treatments-xml/data/68/70/E7/6870E7405A5651009774107EC7CA6B44.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.955.51983" ID-GBIF-Dataset="45a98afc-af5a-4734-93d1-205e1ac3694b" ID-PMC="PMC7423785" ID-Pensoft-Pub="1313-2970-955-1" ID-Pensoft-UUID="F7018CF19C6158309D57C83A2F5E9478" ID-PubMed="32855592" ID-ZooBank="C8EF4A4B6F054621B9B5AE47FEE7C217" ModsDocID="1313-2970-955-1" checkinTime="1596691600214" checkinUser="pensoft" docAuthor="Grego, Jozef, Mumladze, Levan, Falniowski, Andrzej, Osikowski, Artur, Rysiewska, Aleksandra, Palatov, Dimitry M. &amp; Hofman, Sebastian" docDate="2020" docId="6870E7405A5651009774107EC7CA6B44" docLanguage="en" docName="ZooKeys 955: 1-77" docOrigin="ZooKeys 955" docSource="http://dx.doi.org/10.3897/zookeys.955.51983" docTitle="Caucasogeyeria chrysomallos Grego &amp; Mumladze 2020, sp. nov." docType="treatment" docUuid="5837CAA1-020A-46D1-9168-E1881C232D9F" docUuidSource="ZooBank" docVersion="5" id="F7018CF19C6158309D57C83A2F5E9478" lastPageNumber="1" masterDocId="F7018CF19C6158309D57C83A2F5E9478" masterDocTitle="Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="1" updateTime="1668169198162" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, 97401, Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mumladze, Levan</mods:namePart>
<mods:affiliation>Institute of Zoology, Ilia State University, Kakutsa Cholokashvili Ave 3 / 5, Tbilisi 0162, Georgia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2172-6973</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Falniowski, Andrzej</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Osikowski, Artur</mods:namePart>
<mods:affiliation>Department of Animal Reproduction, Anatomy and Genomics, University of Agriculture in Krakow, al. Mickiewicza 24 / 28, 30 - 059, Krakow, Poland</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6646-2687</mods:nameIdentifier>
<mods:nameIdentifier type="email">a.osikowski@urk.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rysiewska, Aleksandra</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Palatov, Dimitry M.</mods:namePart>
<mods:affiliation>Department of Hydrobiology, Biological Faculty, Moscow State University, 1 - 12 Leninskie Gory, 119991, Moscow, Russian Federation</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hofman, Sebastian</mods:namePart>
<mods:affiliation>Department of Comparative Anatomy, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>955</mods:number>
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<mods:start>1</mods:start>
<mods:end>77</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.955.51983</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.955.51983</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-955-1</mods:identifier>
<mods:identifier type="ZooBank">C8EF4A4B6F054621B9B5AE47FEE7C217</mods:identifier>
<mods:identifier type="Pensoft-UUID">F7018CF19C6158309D57C83A2F5E9478</mods:identifier>
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<treatment ID-GBIF-Taxon="166096066" LSID="urn:lsid:zoobank.org:act:5837CAA1-020A-46D1-9168-E1881C232D9F" httpUri="http://treatment.plazi.org/id/6870E7405A5651009774107EC7CA6B44" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="6870E740-5A56-5100-9774-107EC7CA6B44" authority="Grego &amp; Mumladze" authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria chrysomallos" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysomallos" status="sp. nov.">Caucasogeyeria chrysomallos Grego &amp; Mumladze</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Plate 16" captionStartId="F30" captionText="Plate 16. 1 - 6 Caucasogeyeria chrysomallos sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring 1 holotype 2 - 6 paratypes 7, 8 Caucasogeyeria chrysomallos sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring 9 Caucasogeyeria cf. gloeri, Samegrelo, Mukhuri, Mapeli Cave, paratype. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego." figureDoi="10.3897/zookeys.955.51983.plate16" httpUri="https://binary.pensoft.net/fig/439182" pageId="0" pageNumber="1">Plates 16</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F29" captionText="Figure 14. A-C Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave D C. chrysomallos sp. nov., Kanti, Mapeli Spring, morphology of the penis. Photograph A. Falniowski and A. Rysiewska." figureDoi="10.3897/zookeys.955.51983.figure14" httpUri="https://binary.pensoft.net/fig/439181" pageId="0" pageNumber="1">(1-8); 17(3); 18(1); Fig. 14D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Georgia • Samegrelo, Kanti Village near Mukhuri, Mapeli Spring (სოფელი 
<normalizedToken originalValue="კანტი">კანტი</normalizedToken>
, 
<normalizedToken originalValue="მაპელის">მაპელის</normalizedToken>
<normalizedToken originalValue="წყარო">წყარო</normalizedToken>
); 
<geoCoordinate degrees="42" direction="north" minutes="38" orientation="latitude" precision="15" seconds="23" value="42.639725">42°38'23&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="42" direction="east" minutes="10" orientation="longitude" precision="15" seconds="08" value="42.16889">42°10'08&quot;E</geoCoordinate>
; 290 m a.s.l.
