,1910,1935]" captionTargetBox="[213,1374,200,1864]" captionTargetId="figure@10.[196,1391,181,1885]" captionTargetPageId="10" captionText="FIGURES 1525: 15 T. sieberti genitalia ventral view; 16 T. ikariae genitalia ventral view; 1721 T. italiae, genitalia ventral view of specimen from northern Italy (17), Sicily (18), aedeagus of specimen from N Italy (19), Sicily, Villa doso (20) and Sicily Castelbuono (21); 22 T. turcica genitalia ventral view; 23 T. rhodesica genitalia ventral view; 24 T syriensis genitalia ventral view, 25 T. syriensis epandrium and epandrial processes oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621123" httpUri="https://zenodo.org/record/3621123/files/figure.png" pageId="9" pageNumber="192">Fig. 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B8FE65166005211C8FF10474FCAB8B11" box="[151,793,1048,1074]" pageId="9" pageNumber="192" type="reference_group">
<paragraph id="F05B369D6005211C8FF10474FCAB8B11" blockId="9.[151,793,1048,1074]" box="[151,793,1048,1074]" pageId="9" pageNumber="192">
<taxonomicName id="37E44D1E6005211C8FF10474FE4E8B11" ID-CoL="7CHTP" authority="Wagner, 1993" authorityName="Wagner" authorityYear="1993" box="[151,508,1049,1074]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="192" phylum="Arthropoda" rank="species" species="sieberti">
<emphasis id="C290EA8F6005211C8FF10474FED48B11" box="[151,358,1049,1073]" italics="true" pageId="9" pageNumber="192">Tonnoiriella sieberti</emphasis>
<bibRefCitation id="94754B6C6005211C8E0A0477FE4E8B11" author="Wagner, R." box="[364,508,1049,1074]" pageId="9" pageNumber="192" pagination="401 - 416" refId="ref20909" refString="Wagner, R. (1993) Einige neue europaische Psychodiden. Entomofauna, 14 (24), 401 - 416." type="journal article" year="1993">Wagner, 1993</bibRefCitation>
</taxonomicName>
: Entomofauna 14 (24): 407.
</paragraph>
</subSubSection>
<subSubSection id="B8FE65166005211C8FF10432FF6F8BBE" pageId="9" pageNumber="192" type="materials_examined">
<paragraph id="F05B369D6005211C8FF10432FF6F8BBE" blockId="9.[151,1437,1119,1974]" pageId="9" pageNumber="192">
<emphasis id="C290EA8F6005211C8FF10432FF4D8B5A" bold="true" box="[151,255,1119,1146]" pageId="9" pageNumber="192">Material</emphasis>
:
<materialsCitation id="408C3CC06005211C8E69040DFCE98B59" ID-GBIF-Occurrence="2556505403" box="[271,859,1119,1146]" collectingDate="1976-04" country="Macedonia" location="North" municipality="Skopje" pageId="9" pageNumber="192" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="E6E2FD146005211C8E69040DFE808B5A" box="[271,306,1120,1146]" pageId="9" pageNumber="192" type="male">1♂</specimenCount>
<typeStatus id="2F5F883F6005211C8E5D0432FE128B59" box="[315,416,1119,1145]" pageId="9" pageNumber="192" type="holotype">holotype</typeStatus>
,
<location id="F53B60466005211C8ECA0432FE5F8B59" LSID="urn:lsid:plazi:treatment:784D878B6005211C8FF103DCFD5A8896:F53B60466005211C8ECA0432FE5F8B59" box="[428,493,1119,1145]" country="Macedonia" municipality="Skopje" name="North" pageId="9" pageNumber="192">North</location>
<collectingCountry id="88F3760D6005211C8E930432FDC68B59" box="[501,628,1119,1145]" name="Macedonia" pageId="9" pageNumber="192">Macedonia</collectingCountry>
,
<collectingMunicipality id="103FACE76005211C8DE70432FD608B59" box="[641,722,1119,1145]" pageId="9" pageNumber="192">Skopje</collectingMunicipality>
,
<date id="845A105D6005211C8DBB0432FCE98B59" box="[733,859,1119,1145]" pageId="9" pageNumber="192" value="1976-04">
<collectingDate id="941EE9B56005211C8DBB0432FCE98B59" box="[733,859,1119,1145]" pageId="9" pageNumber="192" value="1976-04">April 1976</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="408C3CC06005211C8C0E040DFF6B8BBE" ID-GBIF-Occurrence="2556505401" collectingDate="1976-04" collectionCode="RW" collectorName="Siebert" country="Macedonia" location="North" municipality="Skopje" pageId="9" pageNumber="192" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="E6E2FD146005211C8C0E040DFC398B5A" box="[872,907,1120,1146]" pageId="9" pageNumber="192" type="male">1♂</specimenCount>
<typeStatus id="2F5F883F6005211C8CF5040DFC448B5A" box="[915,1014,1120,1146]" pageId="9" pageNumber="192" type="paratype">paratype</typeStatus>
, collected with holotype, leg.
