465 lines
56 KiB
XML
465 lines
56 KiB
XML
<document id="184982599C9A9D633CA356772A872D86" ID-CLB-Dataset="28983" ID-DOI="10.11646/zootaxa.4461.2.2" ID-GBIF-Dataset="ea5796e1-de50-47b0-908a-2fbc88e25c98" ID-ISSN="1175-5326" ID-Zenodo-Dep="1460018" ID-ZooBank="F2FE69F6-C518-4E38-A362-09E40AF31520" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539336748549" checkinUser="plazi" docAuthor="Mora-Aguilar, Eder F., Curoe, Daniel, Delgado, Leonardo & Ramírez-Ponce, Andrés" docDate="2018" docId="683E5410FFF404709BA7FD5D8617FD44" docLanguage="en" docName="zootaxa.4461.2.2.pdf" docOrigin="Zootaxa 4461 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chrysina chimalapensis Mora-Aguilar, Curoe, Delgado" docType="treatment" docVersion="7" lastPageNumber="200" masterDocId="94072C68FFF504749B30FF878464FFEC" masterDocTitle="A new Mexican species of Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae), with nomenclatural changes, new records, and a key to the C. quiche species group" masterLastPageNumber="204" masterPageNumber="196" pageNumber="197" updateTime="1698579704713" updateUser="ExternalLinkService">
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<mods:mods id="B1310FE4F8BEFB3E845DC131436FCF03" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7CB1D39C7298934887CFA53C1833CD7A">
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<mods:title id="E9B19F9C964A9CEFC46A65C41D140016">A new Mexican species of Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae), with nomenclatural changes, new records, and a key to the C. quiche species group</mods:title>
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</mods:titleInfo>
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<mods:name id="FAA12D5E5363A073C45D43AAB81E2F66" type="personal">
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<mods:roleTerm id="6A16DFEC724A5C46E95DFC1A0161CE4A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8977A00D507A2F4B9F2DF0B0B05BA68F">Mora-Aguilar, Eder F.</mods:namePart>
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<mods:roleTerm id="672FDD965649908D6BC157882C401AE7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F1E91B8174DA5119E85DBABDE0E1882F">Curoe, Daniel</mods:namePart>
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</mods:name>
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<mods:name id="7685766B066E82DB368D2FC8B94CF0FE" type="personal">
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<mods:roleTerm id="19D9FA27C3AD122FBE380BFEC60EFB06">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9D9B44ACD9EB14A9A101D24D2AF99BE7">Delgado, Leonardo</mods:namePart>
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</mods:name>
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<mods:name id="A2184C400A70E301CFF1F763FF57CB17" type="personal">
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<mods:role id="41223E99488609855DAA5BFE01D4017F">
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<mods:roleTerm id="71FED114B8997DABF21DCDDDCBD5A097">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AAFF19D2BF93F29340B76CCF0A00077A">Ramírez-Ponce, Andrés</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="FC2B8322817D13CAA3171575B5BE3647">text</mods:typeOfResource>
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<mods:title id="D5CFD2DF3F8C49A59BBED508B6746C94">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="005A2B7ADFE1909E244CF00E36E3692B">
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<mods:date id="7A1045A3A012085FA1F25A85828C3A20">2018</mods:date>
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<mods:detail id="569D51B72714CF97E3D973EE008504D3" type="pubDate">
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<mods:number id="6D7A089182E7BFD795FEA250F91226DA">2018-08-21</mods:number>
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<mods:number id="D81C7B4896CADE5BD4F278F705DC52A0">4461</mods:number>
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<mods:detail id="95A407DCB020317E5D081D7BA520BADB" type="issue">
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<mods:number id="C88814943C84043A2312B8D792EDFAFE">2</mods:number>
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<treatment id="683E5410FFF404709BA7FD5D8617FD44" ID-DOI="http://doi.org/10.5281/zenodo.6490263" ID-GBIF-Taxon="148701895" ID-Zenodo-Dep="6490263" LSID="urn:lsid:plazi:treatment:683E5410FFF404709BA7FD5D8617FD44" httpUri="http://treatment.plazi.org/id/683E5410FFF404709BA7FD5D8617FD44" lastPageId="4" lastPageNumber="200" pageId="1" pageNumber="197">
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<subSubSection id="A88DB68DFFF404759BA7FD5D8570FCF9" pageId="1" pageNumber="197" type="nomenclature">
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<paragraph id="E028E506FFF404759BA7FD5D80A8FD18" blockId="1.[151,1228,730,789]" box="[151,1228,730,756]" pageId="1" pageNumber="197">
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<heading id="BB60526AFFF404759BA7FD5D80A8FD18" bold="true" box="[151,1228,730,756]" fontSize="11" level="1" pageId="1" pageNumber="197" reason="1">
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<emphasis id="D2E33914FFF404759BA7FD5D80A8FD18" bold="true" box="[151,1228,730,756]" pageId="1" pageNumber="197">
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<taxonomicName id="27979E85FFF404759BA7FD5D8725FD18" ID-CoL="5YKKG" authority="Mora-Aguilar, Curoe, Delgado" authorityName="Mora-Aguilar, Curoe, Delgado" box="[151,833,730,756]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="chimalapensis">
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<emphasis id="D2E33914FFF404759BA7FD5D85DEFD18" bold="true" box="[151,442,730,756]" italics="true" pageId="1" pageNumber="197">Chrysina chimalapensis</emphasis>
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Mora-Aguilar, Curoe, Delgado
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</taxonomicName>
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, & Ramírez-Ponce, new species
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</emphasis>
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||
</heading>
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</paragraph>
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<paragraph id="E028E506FFF404759BA7FD7B8570FCF9" blockId="1.[151,1228,730,789]" box="[151,276,764,789]" pageId="1" pageNumber="197">
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<figureCitation id="78ACF983FFF404759BA7FD7B8574FCF9" box="[151,272,764,789]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[237,1344,325,1876]" captionTargetId="figure@3.[151,1436,229,1892]" captionTargetPageId="3" captionText="FIGURES 1–10. Morphological characters of Chrysina chimalapensis. 1, Habitus of male; 2, lateral view of male; 3, ventral view of male; 4, head of male; 5, distal view of parameres; 6, ventral view of parameres; 7, lateral view of parameres; 8, habitus of female; 9, head of female; 10, genital plates." httpUri="https://zenodo.org/record/1460020/files/figure.png" pageId="1" pageNumber="197">Figs. 1–10</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="A88DB68DFFF404759BA7FCC387AEFC25" pageId="1" pageNumber="197" type="materials_examined">
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<paragraph id="E028E506FFF404759BA7FCC387AEFC25" blockId="1.[151,1437,836,2014]" pageId="1" pageNumber="197">
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<materialsCitation id="50FFEF5BFFF404759BA7FCC38689FCB1" ID-GBIF-Occurrence="1919493316" box="[151,749,836,861]" country="Mexico" location="MEXICO" pageId="1" pageNumber="197" specimenCount="2" stateProvince="MEXICO" typeStatus="holotype">
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<emphasis id="D2E33914FFF404759BA7FCC3852DFCB1" bold="true" box="[151,329,836,861]" pageId="1" pageNumber="197">
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<typeStatus id="3F2C5BA4FFF404759BA7FCC384B6FCB1" box="[151,210,836,861]" pageId="1" pageNumber="197">Type</typeStatus>
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material.