</paragraph>
<caption doi="10.3897/zookeys.955.51983.plate18" httpUri="https://binary.pensoft.net/fig/439184" pageId="0" pageNumber="1" start="Plate 18" startId="F32">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Plate 18.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="1">1</emphasis>
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria chrysomallos" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysomallos">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria chrysomallos</emphasis>
</taxonomicName>
sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, paratype SBMNH 635903 
<emphasis bold="true" pageId="0" pageNumber="1">2</emphasis>
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria pseudocolchis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pseudocolchis">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria pseudocolchis</emphasis>
</taxonomicName>
sp. nov., Samegrelo, Mukhuri, Shisha Spring, paratype SBMNH 635902 
<emphasis bold="true" pageId="0" pageNumber="1">3</emphasis>
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria gloeri" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gloeri">Caucasogeyeria cf. gloeri</taxonomicName>
, Samegrelo, Mukhuri, Mapeli Cave, paratype SBMNH 635901. Scale bars: 1 mm. SEM SBMNH Vanessa Delnavaz.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Georgia • 1 adult, dry; type locality; 12 October 2019; J. Grego, L. Mumladze and G. Bananashvili leg.; ISU FM-T001-H. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Georgia • same as for holotype; ISU FM-T001-P1/12 dry and 24 wet, SBMNH 633099/2 dry, NHMW 113365/1 dry, HNHM 104678/1 dry, NHMUK 20191334/2 dry, NHMB 563964/1 dry, SMF 358925/1 dry, NHMW 113365, ZMH 140630/1 dry, MNHN-IM-2016-7895, ZIN 1/507-2020/1 dry, coll. JG F1414/15 dry and 24 wet, coll. 
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
/1 dry. • same as for holotype; 10 May 2018; J. Grego, L. Mumladze and M. 
<normalizedToken originalValue="Olšavský">Olsavsky</normalizedToken>
leg.; coll. JG F1041/2 dry. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Other material</emphasis>
: Georgia • Samegrelo, Mapeli Cave, Mukhuri, water catchment (above cemetery); 
<geoCoordinate degrees="42" direction="north" minutes="38" orientation="latitude" precision="15" seconds="22" value="42.639446">42°38'22&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="42" direction="east" minutes="11" orientation="longitude" precision="15" seconds="39" value="42.194168">42°11'39&quot;E</geoCoordinate>
; 325 m a.s.l.; 10 May 2018; J. Grego, L. Mumladze and M. 
<normalizedToken originalValue="Olšavský">Olsavsky</normalizedToken>
leg.; coll. JG/3 dry.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
The new species differs from all congeners by its smaller more elongate-conical shell with more numerous whorls combined with a smaller umbilicus. The aperture sinuation shows similarity with 
<taxonomicName lsidName="C. gloeri" pageId="0" pageNumber="1" rank="species" species="gloeri">
<emphasis italics="true" pageId="0" pageNumber="1">C. gloeri</emphasis>
</taxonomicName>
sp. nov., however, the latter has a larger, more robust and less elevated shell shape with the columellar sinuation. 
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria colchis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="colchis">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria colchis</emphasis>
</taxonomicName>
sp. nov. differs from new species by its larger size and more deeply sinuated labral margin. Measurement comparison of 
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria</emphasis>
</taxonomicName>
species is given in Table 
<tableCitation captionStart="Table 8" captionStartId="T8" captionText="Table 8. Measurement comparison of species from genus Caucasogeyeria gen. nov." httpUri="http://table.plazi.org/id/9CCA906293F80800F9D00A1C34855C9A" pageId="0" pageNumber="1" tableUuid="9CCA906293F80800F9D00A1C34855C9A">8</tableCitation>
.