<collectorName id="5D11534B6005211C8A2B0432FA2E8B59" box="[1357,1436,1119,1145]" pageId="9" pageNumber="192">Siebert</collectorName>
(
<collectionCode id="96F5AE586005211C8FF904E9FF638BBE" box="[159,209,1156,1182]" pageId="9" pageNumber="192">RW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B8FE65166005211C8FA104CAFAA68821" pageId="9" pageNumber="192" type="description">
<paragraph id="F05B369D6005211C8FA104CAFBF48A72" blockId="9.[151,1437,1119,1974]" pageId="9" pageNumber="192">
<emphasis id="C290EA8F6005211C8FA104CAFEE38BE2" bold="true" box="[199,337,1191,1218]" pageId="9" pageNumber="192">Description</emphasis>
: Head round, eyebridge of 3 facet rows; distance between the eyes 5 facet diameters; 6 or 7 postocular bristles. Antenna with scape, pedicel and 14 flagellomeres; absolute length 0.044-0.056-0.044-0.048- 0.054-0.048-0.05-0.05-0.046-0.04
<geoCoordinate id="95D0505A6005211C8D700482FDED8A2A" box="[534,607,1263,1290]" degrees="0.038" direction="south" orientation="latitude" pageId="9" pageNumber="192" precision="55" value="-0.038">-0.038</geoCoordinate>
<geoCoordinate id="95D0505A6005211C8D380482FD1A8A2A" box="[606,680,1263,1290]" degrees="0.042" direction="west" orientation="longitude" pageId="9" pageNumber="192" precision="55" value="-0.042">-0.042</geoCoordinate>
-0.03-0.024-
<quantity id="371C9B786005211C8C570482FC508A2A" box="[817,994,1263,1290]" metricMagnitude="-5" metricUnit="m" metricValue="2.4" metricValueMax="2.8" metricValueMin="2.0" pageId="9" pageNumber="192" unit="mm" value="0.024" valueMax="0.028" valueMin="0.02">0.02-0.028 mm</quantity>
; relative length 22-28-22-24-27-24-25- 25-23-20-
<date id="845A105D6005211C8E6A057EFE258A0E" box="[268,407,1299,1326]" pageId="9" pageNumber="192" value="2019-12-21" valueMax="2019-12-15" valueMin="2019-12-21">19-21-15-12</date>
-10-14. Flagellomeres 511 with a pair of digitiform curved ascoids in the distal third. Palpus segments: absolute length 0.06-0.088-
<quantity id="371C9B786005211C8D27055AFD4D8A72" box="[577,767,1335,1362]" metricMagnitude="-4" metricUnit="m" metricValue="1.37" metricValueMax="1.7599999999999998" metricValueMin="0.98" pageId="9" pageNumber="192" unit="mm" value="0.137" valueMax="0.176" valueMin="0.098">0.098-0.176 mm</quantity>
; relative length 30-44-49-88.