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</emphasis>
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||
<typeStatus id="3F2C5BA4FFF404759A64FCC385D8FCB1" box="[340,444,836,861]" pageId="1" pageNumber="197" type="holotype">Holotype</typeStatus>
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male labeled: “
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<collectingRegion id="22532BE4FFF404759948FCC28689FCB1" box="[632,749,837,861]" country="Mexico" name="Mexico" pageId="1" pageNumber="197">
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<collectingCountry id="9880A596FFF404759948FCC28689FCB1" box="[632,749,837,861]" name="Mexico" pageId="1" pageNumber="197">MEXICO</collectingCountry>
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</collectingRegion>
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</materialsCitation>
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||
:
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<materialsCitation id="50FFEF5BFFF4047599CCFCC286C7FC6D" ID-GBIF-Occurrence="1919493314" collectingDate="2017-07-24" collectorName="San Miguel Chimalapa & San Antonio & Camino & Reten & la Torre" country="Mexico" latitude="16.6637" location="Oaxaca" pageId="1" pageNumber="197" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
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<collectingRegion id="22532BE4FFF4047599CCFCC28737FCB1" box="[764,851,837,861]" country="Mexico" name="Oaxaca" pageId="1" pageNumber="197">Oaxaca</collectingRegion>
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,
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<collectorName id="4D6280D0FFF404759852FCC28014FCB1" box="[866,1136,836,861]" pageId="1" pageNumber="197">San Miguel Chimalapa</collectorName>
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,
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<collectorName id="4D6280D0FFF404759FB0FCC28170FCB1" box="[1152,1300,836,861]" pageId="1" pageNumber="197">San Antonio</collectorName>
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,
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<collectorName id="4D6280D0FFF404759E14FCC38119FCB1" box="[1316,1405,836,861]" pageId="1" pageNumber="197">Camino</collectorName>
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al
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<collectorName id="4D6280D0FFF404759BA7FCEE84BCFC6D" box="[151,216,873,897]" pageId="1" pageNumber="197">Retén</collectorName>
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(a
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<collectorName id="4D6280D0FFF404759A34FCEE8507FC6D" box="[260,355,873,898]" pageId="1" pageNumber="197">la Torre</collectorName>
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),
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<collectingDate id="846D3A2EFFF404759A4AFCEE866DFC6D" box="[378,521,873,897]" pageId="1" pageNumber="197" value="2017-07-24">
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<date id="9429C3C6FFF404759A4AFCEE866DFC6D" box="[378,521,873,897]" pageId="1" pageNumber="197" value="2017-07-24">24-VII-2017</date>
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</collectingDate>
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,
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<geoCoordinate id="85A383C1FFF404759929FCEE86FAFC6D" box="[537,670,873,898]" direction="north" orientation="latitude" pageId="1" pageNumber="197" precision="5" value="16.6637">16.6637° N</geoCoordinate>
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,
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</materialsCitation>
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||
-
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||
<materialsCitation id="50FFEF5BFFF404759985FCEE85D3FC25" ID-GBIF-Occurrence="1919493323" collectingDate="2017-07-26" collectionCode="BMM" collectorName="Luz & D. Curoe & One" country="Mexico" elevation="1760" location="Luz" longitude="-94.2335" pageId="1" pageNumber="197" specimenCount="1" stateProvince="MEXICO" typeStatus="paratype">
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||
<geoCoordinate id="85A383C1FFF404759985FCEE872FFC6D" box="[693,843,873,898]" direction="west" orientation="longitude" pageId="1" pageNumber="197" precision="5" value="-94.2335">94.2335 ° W</geoCoordinate>
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,
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<elevation id="6BBA0235FFF40475986AFCEE87CEFC6E" box="[858,938,873,898]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="1" pageNumber="197" unit="m" value="1760.0">
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||
<quantity id="276F48E3FFF40475986AFCEE87CEFC6E" box="[858,938,873,898]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="1" pageNumber="197" unit="m" value="1760.0">1760m</quantity>
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</elevation>
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,
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<collectorName id="4D6280D0FFF404759889FCED8781FC6D" box="[953,997,874,897]" pageId="1" pageNumber="197">Luz</collectorName>
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mercurial, 1am,
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<collectionCode id="86867DC3FFF404759F9FFCED8092FC6D" box="[1199,1270,874,897]" country="Germany" httpUri="http://grbio.org/cool/e0n2-eyvk" name="Buergermeister Mueller, Museum" pageId="1" pageNumber="197">BMM</collectionCode>
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,
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<collectorName id="4D6280D0FFF404759E35FCED8116FC6D" box="[1285,1394,873,897]" pageId="1" pageNumber="197">D. Curoe</collectorName>
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, A. Ramirez-P, E. Mora-A. cols”.
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<collectorName id="4D6280D0FFF404759AC0FC0A867BFC49" box="[496,543,909,933]" pageId="1" pageNumber="197">One</collectorName>
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female
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<typeStatus id="3F2C5BA4FFF40475994CFC0986B8FC4A" box="[636,732,910,934]" pageId="1" pageNumber="197" type="paratype">paratype</typeStatus>
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with same data as holotype except: “
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<collectingDate id="846D3A2EFFF404759FB9FC0B817FFC49" box="[1161,1307,908,933]" pageId="1" pageNumber="197" value="2017-07-26">
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||
<date id="9429C3C6FFF404759FB9FC0B817FFC49" box="[1161,1307,908,933]" pageId="1" pageNumber="197" value="2017-07-26">26-VII-2017</date>
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</collectingDate>
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, 1 800,
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<location id="E548B3DDFFF404759E40FC0981F8FC49" LSID="urn:lsid:plazi:treatment:683E5410FFF404709BA7FD5D8617FD44:E548B3DDFFF404759E40FC0981F8FC49" box="[1392,1436,910,933]" longitude="-94.2335" name="Luz" pageId="1" pageNumber="197">Luz</location>
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aditivos metálicos, 4 am”
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</materialsCitation>
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. Holotype and paratype are deposited in IEXA.