</paragraph>
<caption ID-Table-UUID="9CCA906293F80800F9D00A1C34855C9A" httpUri="http://table.plazi.org/id/9CCA906293F80800F9D00A1C34855C9A" pageId="0" pageNumber="1" start="Table 8" startId="T8">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Table 8.</emphasis>
Measurement comparison of species from genus 
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria</emphasis>
</taxonomicName>
gen. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">
<emphasis italics="true" pageId="0" pageNumber="1">Genus species</emphasis>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">H</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">W</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">BH</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">BW</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">AH</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">AW</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">CA</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">H/W</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">AH / AW</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">W / BW</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">H/BH</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">H/AH</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">W / AW</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">H/(W- BW)</th>
</tr>
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">deg.</th>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Palatov" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria ignidona" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ignidona">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria ignidona</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.10</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.90</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.85</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.70</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">28</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.45</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.21</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.22</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.39</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.88</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.57</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8.00</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria colchis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="colchis">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria colchis</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.80</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.10</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.85</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.65</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">37</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.29</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.31</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.27</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.57</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.12</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6.00</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria colchis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="colchis">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria colchis</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Paratype</emphasis>
Priveli Balda
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.30</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.95</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.90</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.75</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.55</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">31</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.04</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.36</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.44</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.42</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.80</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.36</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.38</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria colchis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="colchis">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria colchis</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Paratype</emphasis>
Nakriduri 2 spring
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.20</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.05</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.90</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.75</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.33</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.33</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.52</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.13</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.00</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5.33</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria pseudocolchis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pseudocolchis">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria pseudocolchis</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
Shisha Spring
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.45</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.95</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.70</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">28</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.26</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.17</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.21</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.16</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.07</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.92</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7.25</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria pseudocolchis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pseudocolchis">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria pseudocolchis</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Paratype</emphasis>
Shisha Spring
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.55</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.20</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.10</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.95</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.76</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.57</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">24</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.29</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.33</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.26</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.41</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.04</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.11</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6.20</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria chrysomallos" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysomallos">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria chrysomallos</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.93</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.21</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.21</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.07</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.84</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.65</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.29</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.13</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.31</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.86</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">13.83</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="7">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Caucasogeyeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caucasogeyeria chrysomallos" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysomallos">
<emphasis italics="true" pageId="0" pageNumber="1">Caucasogeyeria chrysomallos</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Paratype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.86</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.21</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.16</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.98</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.79</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.65</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">28</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.54</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.21</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.24</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.86</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8.00</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.77</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.23</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.98</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.95</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.74</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.63</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">33</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.43</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.29</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.81</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.38</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.96</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6.33</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.77</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.16</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.07</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.93</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.74</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">29</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.52</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.23</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.65</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.38</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.92</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7.60</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.67</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.16</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.05</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.93</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.72</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">33</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.44</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.32</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.92</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7.20</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.77</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.14</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.98</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.88</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.79</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">28</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.55</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.31</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.29</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.81</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.24</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.88</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6.91</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.23</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.98</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.98</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.70</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.65</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.13</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.07</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.26</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.43</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.00</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.89</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5.45</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.58</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.14</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.02</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.93</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.74</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.56</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">28</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.39</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.33</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.23</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.55</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.13</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.04</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7.56</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
: elongate conical with five tumid whorls, a semi-deep suture, a blunt apex and a narrow umbilicus. Height 1.40-1.93 mm. Shell surface smooth, covered by a milky white periostracum, frequently overlaid by thick dark brown-black inorganic precipitate. The expanded aperture irregularly pear shaped. Labral margin with a weak but broad negative sinuation near the body whorl junction, followed by a characteristic inward curved but shallow labral fold. Columellar margin is straight, not sinuated. Protoconch surface regularly pitted, pitting fading out at the nucleus.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Operculum</emphasis>
: light yellow, paucispiral with central nucleus.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Animal body</emphasis>
: without eye spots, not pigmented, whitish translucent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype measurements</emphasis>
: H-1.93 mm; W-1.21 mm; BH-1.21 mm; BW-1.07 mm; AH-0.84 mm; AW-0.65 mm; CA: 30°.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Anatomy</emphasis>
: penis (Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F29" captionText="Figure 14. A-C Caucasogeyeria ignidona sp. nov., Imereti, Kumistavi, Prometheus Cave D C. chrysomallos sp. nov., Kanti, Mapeli Spring, morphology of the penis. Photograph A. Falniowski and A. Rysiewska." figureDoi="10.3897/zookeys.955.51983.figure14" httpUri="https://binary.pensoft.net/fig/439181" pageId="0" pageNumber="1">14D</figureCitation>
) straight, simple, without any outgrowth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Name derived from the Greek name 
<taxonomicName authorityName="Coenen &amp; Gibeaux" authorityYear="2014" class="Insecta" family="Erebidae" genus="Chrysomallos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chrysomallos" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Chrysomallos</emphasis>
</taxonomicName>
, meaning Golden Fleece (symbol of authority and monarchy), which, according to Greek mythology, was held in Colchis. Jason and his crew of Argonauts were sent out on a quest for the Golden Fleece by order of King Pelias.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">Stygobiotic species. Live individuals as well as empty shells were washed out from its subterranean habitat through a small spring in Mapeli emerging near the road in village Kanti. The dense brown-black deposits on most of individuals indicates a subterranean habitat with chemolithotrophic bacteria. The second known population was found in the sediments of a subterranean cave stream inside Mapeli Cave, ca. 30 m from its entrance</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Only known from the type locality and from Mapeli Cave.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="conservation status">
<paragraph pageId="0" pageNumber="1">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="1">The number of known locations (2) is no more than 5 and EOO is smaller than 20 km2. There is no reason to suppose that AOO, EOO, number of locations, number of subpopulations or the number or mature individuals are declining however due to its extremely small EOO we assessed as Vulnerable (VU) D2.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">The population in Mapeli Cave is typical but has a lower spire and fewer whorls. Its taxonomic position will be clarified after the collection of live individuals.</paragraph>
</subSubSection>
</treatment>
</document>