</paragraph>
<paragraph id="F05B369D6005211C8FA10536FBF98A56" blockId="9.[151,1437,1119,1974]" box="[199,1099,1371,1398]" pageId="9" pageNumber="192">
Wing translucent, 2.86 times longer than wide; length
<quantity id="371C9B786005211C8C4E0531FC3C8A56" box="[808,910,1372,1398]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="9" pageNumber="192" unit="mm" value="2.18">2.18 mm</quantity>
, width
<quantity id="371C9B786005211C8C860536FBF58A56" box="[992,1095,1371,1398]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="9" pageNumber="192" unit="mm" value="0.76">0.76 mm</quantity>
.
</paragraph>
<paragraph id="F05B369D6005211C8FA10512FC8E8AC1" blockId="9.[151,1437,1119,1974]" pageId="9" pageNumber="192">
Male terminalia (
<figureCitation id="68DF2A186005211C8EEE0512FE6B8ABA" box="[392,473,1407,1434]" captionStart="FIGURES 1525" captionStartId="10.[151,264,1910,1935]" captionTargetBox="[213,1374,200,1864]" captionTargetId="figure@10.[196,1391,181,1885]" captionTargetPageId="10" captionText="FIGURES 1525: 15 T. sieberti genitalia ventral view; 16 T. ikariae genitalia ventral view; 1721 T. italiae, genitalia ventral view of specimen from northern Italy (17), Sicily (18), aedeagus of specimen from N Italy (19), Sicily, Villa doso (20) and Sicily Castelbuono (21); 22 T. turcica genitalia ventral view; 23 T. rhodesica genitalia ventral view; 24 T syriensis genitalia ventral view, 25 T. syriensis epandrium and epandrial processes oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621123" httpUri="https://zenodo.org/record/3621123/files/figure.png" pageId="9" pageNumber="192">Fig. 15</figureCitation>
) with hypandrium broadened in the middle with edges sinuously curved. Gonocoxites approximately
<date id="845A105D6005211C8E5905C9FEE98A9E" box="[319,347,1444,1470]" pageId="9" pageNumber="192">2x</date>
longer than wide, straight; inner edge basally with a group of 57 short setae at about middle with a single long seta; gonostylus curved, as long as gonocoxite.
</paragraph>
<paragraph id="F05B369D6005211C8FA10581FBAF896D" blockId="9.[151,1437,1119,1974]" pageId="9" pageNumber="192">Parameres with a broad ventral bridge, lobes do not reach middle of gonocoxite; ventrally in the midst develops the aedeagus sheath with a racket-shaped brownish mark; above a pair of broad bilobed processes that do not meet in the middle; further dorsal large rhomboid condyles that form the median keel.</paragraph>
<paragraph id="F05B369D6005211C8FA1063AFCC389FD" blockId="9.[151,1437,1119,1974]" pageId="9" pageNumber="192">Basiphallus bilobed, ventral lobe bifid, ventral branch connected to the distiphallus, longer part slightly bent with blunt tip; distiphallus with both ends rounded, the broader end with a basal triangular projection, the smaller end rounded bifid. The dorsal lobe of the basiphallus is distally fixed to the anchor-shaped distiphallus with cross end, one side broad the other side a thin band, no apical foldings.</paragraph>
<paragraph id="F05B369D6005211C8FA1068AFAA68821" blockId="9.[151,1437,1119,1974]" box="[199,1300,1767,1794]" pageId="9" pageNumber="192">Epandrium rhomboid, epandrial processes almost straight, apically with 1012 feathered tenacula.</paragraph>
</subSubSection>
<subSubSection id="B8FE65166005211C8FA10766FBE88805" box="[199,1114,1803,1830]" pageId="9" pageNumber="192" type="distribution">
<paragraph id="F05B369D6005211C8FA10766FBE88805" blockId="9.[151,1437,1119,1974]" box="[199,1114,1803,1830]" pageId="9" pageNumber="192">
<emphasis id="C290EA8F6005211C8FA10766FEEB8806" bold="true" box="[199,345,1803,1830]" pageId="9" pageNumber="192">Distribution</emphasis>
:
<collectingCountry id="88F3760D6005211C8E010766FE718805" box="[359,451,1803,1829]" name="Albania" pageId="9" pageNumber="192">Albania</collectingCountry>
,
<collectingCountry id="88F3760D6005211C8EAB0766FD308805" box="[461,642,1803,1829]" name="Czech Republic" pageId="9" pageNumber="192">Czech Republic</collectingCountry>
,
<collectingCountry id="88F3760D6005211C8DEB0761FD6E8806" box="[653,732,1804,1830]" name="France" pageId="9" pageNumber="192">France</collectingCountry>
,
<collectingCountry id="88F3760D6005211C8D8E0761FC888806" box="[744,826,1804,1830]" name="Greece" pageId="9" pageNumber="192">Greece</collectingCountry>
, North
<collectingCountry id="88F3760D6005211C8CEB0766FBBE8805" box="[909,1036,1803,1829]" name="Macedonia" pageId="9" pageNumber="192">Macedonia</collectingCountry>
,
<collectingCountry id="88F3760D6005211C8B7E0766FBE58805" box="[1048,1111,1803,1829]" name="Syria" pageId="9" pageNumber="192">Syria</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B8FE65166005211C8FA1075DFD5A8896" pageId="9" pageNumber="192" type="discussion">
<paragraph id="F05B369D6005211C8FA1075DFD5A8896" blockId="9.[151,1437,1119,1974]" pageId="9" pageNumber="192">
<emphasis id="C290EA8F6005211C8FA1075DFE86886A" bold="true" box="[199,308,1840,1866]" pageId="9" pageNumber="192">Remarks</emphasis>
:
<taxonomicName id="37E44D1E6005211C8E22075DFE06886A" authorityName="Wagner" authorityYear="1993" box="[324,436,1840,1866]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="192" phylum="Arthropoda" rank="species" species="sieberti">
<emphasis id="C290EA8F6005211C8E22075DFE06886A" box="[324,436,1840,1866]" italics="true" pageId="9" pageNumber="192">T. sieberti</emphasis>
</taxonomicName>
is distinguished from the other group members by the shape of the median hypandrium, the large ventral lobes of the parameres, and the apically rounded ventral distiphallus.
<bibRefCitation id="94754B6C6005211C8B02073EFB47884E" author="Jezek, J." box="[1124,1269,1875,1902]" pageId="9" pageNumber="192" pagination="7 - 18" refId="ref20095" refString="Jezek, J. (1999) Rare and new Palaearctic Tonnoiriella - species (Diptera, Psychodidae). Casopis Narodniho muzea Rada prirodovedna, 168 (1 - 4), 7 - 18." type="journal article" year="1999">Ježek (1999)</bibRefCitation>
mentioned the species from the
<collectingCountry id="88F3760D6005211C8E39071AFDA588B1" box="[351,535,1911,1937]" name="Czech Republic" pageId="9" pageNumber="192">Czech Republic</collectingCountry>
, North
<collectingCountry id="88F3760D6005211C8D17071AFD4288B1" box="[625,752,1911,1937]" name="Macedonia" pageId="9" pageNumber="192">Macedonia</collectingCountry>
,
<collectingCountry id="88F3760D6005211C8D9B071AFCEB88B1" box="[765,857,1911,1937]" name="Albania" pageId="9" pageNumber="192">Albania</collectingCountry>
,
<collectingCountry id="88F3760D6005211C8C010715FC0488B2" box="[871,950,1912,1938]" name="Greece" pageId="9" pageNumber="192">Greece</collectingCountry>
and
<collectingCountry id="88F3760D6005211C8C95071AFB8088B1" box="[1011,1074,1911,1937]" name="Syria" pageId="9" pageNumber="192">Syria</collectingCountry>
. However, it is possible that in
<collectingCountry id="88F3760D6005211C8FF107F1FF548896" box="[151,230,1948,1974]" name="Greece" pageId="9" pageNumber="192">Greece</collectingCountry>
and in particular in
<collectingCountry id="88F3760D6005211C8EAF07F6FDB68895" box="[457,516,1947,1973]" name="Syria" pageId="9" pageNumber="192">Syria</collectingCountry>
other species occur.
</paragraph>
</subSubSection>
</treatment>
</document>