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</paragraph>
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</subSubSection>
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<subSubSection id="A88DB68DFFF404769BF7FC528564FC6C" lastPageId="2" lastPageNumber="198" pageId="1" pageNumber="197" type="description">
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||
<paragraph id="E028E506FFF404769BF7FC5284B5FE3C" blockId="1.[151,1437,836,2014]" lastBlockId="2.[151,1437,151,1977]" lastPageId="2" lastPageNumber="198" pageId="1" pageNumber="197">
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<emphasis id="D2E33914FFF404759BF7FC52853DFC02" bold="true" box="[199,345,981,1006]" pageId="1" pageNumber="197">Description.</emphasis>
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||
<typeStatus id="3F2C5BA4FFF404759A51FC5285ADFC02" box="[353,457,981,1006]" pageId="1" pageNumber="197" type="holotype">Holotype</typeStatus>
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||
male (
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||
<figureCitation id="78ACF983FFF404759928FC52861BFC01" box="[536,639,981,1006]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[237,1344,325,1876]" captionTargetId="figure@3.[151,1436,229,1892]" captionTargetPageId="3" captionText="FIGURES 1–10. Morphological characters of Chrysina chimalapensis. 1, Habitus of male; 2, lateral view of male; 3, ventral view of male; 4, head of male; 5, distal view of parameres; 6, ventral view of parameres; 7, lateral view of parameres; 8, habitus of female; 9, head of female; 10, genital plates." httpUri="https://zenodo.org/record/1460020/files/figure.png" pageId="1" pageNumber="197">Figs 1–7</figureCitation>
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||
). Length
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||
<quantity id="276F48E3FFF4047599DCFC528731FC01" box="[748,853,981,1006]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" pageId="1" pageNumber="197" unit="mm" value="26.8">26.8 mm</quantity>
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||
; width at elytral humeri
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||
<quantity id="276F48E3FFF404759F5DFC5280B2FC01" box="[1133,1238,981,1006]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="197" unit="mm" value="12.6">12.6 mm</quantity>
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||
; maximum width (middle of elytra)
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||
<quantity id="276F48E3FFF404759A5BFC7E85B0FBFD" box="[363,468,1017,1041]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="1" pageNumber="197" unit="mm" value="14.2">14.2 mm</quantity>
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||
.
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<emphasis id="D2E33914FFF404759AD1FC7E8646FBFD" box="[481,546,1017,1041]" italics="true" pageId="1" pageNumber="197">Color</emphasis>
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: Dorsum mostly shiny green; external margin and frontal surface of clypeus, labrum, and mandibles yellowish golden; ocular canthi and periocular areas yellowish orange to pink; antennomeres reddish brown with scape yellowish brown dorsally, antennomere 2 with a greenish patch dorsally, club slightly lighter; pronotal disc yellowish green, pronotal lateral margins yellowish pink, anterior and posterior margins yellowish medially; scutellum yellowish green on base, with apical margins edged with yellowish orange and narrow golden band; elytra with epipleuron and external margin yellowish golden, humeral and apical umbones green, elytral suture and striae yellow; propygidium iridescent yellowish green; pygidium green with margins iridescent yellowish golden. Venter yellowish green with golden reflections; sternites with apical margins iridescent yellowish pink; mesometasternal projection iridescent golden at apex. Legs with coxae yellowish green; trochanters iridescent yellowish green; femorae, tibiae dorsally olive green with orange reflections; tarsi golden olive green, tarsomeres 5 with orange reflections; claws reddish brown with green reflections dorsally. Surface entirely microreticulate.
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<emphasis id="D2E33914FFF404759A9AFA028582FA71" box="[426,486,1413,1437]" italics="true" pageId="1" pageNumber="197">Head</emphasis>
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||
: Clypeus subparabolic; lateral margins almost straight and convergent, apex rounded and slightly reflexed at margin; disc slightly concave; surface densely rugopunctate; punctures denser and confluent near apex. Frons slightly convex with sides straight in lateral view; disc surface more sparsely punctate than on clypeal disc. Interocular distance 1.5 times greater than antennal club length. Antennal club longer than previous six antennomeres; antennomere 2 swollen and subcylindrical (
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<figureCitation id="78ACF983FFF404759887F9928798F9C1" box="[951,1020,1557,1582]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[237,1344,325,1876]" captionTargetId="figure@3.[151,1436,229,1892]" captionTargetPageId="3" captionText="FIGURES 1–10. Morphological characters of Chrysina chimalapensis. 1, Habitus of male; 2, lateral view of male; 3, ventral view of male; 4, head of male; 5, distal view of parameres; 6, ventral view of parameres; 7, lateral view of parameres; 8, habitus of female; 9, head of female; 10, genital plates." httpUri="https://zenodo.org/record/1460020/files/figure.png" pageId="1" pageNumber="197">Fig. 4</figureCitation>
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). Labrum slightly concave at middle, and slightly produced and rounded at middle of posterior margin. Mentum subquadrate; apical margin broadly emarginate with two rounded denticles at middle; distally with long and narrow transverse, rugose depression; basally with longitudinal, foveate depression; surface otherwise setigerously punctate; punctures sparse, large, and small, larger on sides; setae long, pale.
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<emphasis id="D2E33914FFF404759958F92186B3F951" box="[616,727,1702,1725]" italics="true" pageId="1" pageNumber="197">Pronotum</emphasis>
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: Punctures on disc finer and sparser than on head becoming denser laterally; pronotal margins completely beaded except behind frons and anterior to scutellum; pronotal width at base equal to 2.3 times interocular distance.
|
||
<emphasis id="D2E33914FFF4047599F5F96A876FF8E9" box="[709,779,1773,1797]" italics="true" pageId="1" pageNumber="197">Elytra</emphasis>
|
||
: Punctate-striate; striae shallow; punctures in striae and intervals small in size, separated by 2–4 puncture diameters; intervals moderately convex; second, fourth, and sixth interstriae almost impunctate on basal two thirds. Epipleuron extending to level of apical margin of the second abdominal sternite. Elytron 3.1 times as long as pronotum; lateral margin almost completely beaded; effaced at apex.
|
||
<emphasis id="D2E33914FFF404759BEBF8FA8522F879" box="[219,326,1917,1941]" italics="true" pageId="1" pageNumber="197">Pygidium</emphasis>
|
||
: Surface convex, strongly so in apical third; disc surface rugopunctate, denser on base and sides; base and sides reticulate; apex impunctate; apical margin and sides with scattered setose punctures; setae short and pale.
|
||
<emphasis id="D2E33914FFF404759BE4F841857DF831" box="[212,281,1990,2013]" italics="true" pageId="1" pageNumber="197">Venter</emphasis>
|
||
: Mesometasternal projection acute and slightly directed away from body (lateral view), apex rounded (ventral view), almost reaching base of prosternal projection (
|
||
<figureCitation id="78ACF983FFF704769894FF108078FF5C" box="[932,1052,151,176]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[237,1344,325,1876]" captionTargetId="figure@3.[151,1436,229,1892]" captionTargetPageId="3" captionText="FIGURES 1–10. Morphological characters of Chrysina chimalapensis. 1, Habitus of male; 2, lateral view of male; 3, ventral view of male; 4, head of male; 5, distal view of parameres; 6, ventral view of parameres; 7, lateral view of parameres; 8, habitus of female; 9, head of female; 10, genital plates." httpUri="https://zenodo.org/record/1460020/files/figure.png" pageId="2" pageNumber="198">Figs. 2–3</figureCitation>
|
||
). Mesoepimeral disc reticulate. Mesosternum and metasternum, metaepisternum, and metaepimeron densely, setigerously punctate; setae dense, long and pale. Apical ventrite concave, with a shallow, transversal depression.
|
||
<emphasis id="D2E33914FFF704769F2CFF668035FF14" box="[1052,1105,225,248]" italics="true" pageId="2" pageNumber="198">Legs</emphasis>
|
||
: Profemora and mesofemora with scattered long, pale setae. Mesotibia with one transversal carina; metatibia with two carinae, basal carina weak; tibial carinae bordered with spiniform setae. Protibia distinctly tridentate, dorsal and ventral surfaces rugopunctate; protarsus and spur articulate in apical fifth. Protarsomeres 3–5 longitudinally striated on their inner side.
|
||
<emphasis id="D2E33914FFF704769BE4FEF7855AFE64" box="[212,318,368,392]" italics="true" pageId="2" pageNumber="198">Genitalia</emphasis>
|
||
: Parameres short, nearly symmetrical, deeply and narrowly notched; each lateral external margin with three subtriangular denticles; color reddish brown with coppery cast; length of genital capsule
|
||
<quantity id="276F48E3FFF704769FCDFE138130FE40" box="[1277,1364,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="2" pageNumber="198" unit="mm" value="8.6">8.6 mm</quantity>
|
||
(
|
||
<figureCitation id="78ACF983FFF704769E54FE1384A1FE3C" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[237,1344,325,1876]" captionTargetId="figure@3.[151,1436,229,1892]" captionTargetPageId="3" captionText="FIGURES 1–10. Morphological characters of Chrysina chimalapensis. 1, Habitus of male; 2, lateral view of male; 3, ventral view of male; 4, head of male; 5, distal view of parameres; 6, ventral view of parameres; 7, lateral view of parameres; 8, habitus of female; 9, head of female; 10, genital plates." httpUri="https://zenodo.org/record/1460020/files/figure.png" pageId="2" pageNumber="198">Figs. 5–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E028E506FFF704769BF7FE5B8564FC6C" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="198">
|
||
<emphasis id="D2E33914FFF704769BF7FE5B855FFE19" bold="true" box="[199,315,476,501]" pageId="2" pageNumber="198">
|
||
<typeStatus id="3F2C5BA4FFF704769BF7FE5B8552FE19" box="[199,310,476,501]" pageId="2" pageNumber="198" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Female (
|
||
<figureCitation id="78ACF983FFF704769A92FE5B8678FE18" box="[418,540,476,501]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[237,1344,325,1876]" captionTargetId="figure@3.[151,1436,229,1892]" captionTargetPageId="3" captionText="FIGURES 1–10. Morphological characters of Chrysina chimalapensis. 1, Habitus of male; 2, lateral view of male; 3, ventral view of male; 4, head of male; 5, distal view of parameres; 6, ventral view of parameres; 7, lateral view of parameres; 8, habitus of female; 9, head of female; 10, genital plates." httpUri="https://zenodo.org/record/1460020/files/figure.png" pageId="2" pageNumber="198">Figs. 8–10</figureCitation>
|
||
). Length
|
||
<quantity id="276F48E3FFF7047699B6FE5B8688FE18" box="[646,748,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="2" pageNumber="198" unit="mm" value="28.4">28.4 mm</quantity>
|
||
; width at elytral humeral umbones
|
||
<quantity id="276F48E3FFF704769F43FE5B80BEFE18" box="[1139,1242,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="2" pageNumber="198" unit="mm" value="12.1">12.1 mm</quantity>
|
||
; maximum width (middle of elytra)
|
||
<quantity id="276F48E3FFF704769A5DFD8785B3FDFB" box="[365,471,512,536]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="2" pageNumber="198" unit="mm" value="14.2">14.2 mm</quantity>
|
||
. Similar to male except in the following respects: Head darker, clypeus longer and subtriangular, apex in frontal view slightly reflexed dorsally, clypeus less convex; interocular distance 2.0 times wider than antennal club length; antennal club subequal in length to previous six antennomeres. Pronotal width at base equal to 2.0 times interocular distance. Elytra 3.0 times as long as pronotum; epipleuron expanded, reaching level of apical margin of second sternite. Last sternite with a narrow transversal depression in medial third, concave near apex. Pygidium with surface more strong declivous laterally; punctate at disc, and sides rugopunctate to reticulated. Tibiae with punctation denser and coarser. Protibia and spur wider and shorter, protarsus and spur articulated in apical fourth. Protarsus slender with only protarsomeres 4 and 5 longitudinally striated. Mesotibia and metatibia wider and shorter. Metatibial spurs wider. Genital plates inverted anchor-like in shape, with setae scattered, short, and pale; each plate with a long and slender posteriorly directed spine, length of plates
|
||
<quantity id="276F48E3FFF704769E74FCC381F9FCB0" box="[1348,1437,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="198" unit="mm" value="3.4">3.4 mm</quantity>
|
||
(
|
||
<figureCitation id="78ACF983FFF704769BAFFCE08497FC6C" box="[159,243,871,896]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[237,1344,325,1876]" captionTargetId="figure@3.[151,1436,229,1892]" captionTargetPageId="3" captionText="FIGURES 1–10. Morphological characters of Chrysina chimalapensis. 1, Habitus of male; 2, lateral view of male; 3, ventral view of male; 4, head of male; 5, distal view of parameres; 6, ventral view of parameres; 7, lateral view of parameres; 8, habitus of female; 9, head of female; 10, genital plates." httpUri="https://zenodo.org/record/1460020/files/figure.png" pageId="2" pageNumber="198">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A88DB68DFFF704769BF7FC0B87A6FBFC" pageId="2" pageNumber="198" type="diagnosis">
|
||
<paragraph id="E028E506FFF704769BF7FC0B87A6FBFC" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="198">
|
||
<emphasis id="D2E33914FFF704769BF7FC0B8525FC49" bold="true" box="[199,321,908,933]" pageId="2" pageNumber="198">Diagnosis.</emphasis>
|
||
<taxonomicName id="27979E85FFF704769A77FC0B8630FC48" box="[327,596,908,932]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="chimalapensis">
|
||
<emphasis id="D2E33914FFF704769A77FC0B8630FC48" box="[327,596,908,932]" italics="true" pageId="2" pageNumber="198">Chrysina chimalapensis</emphasis>
|
||
</taxonomicName>
|
||
is recognized from the remaining species of the
|
||
<taxonomicName id="27979E85FFF704769F40FC0B80BFFC48" box="[1136,1243,908,932]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="quiche">
|
||
<emphasis id="D2E33914FFF704769F40FC0B80E7FC4F" box="[1136,1155,908,931]" italics="true" pageId="2" pageNumber="198">C</emphasis>
|
||
.
|
||
<emphasis id="D2E33914FFF704769FA0FC0B80BFFC48" box="[1168,1243,908,932]" italics="true" pageId="2" pageNumber="198">quiche</emphasis>
|
||
</taxonomicName>
|
||
species group by the following combination of characters: antennomeres 2 subcylindrical without protrusion, basal pronotal bead effaced medially, males with parameres with three denticles on both external margins, and females with sides of plates convergent apically and with anterolateral angles widely rounded.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A88DB68DFFF704769BF7FB9B8712F974" pageId="2" pageNumber="198" type="discussion">
|
||
<paragraph id="E028E506FFF704769BF7FB9B80B6FBB4" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="198">
|
||
<emphasis id="D2E33914FFF704769BF7FB9B855FFBD9" bold="true" box="[199,315,1052,1077]" pageId="2" pageNumber="198">Remarks.</emphasis>
|
||
<taxonomicName id="27979E85FFF704769A73FB9B8637FBD8" box="[323,595,1052,1076]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="chimalapensis">
|
||
<emphasis id="D2E33914FFF704769A73FB9B8637FBD8" box="[323,595,1052,1076]" italics="true" pageId="2" pageNumber="198">Chrysina chimalapensis</emphasis>
|
||
</taxonomicName>
|
||
is placed in the
|
||
<taxonomicName id="27979E85FFF704769821FB9B871AFBD8" box="[785,894,1052,1076]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="quiche">
|
||
<emphasis id="D2E33914FFF704769821FB9B8740FBDF" box="[785,804,1052,1075]" italics="true" pageId="2" pageNumber="198">C</emphasis>
|
||
.
|
||
<emphasis id="D2E33914FFF704769804FB9B871AFBD8" box="[820,894,1052,1076]" italics="true" pageId="2" pageNumber="198">quiche</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<emphasis id="D2E33914FFF704769F04FB9A8017FBD8" box="[1076,1139,1053,1076]" italics="true" pageId="2" pageNumber="198">sensu</emphasis>
|
||
<bibRefCitation id="840698F7FFF704769F4BFB9B8169FBD8" box="[1147,1293,1052,1077]" pageId="2" pageNumber="198" refId="ref5283" refString="Hawks, D. (2001) Taxonomic and nomenclatural changes in Chrysina and a synonymic checklist of species (Scarabaeidae: Rutelinae). Occasional Papers of the Consortium Coleopterorum, 4, 1 - 8." type="journal article">Hawks 2001</bibRefCitation>
|
||
), which also comprises
|
||
<taxonomicName id="27979E85FFF704769A3FFBC78587FBB4" authority="(Moron)" authorityName="Moron" box="[271,483,1088,1112]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="quiche">
|
||
<emphasis id="D2E33914FFF704769A3FFBC7851FFBB4" box="[271,379,1088,1112]" italics="true" pageId="2" pageNumber="198">C. quiche</emphasis>
|
||
(Morón)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="27979E85FFF704769ADEFBC7873DFBB4" authority="(Cano & Moron)" authorityName="Cano & Moron" box="[494,857,1088,1112]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="tecunumani">
|
||
<emphasis id="D2E33914FFF704769ADEFBC786F6FBBB" box="[494,658,1088,1111]" italics="true" pageId="2" pageNumber="198">C. tecunumani</emphasis>
|
||
(Cano & Morón)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="27979E85FFF7047698A4FBC780B6FBB4" authority="Pokorny & Curoe." authorityName="Pokorny & Curoe." box="[916,1234,1087,1112]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="benesi">
|
||
<emphasis id="D2E33914FFF7047698A4FBC78798FBB4" box="[916,1020,1088,1112]" italics="true" pageId="2" pageNumber="198">C. benesi</emphasis>
|
||
Pokorný & Curoe.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="E028E506FFF704769BF7FBE3866DFA71" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="198">
|
||
The
|
||
<taxonomicName id="27979E85FFF704769BC9FBE38500FB90" box="[249,356,1124,1148]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="quiche">
|
||
<emphasis id="D2E33914FFF704769BC9FBE38568FB97" box="[249,268,1124,1147]" italics="true" pageId="2" pageNumber="198">C</emphasis>
|
||
.
|
||
<emphasis id="D2E33914FFF704769A29FBE38500FB90" box="[281,356,1124,1148]" italics="true" pageId="2" pageNumber="198">quiche</emphasis>
|
||
</taxonomicName>
|
||
species group of
|
||
<taxonomicName id="27979E85FFF704769916FBE386E9FB90" authorityName="Kirby" authorityYear="1828" box="[550,653,1124,1148]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D2E33914FFF704769916FBE386E9FB90" box="[550,653,1124,1148]" italics="true" pageId="2" pageNumber="198">Chrysina</emphasis>
|
||
</taxonomicName>
|
||
is morphologically homogeneous and can be distinguished from other
|
||
<taxonomicName id="27979E85FFF704769BA7FB0F849AFB4C" authorityName="Kirby" authorityYear="1828" box="[151,254,1160,1184]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D2E33914FFF704769BA7FB0F849AFB4C" box="[151,254,1160,1184]" italics="true" pageId="2" pageNumber="198">Chrysina</emphasis>
|
||
</taxonomicName>
|
||
groups by the following combination of characters: medium size (
|
||
<quantity id="276F48E3FFF7047698D2FB0F8004FB4C" box="[994,1120,1160,1184]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.8" metricValueMin="2.4" pageId="2" pageNumber="198" unit="mm" value="26.0" valueMax="28.0" valueMin="24.0">24–28 mm</quantity>
|
||
); habitus shiny green except green-gold to pink external margin of clypeus, periocular areas, pronotal margins, apical margins of scutellum, elytral sutural margins and mesometasternal projection; lateral margin and apical calla of elytra green; elytra with shallow but distinct striae and narrow epipleura, reaching level of second abdominal sternite; mesometasternal projection short with apex rounded (ventral view); male genitalia with parameres short, nearly symmetrical and almost straight in lateral view, and with ventral plates of the phallobase widely separated, slender, apically acute and bent or curved toward one another; female genitalia with inferior plates inverted anchor-shaped, with a basolateral denticle on each side.
|
||
</paragraph>
|
||
<paragraph id="E028E506FFF704769BF7FA2F81E8F9BD" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="198">
|
||
<bibRefCitation id="840698F7FFF704769BF7FA2F851EFA2C" box="[199,378,1447,1472]" pageId="2" pageNumber="198" refId="ref5393" refString="Monzon, J. (2006) El genero Chrysina Kirby (Coleoptera: Scarabaeidae) en Guatemala. In: Cano, E. B. (Ed.), Biodiversidad de Guatemala. Uol. 1. Universidad del Valle de Guatemala, Guatemala City, pp. 393 - 401." type="book chapter">Monzón (2006)</bibRefCitation>
|
||
also included
|
||
<taxonomicName id="27979E85FFF704769917FA2F86D5FA2C" baseAuthorityName="Monzón, Cano & Bailey" baseAuthorityYear="1999" box="[551,689,1448,1472]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="schusteri">
|
||
<emphasis id="D2E33914FFF704769917FA2F86D5FA2C" box="[551,689,1448,1472]" italics="true" pageId="2" pageNumber="198">C. schusteri</emphasis>
|
||
</taxonomicName>
|
||
(Monzón, Cano, & Bailey) within the
|
||
<taxonomicName id="27979E85FFF704769F4EFA2F808AFA2C" box="[1150,1262,1448,1472]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="quiche">
|
||
<emphasis id="D2E33914FFF704769F4EFA2F80F5FA53" box="[1150,1169,1448,1471]" italics="true" pageId="2" pageNumber="198">C</emphasis>
|
||
.
|
||
<emphasis id="D2E33914FFF704769F93FA2F808AFA2C" box="[1187,1262,1448,1472]" italics="true" pageId="2" pageNumber="198">quiche</emphasis>
|
||
</taxonomicName>
|
||
species group. Although externally similar to these species,
|
||
<taxonomicName id="27979E85FFF7047699ABFA4B8746FA08" baseAuthorityName="Monzón, Cano & Bailey" baseAuthorityYear="1999" box="[667,802,1484,1508]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="schusteri">
|
||
<emphasis id="D2E33914FFF7047699ABFA4B86CAFA0F" box="[667,686,1484,1507]" italics="true" pageId="2" pageNumber="198">C</emphasis>
|
||
.
|
||
<emphasis id="D2E33914FFF70476998DFA4B8746FA08" box="[701,802,1484,1508]" italics="true" pageId="2" pageNumber="198">schusteri</emphasis>
|
||
</taxonomicName>
|
||
differs from them by its male genitalia with parameres long (more than half the length of basal piece) and slightly curved at the apical half. Also, the ventral plates are adjacent to each other and rounded apically. The female genitalia have the inferior plates T-shaped and with basolateral lobes. Based on these differences, we prefer to exclude
|
||
<taxonomicName id="27979E85FFF7047698B5F9BF806FF9BC" baseAuthorityName="Monzón, Cano & Bailey" baseAuthorityYear="1999" box="[901,1035,1592,1616]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="schusteri">
|
||
<emphasis id="D2E33914FFF7047698B5F9BF806FF9BC" box="[901,1035,1592,1616]" italics="true" pageId="2" pageNumber="198">C. schusteri</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<taxonomicName id="27979E85FFF704769F49F9BF8080F9BC" box="[1145,1252,1592,1616]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="quiche">
|
||
<emphasis id="D2E33914FFF704769F49F9BF80E8F9A3" box="[1145,1164,1592,1615]" italics="true" pageId="2" pageNumber="198">C</emphasis>
|
||
.
|
||
<emphasis id="D2E33914FFF704769FA9F9BF8080F9BC" box="[1177,1252,1592,1616]" italics="true" pageId="2" pageNumber="198">quiche</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
</paragraph>
|
||
<paragraph id="E028E506FFF704769BF7F9DB8712F974" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="198">
|
||
The four species of the
|
||
<taxonomicName id="27979E85FFF704769ADDF9DB863AF998" box="[493,606,1628,1652]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="quiche">
|
||
<emphasis id="D2E33914FFF704769ADDF9DB8664F99F" box="[493,512,1628,1651]" italics="true" pageId="2" pageNumber="198">C</emphasis>
|
||
.
|
||
<emphasis id="D2E33914FFF704769924F9DB863AF998" box="[532,606,1628,1652]" italics="true" pageId="2" pageNumber="198">quiche</emphasis>
|
||
</taxonomicName>
|
||
species group are externally nearly indistinguishable; examination of genitalia is necessary for determination (
|
||
<bibRefCitation id="840698F7FFF70476996DF9F8870EF974" box="[605,874,1663,1688]" pageId="2" pageNumber="198" refId="ref5682" refString="Pokorny, S. & Curoe, D. J. (2012) A new species of Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae) of the quiche group. Folia Heyrovskyana, Series A, 20, 111 - 116" type="journal article">Pokorný & Curoe 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A88DB68DFFF704769BF7F9238021F933" pageId="2" pageNumber="198" type="etymology">
|
||
<paragraph id="E028E506FFF704769BF7F9238021F933" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="198">
|
||
<emphasis id="D2E33914FFF704769BF7F923852AF951" bold="true" box="[199,334,1700,1725]" pageId="2" pageNumber="198">Etymology.</emphasis>
|
||
The specific epithet is derived from Chimalapas, the region of the state of
|
||
<collectingRegion id="22532BE4FFF704769FF6F923817EF950" box="[1222,1306,1700,1724]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="198">Oaxaca</collectingRegion>
|
||
where this species was collected, combined with the Latin suffix –ensis, meaning belonging to.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A88DB68DFFF704709BF7F96B8617FD44" lastPageId="4" lastPageNumber="200" pageId="2" pageNumber="198" type="distribution">
|
||
<paragraph id="E028E506FFF704769BF7F96B8504F855" blockId="2.[151,1437,151,1977]" pageId="2" pageNumber="198">
|
||
<emphasis id="D2E33914FFF704769BF7F96B8504F8E9" bold="true" box="[199,352,1772,1797]" pageId="2" pageNumber="198">Distribution.</emphasis>
|
||
<taxonomicName id="27979E85FFF704769A58F96B8612F8E8" box="[360,630,1772,1796]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="chimalapensis">
|
||
<emphasis id="D2E33914FFF704769A58F96B8612F8E8" box="[360,630,1772,1796]" italics="true" pageId="2" pageNumber="198">Chrysina chimalapensis</emphasis>
|
||
</taxonomicName>
|
||
is currently known to occur only in the cloud forest of El Retén, in the Chimalapas region located at the southeast end of the state of
|
||
<collectingRegion id="22532BE4FFF704769875F89787FDF8C4" box="[837,921,1808,1832]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="198">Oaxaca</collectingRegion>
|
||
State (
|
||
<figureCitation id="78ACF983FFF7047698D5F888805CF8C4" box="[997,1080,1807,1832]" captionStart="FIGURE 17" captionStartId="7.[151,250,1446,1468]" captionTargetBox="[153,1433,360,1405]" captionTargetId="figure@7.[150,1436,339,1424]" captionTargetPageId="7" captionText="FIGURE 17. Geographical distribution of the Chrysina quiche species group. Chrysina benesi (black triangle), C. chimalapensis (grey triangle), C. quiche (black squares), C. tecunumani (open squares). Elevation limits of 1,200, 1,800, and 2,400 m, darker gray lines indicate lower elevation." httpUri="https://zenodo.org/record/1460026/files/figure.png" pageId="2" pageNumber="198">Fig. 17</figureCitation>
|
||
). The forests at the
|
||
<typeStatus id="3F2C5BA4FFF704769E22F8978126F8C4" box="[1298,1346,1808,1832]" pageId="2" pageNumber="198">type</typeStatus>
|
||
locality and neighboring areas (
|
||
<quantity id="276F48E3FFF704769A92F8B38634F8A0" box="[418,592,1844,1869]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.8" metricValueMin="1.45" pageId="2" pageNumber="198" unit="m" value="1625.0" valueMax="1800.0" valueMin="1450.0">1,450–1,800 m</quantity>
|
||
) present a high degree of conservation. The canopy in the cloud forest of Chimalapas is composed of trees of the genera
|
||
<taxonomicName id="27979E85FFF7047699E5F8DF8074F89C" box="[725,1040,1879,1904]" class="Magnoliopsida" family="Euphorbiaceae" genus="Alchornea" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="198" phylum="Magnoliopsida" rank="genus">
|
||
<emphasis id="D2E33914FFF7047699E5F8DF872EF89C" box="[725,842,1880,1904]" italics="true" pageId="2" pageNumber="198">Alchornea</emphasis>
|
||
(Euphorbiaceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="27979E85FFF704769F14F8DF8144F89C" box="[1060,1312,1879,1904]" class="Magnoliopsida" family="Clethraceae" genus="Clethra" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="198" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="D2E33914FFF704769F14F8DF801DF89C" box="[1060,1145,1880,1904]" italics="true" pageId="2" pageNumber="198">Clethra</emphasis>
|
||
(Clethraceae)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="D2E33914FFF704769E01F8DF81F8F89C" box="[1329,1436,1880,1904]" italics="true" pageId="2" pageNumber="198">Coccolba</emphasis>
|
||
(
|
||
<taxonomicName id="27979E85FFF704769BAEF8FB8559F879" authorityName="" authorityYear="1789" box="[158,317,1916,1941]" family="Polygonaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="198" phylum="Magnoliopsida" rank="family">Polygonaceae</taxonomicName>
|
||
),
|
||
<taxonomicName id="27979E85FFF704769A68F8FA8643F878" box="[344,551,1916,1940]" class="Magnoliopsida" family="Moraceae" genus="Ficus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="2" pageNumber="198" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="D2E33914FFF704769A68F8FA85F2F878" box="[344,406,1917,1940]" italics="true" pageId="2" pageNumber="198">Ficus</emphasis>
|
||
(Moraceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="27979E85FFF704769908F8FB877FF878" authorityName="Aublet" authorityYear="1775" box="[568,795,1916,1940]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" higherTaxonomySource="CoL" kingdom="Plantae" order="Laurales" pageId="2" pageNumber="198" phylum="Magnoliopsida" rank="genus">
|
||
<emphasis id="D2E33914FFF704769908F8FB86EDF87F" box="[568,649,1916,1939]" italics="true" pageId="2" pageNumber="198">Ocotea</emphasis>
|
||
(Lauraceae)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="27979E85FFF70476981EF8FA8075F878" box="[814,1041,1916,1940]" class="Magnoliopsida" family="Lauraceae" genus="Persea" higherTaxonomySource="CoL" kingdom="Plantae" order="Laurales" pageId="2" pageNumber="198" phylum="Magnoliopsida" rank="genus">
|
||
<emphasis id="D2E33914FFF70476981EF8FA8718F878" box="[814,892,1917,1940]" italics="true" pageId="2" pageNumber="198">Persea</emphasis>
|
||
(Lauraceae)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="27979E85FFF704769F66F8FB8158F878" box="[1110,1340,1916,1940]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="198" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="D2E33914FFF704769F66F8FB80D0F87F" box="[1110,1204,1916,1939]" italics="true" pageId="2" pageNumber="198">Quercus</emphasis>
|
||
(Fagaceae)
|
||
</taxonomicName>
|
||
(Lorea-
|
||
<bibRefCitation id="840698F7FFF704769BA7F8278530F855" box="[151,340,1952,1977]" pageId="2" pageNumber="198" refId="ref5351" refString="Lorea-Hernandez, F. (2005) Nuevas especies de Licaria, Ocotea y Persea (Lauraceae) de Mexico. Acta Botanica Mexicana, 71, 61 - 87. https: // doi. org / 10.21829 / abm 71.2005.995" type="journal article">Hernández 2005</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B4E8B58EFFF604779BA7F81B8611F81C" httpUri="https://zenodo.org/record/1460020/files/figure.png" pageId="3" pageNumber="199" startId="3.[151,264,1948,1971]" targetBox="[237,1344,325,1876]" targetPageId="3">
|
||
<paragraph id="E028E506FFF604779BA7F81B8611F81C" blockId="3.[151,1436,1948,2032]" pageId="3" pageNumber="199">
|
||
<emphasis id="D2E33914FFF604779BA7F81B8523F85E" bold="true" box="[151,327,1948,1971]" pageId="3" pageNumber="199">FIGURES 1–10.</emphasis>
|
||
Morphological characters of
|
||
<taxonomicName id="27979E85FFF604779947F81A8708F85F" box="[631,876,1949,1971]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="species" species="chimalapensis">
|
||
<emphasis id="D2E33914FFF604779947F81A8708F85F" box="[631,876,1949,1971]" italics="true" pageId="3" pageNumber="199">Chrysina chimalapensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="D2E33914FFF60477984AF81A87E8F85E" bold="true" box="[890,908,1949,1970]" pageId="3" pageNumber="199">1,</emphasis>
|
||
Habitus of male;
|
||
<emphasis id="D2E33914FFF604779F76F81A803CF85E" bold="true" box="[1094,1112,1949,1970]" pageId="3" pageNumber="199">2,</emphasis>
|
||
lateral view of male;
|
||
<emphasis id="D2E33914FFF604779E0BF81A812BF85E" bold="true" box="[1339,1359,1949,1970]" pageId="3" pageNumber="199">3,</emphasis>
|
||
ventral view of male;
|
||
<emphasis id="D2E33914FFF604779A17F83C855DF83C" bold="true" box="[295,313,1979,2000]" pageId="3" pageNumber="199">4,</emphasis>
|
||
head of male;
|
||
<emphasis id="D2E33914FFF604779AFEF83B8585F83D" bold="true" box="[462,481,1980,2001]" pageId="3" pageNumber="199">5,</emphasis>
|
||
distal view of parameres;
|
||
<emphasis id="D2E33914FFF6047799DAF83C869AF83C" bold="true" box="[746,766,1979,2000]" pageId="3" pageNumber="199">6,</emphasis>
|
||
ventral view of parameres;
|
||
<emphasis id="D2E33914FFF604779F25F83B804CF83D" bold="true" box="[1045,1064,1980,2001]" pageId="3" pageNumber="199">7,</emphasis>
|
||
lateral view of parameres;
|
||
<emphasis id="D2E33914FFF604779E0AF83C8129F83C" bold="true" box="[1338,1357,1979,2000]" pageId="3" pageNumber="199">8,</emphasis>
|
||
habitus of female;
|
||
<emphasis id="D2E33914FFF604779A34F85D857CF803" bold="true" box="[260,280,2010,2031]" pageId="3" pageNumber="199">9,</emphasis>
|
||
head of female;
|
||
<emphasis id="D2E33914FFF604779AF2F85D8585F803" bold="true" box="[450,481,2010,2031]" pageId="3" pageNumber="199">10,</emphasis>
|
||
genital plates.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E028E506FFF104709BF7FF108098FDD1" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="200">
|
||
The species of the
|
||
<taxonomicName id="27979E85FFF104709AA7FF1F8667FF5C" box="[407,515,152,176]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="200" phylum="Arthropoda" rank="species" species="quiche">
|
||
<emphasis id="D2E33914FFF104709AA7FF1F85CEFF43" box="[407,426,152,175]" italics="true" pageId="4" pageNumber="200">C</emphasis>
|
||
.
|
||
<emphasis id="D2E33914FFF104709A88FF1F8667FF5C" box="[440,515,152,176]" italics="true" pageId="4" pageNumber="200">quiche</emphasis>
|
||
</taxonomicName>
|
||
species group are restricted to the cloud forests located above
|
||
<quantity id="276F48E3FFF104709F8DFF10817DFF43" box="[1213,1305,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="200" unit="m" value="1600.0">1,600 m</quantity>
|
||
in southern Mexico (Oaxaca and Chiapas), and Guatemala, in the Chiapas Highlands biogeographic province (
|
||
<emphasis id="D2E33914FFF104709FC4FF3A8156FF38" box="[1268,1330,189,212]" italics="true" pageId="4" pageNumber="200">sensu</emphasis>
|
||
<bibRefCitation id="840698F7FFF104709E0AFF3A84B0FF14" pageId="4" pageNumber="200" refId="ref5646" refString="Morrone, J. J. (2014) Biogeographical regionalization of the Neotropical region. Zootaxa, 3782, 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article">Morrone 2014</bibRefCitation>
|
||
). All species, except
|
||
<taxonomicName id="27979E85FFF104709AF8FF678652FF14" box="[456,566,224,248]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="200" phylum="Arthropoda" rank="species" species="quiche">
|
||
<emphasis id="D2E33914FFF104709AF8FF678652FF14" box="[456,566,224,248]" italics="true" pageId="4" pageNumber="200">C. quiche</emphasis>
|
||
</taxonomicName>
|
||
, show isolated distributions in different geographical areas (
|
||
<figureCitation id="78ACF983FFF104709FDCFF588124FF14" box="[1260,1344,223,248]" captionStart="FIGURE 17" captionStartId="7.[151,250,1446,1468]" captionTargetBox="[153,1433,360,1405]" captionTargetId="figure@7.[150,1436,339,1424]" captionTargetPageId="7" captionText="FIGURE 17. Geographical distribution of the Chrysina quiche species group. Chrysina benesi (black triangle), C. chimalapensis (grey triangle), C. quiche (black squares), C. tecunumani (open squares). Elevation limits of 1,200, 1,800, and 2,400 m, darker gray lines indicate lower elevation." httpUri="https://zenodo.org/record/1460026/files/figure.png" pageId="4" pageNumber="200">Fig. 17</figureCitation>
|
||
), which fits the Nuclear Central America areas of endemism of proposed by
|
||
<bibRefCitation id="840698F7FFF1047098A5FE8380EDFEF0" box="[917,1161,260,285]" pageId="4" pageNumber="200" refId="ref5725" refString="Schuster, J. C., Cano, E. B. & Carmona, C. (2000) Un metodo sencillo para priorizar la conservacion de los bosques nubosos de Guatemala, usando Passalidae (Coleoptera) como organismos indicadores. Acta Zoologica Mexicana, New Series, 80, 197 - 209." type="journal article">
|
||
Schuster
|
||
<emphasis id="D2E33914FFF1047098CCFE828048FEF0" box="[1020,1068,260,284]" italics="true" pageId="4" pageNumber="200">et al</emphasis>
|
||
. (2000)
|
||
</bibRefCitation>
|
||
. The distribution area of
|
||
<taxonomicName id="27979E85FFF104709BA7FEAF853CFEAC" box="[151,344,296,320]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="200" phylum="Arthropoda" rank="species" species="chimalapensis">
|
||
<emphasis id="D2E33914FFF104709BA7FEAF853CFEAC" box="[151,344,296,320]" italics="true" pageId="4" pageNumber="200">C. chimalapensis</emphasis>
|
||
</taxonomicName>
|
||
is isolated from the remaining portion of the Sierra Madre de Chiapas in Mexico by areas below
|
||
<quantity id="276F48E3FFF104709BA7FECB8490FE88" box="[151,244,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="200" unit="m" value="1000.0">1,000 m</quantity>
|
||
elevation.
|
||
<taxonomicName id="27979E85FFF104709A41FECB8643FE88" box="[369,551,332,356]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="200" phylum="Arthropoda" rank="species" species="benesi">
|
||
<emphasis id="D2E33914FFF104709A41FECB8643FE88" box="[369,551,332,356]" italics="true" pageId="4" pageNumber="200">Chrysina benesi</emphasis>
|
||
</taxonomicName>
|
||
is known from one locality in the state of Chiapas, located at the southeast of the Sierra Madre de Chiapas, and, in Guatemala, in the north of the Sierra Los Cuchumatanes.
|
||
<taxonomicName id="27979E85FFF104709E05FEF7857EFE40" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="200" phylum="Arthropoda" rank="species" species="tecunumani">
|
||
<emphasis id="D2E33914FFF104709E05FEF7857EFE40" italics="true" pageId="4" pageNumber="200">Chrysina tecunumani</emphasis>
|
||
</taxonomicName>
|
||
is only known from the Sierra de Chama and Sierra de Las Minas in the central region of Guatemala.
|
||
<taxonomicName id="27979E85FFF104709BA7FE3F8530FE3C" box="[151,340,440,464]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="200" phylum="Arthropoda" rank="species" species="quiche">
|
||
<emphasis id="D2E33914FFF104709BA7FE3F8530FE3C" box="[151,340,440,464]" italics="true" pageId="4" pageNumber="200">Chrysina quiche</emphasis>
|
||
</taxonomicName>
|
||
shows the widest distribution, occurring in the Sierra Madre de Chiapas in Mexico, and, in Guatemala, in the Cordillera Volcánica Occidental, the Sierra de Chama, and the Sierra de Las Minas (J. Monzón, Guatemala City, Guatemala, personal communication). The previous record from the eastern region of the Sierra de los Cuchumatanes for
|
||
<taxonomicName id="27979E85FFF104709AA2FDA386D0FDD1" authority="(Monzon 2006)" baseAuthorityName="Monzon" baseAuthorityYear="2006" box="[402,692,548,573]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="200" phylum="Arthropoda" rank="species" species="quiche">
|
||
<emphasis id="D2E33914FFF104709AA2FDA3859AFDD0" box="[402,510,548,572]" italics="true" pageId="4" pageNumber="200">C. quiche</emphasis>
|
||
(
|
||
<bibRefCitation id="840698F7FFF10470993CFDA386C8FDD1" box="[524,684,548,573]" pageId="4" pageNumber="200" refId="ref5393" refString="Monzon, J. (2006) El genero Chrysina Kirby (Coleoptera: Scarabaeidae) en Guatemala. In: Cano, E. B. (Ed.), Biodiversidad de Guatemala. Uol. 1. Universidad del Valle de Guatemala, Guatemala City, pp. 393 - 401." type="book chapter">Monzón 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
is erroneous (J. Monzón, personal communication).
|
||
</paragraph>
|
||
<paragraph id="E028E506FFF104709BF7FDC08617FD44" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="200">
|
||
<emphasis id="D2E33914FFF104709BF7FDC085E2FD8C" bold="true" box="[199,390,583,608]" pageId="4" pageNumber="200">Natural history.</emphasis>
|
||
<taxonomicName id="27979E85FFF104709ABFFDCF86FAFD8C" box="[399,670,584,608]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="200" phylum="Arthropoda" rank="species" species="chimalapensis">
|
||
<emphasis id="D2E33914FFF104709ABFFDCF86FAFD8C" box="[399,670,584,608]" italics="true" pageId="4" pageNumber="200">Chrysina chimalapensis</emphasis>
|
||
</taxonomicName>
|
||
is attracted at night to mercury vapor and metal halide light and is sympatric with
|
||
<taxonomicName id="27979E85FFF104709A76FDEB865FFD68" authority="Moron" authorityName="Moron" authorityYear="1990" box="[326,571,620,644]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="200" phylum="Arthropoda" rank="species" species="triumphalis">
|
||
<emphasis id="D2E33914FFF104709A76FDEB858CFD68" box="[326,488,620,644]" italics="true" pageId="4" pageNumber="200">C. triumphalis</emphasis>
|
||
Morón
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="27979E85FFF104709943FDEB87EAFD68" authority="(Moron)" authorityName="Moron" box="[627,910,620,644]" class="Insecta" family="Scarabaeidae" genus="Chrysina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="200" phylum="Arthropoda" rank="species" species="quetzalcoatli">
|
||
<emphasis id="D2E33914FFF104709943FDEB8740FD68" box="[627,804,620,644]" italics="true" pageId="4" pageNumber="200">C. quetzalcoatli</emphasis>
|
||
(Morón)
|
||
</taxonomicName>
|
||
, with the new species being the least abundant in this six collecting events at this locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |