801 lines
103 KiB
XML
801 lines
103 KiB
XML
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<mods:title id="281B10D0A659B1E44B4DF8AD3DF0E802">Morphology and taxonomic assessment of eight genetic clades of Mercuria Boeters, 1971 (Caenogastropoda, Hydrobiidae), with the description of five new species</mods:title>
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<mods:affiliation id="8FEF21B64A32767D4FD6F8558E5BECD2">Museo Nacional de Ciencias Naturales (MNCN-CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain. Justus Liebig University Giessen, Department of Animal Ecology & Systematics, Heinrich-Buff-Ring 26 - 32 IFZD, 35392 Giessen, Germany. University of Carthage, Faculty of Sciences of Bizerte, LR 01 ES 14 Environmental Biomonitoring Laboratory, 7021 Jarzouna, Tunisia. Deceased 3 March 2023.</mods:affiliation>
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<mods:title id="343FA79D806530BE9CAED479397ADF3D">European Journal of Taxonomy</mods:title>
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<treatment id="5C7987C4FFF4FFE0EF51FDFA314EF9B4" ID-DOI="http://doi.org/10.5281/zenodo.7891731" ID-GBIF-Taxon="209579758" ID-Zenodo-Dep="7891731" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4" httpUri="http://treatment.plazi.org/id/5C7987C4FFF4FFE0EF51FDFA314EF9B4" lastPageId="25" lastPageNumber="26" pageId="20" pageNumber="21">
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<emphasis id="E6A4EAC0FFF4FFEDEF51FDFA360DFD84" bold="true" box="[572,806,611,638]" italics="true" pageId="20" pageNumber="21">Mercuria balearica</emphasis>
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(
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<bibRefCitation id="B0414B23FFF4FFEDEE58FDFA36C4FD85" author="Paladilhe A." box="[821,1007,612,639]" pageId="20" pageNumber="21" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe, 1869</bibRefCitation>
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<figureCitation id="4CEB2A57FFF4FFEDEF4AFD193786FD5B" box="[551,685,647,673]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="21.[189,232,1892,1918]" captionStartId-1="22.[189,232,1892,1918]" captionStartId-2="23.[189,232,1928,1954]" captionStartId-3="24.[189,232,1891,1917]" captionTargetBox-0="[189,1398,274,1842]" captionTargetBox-1="[189,1398,1115,1846]" captionTargetBox-2="[189,1398,1130,1885]" captionTargetBox-3="[189,1398,1087,1848]" captionTargetId-0="figure-11@21.[189,1398,274,1842]" captionTargetId-1="figure-302@22.[189,1398,1115,1846]" captionTargetId-2="figure-328@23.[189,1398,1130,1885]" captionTargetId-3="figure-272@24.[189,1398,1085,1848]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionText-0="Fig. 11. Shells and operculum of Mercuria balearica (Paladilhe, 1869). A–B, G–H. Stream at Sant Joan de Carbonell, Minorca, Spain. C. Colarsega River, near the Port of Mahon (type locality). D. Fuente Árabe Spring, Mojácar, Granada, Spain (unpigmented specimen). E. Venta El Pilar Spring, Málaga, Spain. F. Spring, Alozaina, Cádiz, Spain. G. Operculum, inner side. H. Operculum, outer side." captionText-1="Fig. 12. Protoconch and radula of Mercuria balearica (Paladilhe, 1869). A–C. Venta El Pilar Spring, Málaga, Spain. A. Details of the protoconch. B. Details of the protoconch nucleus. C. Protoconch microsculpture. D–F. Stream at Sant Joan de Carbonell, Minorca. D. Overview of radular teeth rows. E. Rows of lateral and inner marginal radular teeth. F. Detailed view of the outer marginal teeth." captionText-2="Fig. 13. Anatomy of Mercuria balearica (Paladilhe, 1869). A, D, F. Stream at Sant Joan de Carbonell, Minorca, Spain. B, C, E. Venta El Pilar Spring, Málaga, Spain. A. Female genitalia. B. Penis and head. C. Prostate gland. D. Ctenidium and osphradium. E. Stomach. F. Perioesophageal ring." captionText-3="Fig. 14. Variation of the bursa copulatrix of Mercuria balearica (Paladilhe, 1869). A. Elongate, stream at Sant Joan de Carbonell, Minorca, Spain. B. Pyriform, Venta El Pilar Spring, Málaga, Spain. C–D. Pyriform, Fuente Valentín Spring, Alozaina, Cádiz, Spain. E–G. Detailed views of the seminal receptacle (Sr) and renal oviduct loops." figureDoi-0="http://doi.org/10.5281/zenodo.7867385" figureDoi-1="http://doi.org/10.5281/zenodo.7867387" figureDoi-2="http://doi.org/10.5281/zenodo.7867389" figureDoi-3="http://doi.org/10.5281/zenodo.7867391" httpUri-0="https://zenodo.org/record/7867385/files/figure.png" httpUri-1="https://zenodo.org/record/7867387/files/figure.png" httpUri-2="https://zenodo.org/record/7867389/files/figure.png" httpUri-3="https://zenodo.org/record/7867391/files/figure.png" pageId="20" pageNumber="21">Figs 11–14</figureCitation>
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; Supp. file 2: Tables S10–S13
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</paragraph>
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<subSubSection id="9CCA6559FFF4FFEDEDD0FD573612FCBD" pageId="20" pageNumber="21" type="reference_group">
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<paragraph id="D46F36D2FFF4FFEDEDD0FD57378EFD19" blockId="20.[189,677,713,739]" box="[189,677,713,739]" pageId="20" pageNumber="21">
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<treatmentCitation id="557110C3FFF4FFEDEDD0FD57378BFD19" author="Paladilhe A." box="[189,672,713,739]" page="113" pageId="20" pageNumber="21" year="1869">
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<taxonomicName id="13D04D51FFF4FFEDEDD0FD57378BFD19" authority="Paladilhe, 1869: 113" authorityName="Paladilhe" authorityPageNumber="113" authorityYear="1869" box="[189,672,713,739]" class="Gastropoda" family="Amnicolidae" genus="Amnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="balearica">
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<emphasis id="E6A4EAC0FFF4FFEDEDD0FD57348EFD19" box="[189,421,713,739]" italics="true" pageId="20" pageNumber="21">Amnicola balearica</emphasis>
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<bibRefCitation id="B0414B23FFF4FFEDECC1FD57378BFD19" author="Paladilhe A." box="[428,672,713,739]" pageId="20" pageNumber="21" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe, 1869: 113</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="D46F36D2FFF4FFEDEDD0FC95366DFCDE" blockId="20.[189,838,778,840]" box="[189,838,778,805]" pageId="20" pageNumber="21">
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<treatmentCitationGroup id="F4C011FCFFF4FFEDEDD0FC95366DFCDE" box="[189,838,778,805]" pageId="20" pageNumber="21">
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<taxonomicName id="13D04D51FFF4FFEDEDD0FC953489FCDE" ID-CoL="3ZTH3" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[189,418,778,804]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="balearica">
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<emphasis id="E6A4EAC0FFF4FFEDEDD0FC953489FCDE" box="[189,418,778,804]" italics="true" pageId="20" pageNumber="21">Mercuria balearica</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="557110C3FFF4FFEDECADFC9537B1FCDE" author="Boeters H." box="[448,666,778,805]" page="207" pageId="20" pageNumber="21" year="1988">
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<bibRefCitation id="B0414B23FFF4FFEDECADFC9537B1FCDE" author="Boeters H." box="[448,666,778,805]" pageId="20" pageNumber="21" pagination="181 - 261" refId="ref28493" refString="Boeters H. 1988. Moitessieriidae und Hydrobiidae in Spanien und Portugal (Gastropoda: Prosobranchia). Archiv fur Molluskenkunde 118: 181 - 261." type="journal article" year="1988">Boeters 1988: 207</bibRefCitation>
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</treatmentCitation>
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||
, figs 106–107.
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</paragraph>
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<paragraph id="D46F36D2FFF4FFEDEDD0FCB33612FCBD" blockId="20.[189,838,778,840]" box="[189,825,813,840]" pageId="20" pageNumber="21">
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<treatmentCitationGroup id="F4C011FCFFF4FFEDEDD0FCB33612FCBD" box="[189,825,813,840]" pageId="20" pageNumber="21">
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<taxonomicName id="13D04D51FFF4FFEDEDD0FCB334C2FCBD" ID-CoL="99393" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[189,489,813,839]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="balearica">
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||
<emphasis id="E6A4EAC0FFF4FFEDEDD0FCB334C2FCBD" box="[189,489,813,839]" italics="true" pageId="20" pageNumber="21">Pseudamnicola balearica</emphasis>
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</taxonomicName>
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||
–
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||
<treatmentCitation id="557110C3FFF4FFEDEF6AFCB337C8FCBD" author="Quintana J." box="[519,739,813,839]" page="19" pageId="20" pageNumber="21" year="2019">
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<bibRefCitation id="B0414B23FFF4FFEDEF6AFCB337C8FCBD" author="Quintana J." box="[519,739,813,839]" pageId="20" pageNumber="21" pagination="15 - 23" refId="ref30612" refString="Quintana J. 2019. Los moluscos del nivel B de la Cova Murada (Barranc d'Algendar, Ciutadella de Menorca, Illes Balears). Implicaciones tafonomicas. Bolleti de la Societat d'Historia Natural de les Balears 62: 15 - 23." type="journal article" year="2019">Quintana 2019: 19</bibRefCitation>
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</treatmentCitation>
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, fig 1a.
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</paragraph>
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|
||
<emphasis id="E6A4EAC0FFF4FFEDEDD0FCF034B1FC72" bold="true" box="[189,410,878,904]" pageId="20" pageNumber="21">Revised diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF4FFEDEDD0FC0331DBFB92" blockId="20.[189,1400,925,1128]" pageId="20" pageNumber="21">Shell ovate-conic; aperture obliquely broad ovate; protoconch microsculpture granulated; central radular tooth formula (4)3-C-3(4)/1-1; glandular oviduct 2.5 times as long as wide; albumen gland longer than capsule gland; bursa copulatrix pyriform to elongate; seminal receptacle elongate, with a short duct; penis darkly pigmented, as long as the penial appendix; penial appendix unpigmented or slightly pigmented at the junction with the penis, rounded, small, base narrow, medially positioned on the inner edge of the penis; nervous system pigmented, elongated (mean RPG ratio = 0.53).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CCA6559FFF4FFEDEDD0FB1136F4F865" pageId="20" pageNumber="21" type="materials_examined">
|
||
<paragraph id="D46F36D2FFF4FFEDEDD0FB11370DFB50" blockId="20.[189,550,1166,1194]" box="[189,550,1166,1194]" pageId="20" pageNumber="21">
|
||
<emphasis id="E6A4EAC0FFF4FFEDEDD0FB113447FB52" bold="true" box="[189,364,1166,1193]" pageId="20" pageNumber="21">Type material</emphasis>
|
||
(not examined)
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF4FFEDED87FB233168FB01" blockId="20.[189,1091,1213,1275]" pageId="20" pageNumber="21">
|
||
<materialsCitation id="64B83C8FFFF4FFEDED87FB233168FB01" ID-GBIF-Occurrence="4098914360" collectionCode="UM" country="France" location="FRANCE" pageId="20" pageNumber="21" specimenCode="UM.PLD.005" specimenCount="1" typeStatus="syntype">
|
||
<emphasis id="E6A4EAC0FFF4FFEDED87FB233464FB2D" bold="true" box="[234,335,1213,1239]" pageId="20" pageNumber="21">
|
||
<typeStatus id="0B6B8870FFF4FFEDED87FB233464FB2D" box="[234,335,1213,1239]" pageId="20" pageNumber="21" type="syntype">Syntype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="ACC77642FFF4FFEDEDD0FB7F341AFB01" box="[189,305,1249,1275]" name="France" pageId="20" pageNumber="21">FRANCE</collectingCountry>
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEC27FB7F34B0FB01" box="[330,411,1249,1275]" pageId="20" pageNumber="21" type="generic">1 spec.</specimenCount>
|
||
(sex unknown);
|
||
<specimenCode id="84769EA9FFF4FFEDEF0CFB7E3629FB01" box="[609,770,1248,1275]" pageId="20" pageNumber="21">UM.PLD.005</specimenCode>
|
||
(
|
||
<bibRefCitation id="B0414B23FFF4FFEDEE7FFB7F311CFB01" author="Breure A. & Audibert C." box="[786,1079,1248,1275]" pageId="20" pageNumber="21" pagination="28 - 39" refId="ref28746" refString="Breure A. & Audibert C. 2017. Under the spell of Bourguignat: Alcide Paladilhe's malacological work, with notes on his collection. Folia Conchyliologica 41: 28 - 39." type="journal article" year="2017">Breure & Audibert 2017</bibRefCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF4FFEDEDD0FABC3476FAC6" blockId="20.[189,349,1314,1340]" box="[189,349,1314,1340]" pageId="20" pageNumber="21">
|
||
<heading id="8F2781BEFFF4FFEDEDD0FABC3476FAC6" bold="true" box="[189,349,1314,1340]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
|
||
<emphasis id="E6A4EAC0FFF4FFEDEDD0FABC3476FAC6" bold="true" box="[189,349,1314,1340]" pageId="20" pageNumber="21">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF4FFEDEDD0FACE3740FA74" blockId="20.[187,1399,1360,1422]" pageId="20" pageNumber="21">
|
||
According to the original description (
|
||
<bibRefCitation id="B0414B23FFF4FFEDEF1EFACE360EFA91" author="Paladilhe A." box="[627,805,1360,1387]" pageId="20" pageNumber="21" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe 1869</bibRefCitation>
|
||
), the
|
||
<typeStatus id="0B6B8870FFF4FFEDEE0CFACF36BFFA91" box="[865,916,1361,1387]" pageId="20" pageNumber="21">type</typeStatus>
|
||
locality is Port Mahon,
|
||
<collectingCountry id="ACC77642FFF4FFEDE9C8FACE3020FA90" box="[1189,1291,1360,1386]" name="Spain" pageId="20" pageNumber="21">Minorca</collectingCountry>
|
||
,
|
||
<collectingRegion id="1614F830FFF4FFEDE878FACE3425FA74" country="Spain" name="Illes Baleares" pageId="20" pageNumber="21">Balearic Islands</collectingRegion>
|
||
[Port-Mahon (
|
||
<collectingRegion id="1614F830FFF4FFEDECD1FAEA3773FA74" box="[444,600,1395,1422]" country="Spain" name="Illes Baleares" pageId="20" pageNumber="21">Illes Balears</collectingRegion>
|
||
)].
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF4FFEDEDD0FA2B3481FA35" blockId="20.[189,426,1461,1487]" box="[189,426,1461,1487]" pageId="20" pageNumber="21">
|
||
<heading id="8F2781BEFFF4FFEDEDD0FA2B3481FA35" bold="true" box="[189,426,1461,1487]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
|
||
<emphasis id="E6A4EAC0FFF4FFEDEDD0FA2B3481FA35" bold="true" box="[189,426,1461,1487]" pageId="20" pageNumber="21">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF4FFEDEDD0FA7A37B4F8A4" blockId="20.[189,1399,1507,1886]" pageId="20" pageNumber="21">
|
||
<materialsCitation id="64B83C8FFFF4FFEDEDD0FA7A3048FA04" ID-GBIF-Occurrence="4098914332" box="[189,1379,1507,1534]" collectionCode="MNCN" country="Spain" location="Colarsega River in Port Mahon" municipality="Minorca" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94744" specimenCount="30" stateProvince="Islas Baleares">
|
||
<collectingCountry id="ACC77642FFF4FFEDEDD0FA7A3424FA04" box="[189,271,1508,1534]" name="Spain" pageId="20" pageNumber="21">SPAIN</collectingCountry>
|
||
–
|
||
<emphasis id="E6A4EAC0FFF4FFEDEC5FFA7A34CBFA04" bold="true" box="[306,480,1508,1534]" pageId="20" pageNumber="21">
|
||
<collectingRegion id="1614F830FFF4FFEDEC5FFA7A34CBFA04" box="[306,480,1508,1534]" country="Spain" name="Illes Baleares" pageId="20" pageNumber="21">Islas Baleares</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEC96FA7A374FFA04" box="[507,612,1508,1534]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
;
|
||
<collectingMunicipality id="340BACA8FFF4FFEDEF1FFA7A37F3FA04" box="[626,728,1508,1534]" pageId="20" pageNumber="21">Minorca</collectingMunicipality>
|
||
,
|
||
<location id="D10F6009FFF4FFEDEF8BFA7A314AFA04" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDEF8BFA7A314AFA04" box="[742,1121,1508,1534]" country="Spain" municipality="Minorca" name="Colarsega River in Port Mahon" pageId="20" pageNumber="21" stateProvince="Islas Baleares">Colarsega River in Port Mahon</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDE903FA7A3048FA04" box="[1134,1379,1507,1534]" pageId="20" pageNumber="21">MNCN 15.05/94744</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDE801FA7A3169F9D8" ID-GBIF-Occurrence="4098914358" collectionCode="MNCN" location="Sant Joan de Carbonell" municipality="Minorca" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94745" specimenCount="30">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEDD0F999340EF9DB" box="[189,293,1543,1569]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
;
|
||
<collectingMunicipality id="340BACA8FFF4FFEDEC5DF99934BDF9DB" box="[304,406,1543,1569]" pageId="20" pageNumber="21">Minorca</collectingMunicipality>
|
||
, stream near
|
||
<location id="D10F6009FFF4FFEDEF5FF999366EF9DB" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDEF5FF999366EF9DB" box="[562,837,1543,1569]" municipality="Minorca" name="Sant Joan de Carbonell" pageId="20" pageNumber="21">Sant Joan de Carbonell</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDEE3DF9993169F9D8" box="[848,1090,1543,1570]" pageId="20" pageNumber="21">MNCN 15.05/94745</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDE927F9993700F9BF" ID-GBIF-Occurrence="4098914420" collectionCode="MNCN" location="Cala en Porter" municipality="Minorca" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94746" specimenCount="30">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDE936F99931E8F9DB" box="[1115,1219,1543,1569]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
;
|
||
<collectingMunicipality id="340BACA8FFF4FFEDE9A2F999301EF9DB" box="[1231,1333,1543,1569]" pageId="20" pageNumber="21">Minorca</collectingMunicipality>
|
||
,
|
||
<location id="D10F6009FFF4FFEDE82CF9993407F9BF" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDE82CF9993407F9BF" municipality="Minorca" name="Cala en Porter" pageId="20" pageNumber="21">Cala en Porter</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDEC55F9B43700F9BF" box="[312,555,1578,1605]" pageId="20" pageNumber="21">MNCN 15.05/94746</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDEF5EF9B5304AF9BF" ID-GBIF-Occurrence="4098914308" box="[563,1377,1578,1605]" collectionCode="MNCN" location="Barranc de Bec" municipality="Son Bou" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94747" specimenCount="30" stateProvince="Minorca">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEF28F9B43786F9BF" box="[581,685,1578,1605]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
;
|
||
<collectingRegion id="1614F830FFF4FFEDEFD4F9B43634F9BE" box="[697,799,1578,1604]" pageId="20" pageNumber="21">Minorca</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="340BACA8FFF4FFEDEE46F9B436BEF9BF" box="[811,917,1578,1605]" pageId="20" pageNumber="21">Son Bou</collectingMunicipality>
|
||
,
|
||
<location id="D10F6009FFF4FFEDEECDF9B53172F9BF" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDEECDF9B53172F9BF" box="[928,1113,1578,1605]" municipality="Son Bou" name="Barranc de Bec" pageId="20" pageNumber="21" stateProvince="Minorca">Barranc de Bec</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDE90BF9B43077F9BF" box="[1126,1372,1578,1605]" pageId="20" pageNumber="21">MNCN 15.05/94747</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDE805F9B536CEF992" ID-GBIF-Occurrence="4098914410" collectionCode="MNCN" location="El Chorro Spring" municipality="Ardales" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94732" specimenCount="30" stateProvince="Malaga">
|
||
–
|
||
<emphasis id="E6A4EAC0FFF4FFEDEDD0F9D23437F99D" bold="true" box="[189,284,1612,1639]" pageId="20" pageNumber="21">
|
||
<collectingRegion id="1614F830FFF4FFEDEDD0F9D23437F99D" box="[189,284,1612,1639]" pageId="20" pageNumber="21">Málaga</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEC5BF9D334B4F992" box="[310,415,1613,1640]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
;
|
||
<collectingMunicipality id="340BACA8FFF4FFEDECC7F9D33723F99D" box="[426,520,1613,1639]" pageId="20" pageNumber="21">Ardales</collectingMunicipality>
|
||
,
|
||
<location id="D10F6009FFF4FFEDEF78F9D337CFF99D" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDEF78F9D337CFF99D" box="[533,740,1613,1639]" municipality="Ardales" name="El Chorro Spring" pageId="20" pageNumber="21" stateProvince="Malaga">El Chorro Spring</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDEF9CF9D336CEF992" box="[753,997,1613,1640]" pageId="20" pageNumber="21">MNCN 15.05/94732</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDEE80F9D037EBF971" ID-GBIF-Occurrence="4098914314" collectionCode="MNCN" location="Montecorto" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94731, MNCN 15.05/94737" specimenCount="30">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEE92F9D33143F992" box="[1023,1128,1613,1640]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
; spring in
|
||
<location id="D10F6009FFF4FFEDE988F9D03059F992" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDE988F9D03059F992" box="[1253,1394,1614,1640]" name="Montecorto" pageId="20" pageNumber="21">Montecorto</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDEDD0F9EF3493F971" box="[189,440,1648,1675]" pageId="20" pageNumber="21">MNCN 15.05/94731</specimenCode>
|
||
,
|
||
<specimenCode id="84769EA9FFF4FFEDECA4F9EF37EBF971" box="[457,704,1648,1675]" pageId="20" pageNumber="21">MNCN 15.05/94737</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDEFA0F9EF304BF971" ID-GBIF-Occurrence="4098914406" box="[717,1376,1648,1675]" collectionCode="MNCN" location="Churriana" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94733" specimenCount="30">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEF89F9EF367AF971" box="[740,849,1649,1675]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
; trough in
|
||
<location id="D10F6009FFF4FFEDEEB3F9EE317CF970" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDEEB3F9EE317CF970" box="[990,1111,1648,1674]" name="Churriana" pageId="20" pageNumber="21">Churriana</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDE905F9EF304BF971" box="[1128,1376,1648,1675]" pageId="20" pageNumber="21">MNCN 15.05/94733</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDE801F9EF3114F954" ID-GBIF-Occurrence="4098914313" collectionCode="MNCN" location="Venta El Pilar Spring" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94734, MNCN 15.05/94738" specimenCount="30">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEDD0F90A340CF954" box="[189,295,1684,1710]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
;
|
||
<location id="D10F6009FFF4FFEDEC59F90A371CF954" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDEC59F90A371CF954" box="[308,567,1684,1710]" name="Venta El Pilar Spring" pageId="20" pageNumber="21">Venta El Pilar Spring</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDEF28F90A3617F954" box="[581,828,1683,1710]" pageId="20" pageNumber="21">MNCN 15.05/94734</specimenCode>
|
||
,
|
||
<specimenCode id="84769EA9FFF4FFEDEE27F90A3114F954" box="[842,1087,1683,1710]" pageId="20" pageNumber="21">MNCN 15.05/94738</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDE924F90A3712F928" ID-GBIF-Occurrence="4098914380" collectionCode="MNCN" location="Los Granados" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94735" specimenCount="30">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDE931F90A31EDF954" box="[1116,1222,1684,1710]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
; trough in
|
||
<location id="D10F6009FFF4FFEDE827F90A341AF92B" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDE827F90A341AF92B" name="Los Granados" pageId="20" pageNumber="21">Los Granados</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDEC2CF9293712F928" box="[321,569,1719,1746]" pageId="20" pageNumber="21">MNCN 15.05/94735</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDEF28F92931DAF928" ID-GBIF-Occurrence="4098914390" box="[581,1265,1719,1746]" collectionCode="MNCN" location="Montecorto" municipality="Montecorto" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94736" specimenCount="30">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEF36F92937ECF92B" box="[603,711,1719,1745]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
; stream in
|
||
<collectingMunicipality id="340BACA8FFF4FFEDEE38F92936C9F92B" box="[853,994,1719,1745]" pageId="20" pageNumber="21">Montecorto</collectingMunicipality>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDEE9FF92931C6F928" box="[1010,1261,1719,1746]" pageId="20" pageNumber="21">MNCN 15.05/94736</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDE991F9293677F90F" ID-GBIF-Occurrence="4098914413" collectionCode="MNCN" location="Valentin Spring" municipality="Alozaina" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94739" specimenCount="30" stateProvince="Cadiz">
|
||
–
|
||
<emphasis id="E6A4EAC0FFF4FFEDE87AF928304BF92B" bold="true" box="[1303,1376,1718,1745]" pageId="20" pageNumber="21">
|
||
<collectingRegion id="1614F830FFF4FFEDE87AF928304BF92B" box="[1303,1376,1718,1745]" pageId="20" pageNumber="21">Cádiz</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEDD0F9443408F90F" box="[189,291,1754,1781]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
;
|
||
<collectingMunicipality id="340BACA8FFF4FFEDEC46F94434B3F90E" box="[299,408,1754,1780]" pageId="20" pageNumber="21">Alozaina</collectingMunicipality>
|
||
,
|
||
<location id="D10F6009FFF4FFEDECCCF9443771F90E" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDECCCF9443771F90E" box="[417,602,1754,1780]" municipality="Alozaina" name="Valentin Spring" pageId="20" pageNumber="21" stateProvince="Cadiz">Valentín Spring</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDEF09F9443672F90F" box="[612,857,1754,1781]" pageId="20" pageNumber="21">MNCN 15.05/94739</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDEE0CF945371EF8E2" ID-GBIF-Occurrence="4098914423" collectionCode="MNCN" location="Cano de la Rambla" municipality="Fontanar" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94740" specimenCount="30" stateProvince="Jaen">
|
||
–
|
||
<emphasis id="E6A4EAC0FFF4FFEDEE1EF9473685F90E" bold="true" box="[883,942,1753,1780]" pageId="20" pageNumber="21">
|
||
<collectingRegion id="1614F830FFF4FFEDEE1EF9473685F90E" box="[883,942,1753,1780]" pageId="20" pageNumber="21">Jaén</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEEA8F9443101F90F" box="[965,1066,1754,1781]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
;
|
||
<collectingMunicipality id="340BACA8FFF4FFEDE959F94531B5F90F" box="[1076,1182,1755,1781]" pageId="20" pageNumber="21">Fontanar</collectingMunicipality>
|
||
, stream in
|
||
<location id="D10F6009FFF4FFEDE875F9443413F8ED" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDE875F9443413F8ED" municipality="Fontanar" name="Cano de la Rambla" pageId="20" pageNumber="21" stateProvince="Jaen">Caño de la Rambla</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDEC2EF963371EF8E2" box="[323,565,1789,1816]" pageId="20" pageNumber="21">MNCN 15.05/94740</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDEF56F960305CF8E2" ID-GBIF-Occurrence="4098914415" box="[571,1399,1789,1816]" collectionCode="MNCN" location="Cinco Canos Spring" municipality="Peal de Becerro" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94743" specimenCount="30">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEF21F9633798F8E2" box="[588,691,1789,1816]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
;
|
||
<collectingMunicipality id="340BACA8FFF4FFEDEFD3F9633652F8E2" box="[702,889,1789,1816]" pageId="20" pageNumber="21">Peal de Becerro</collectingMunicipality>
|
||
,
|
||
<location id="D10F6009FFF4FFEDEEE9F9633159F8ED" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDEEE9F9633159F8ED" box="[900,1138,1789,1815]" municipality="Peal de Becerro" name="Cinco Canos Spring" pageId="20" pageNumber="21">Cinco Caños Spring</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDE910F9633058F8E2" box="[1149,1395,1789,1816]" pageId="20" pageNumber="21">MNCN 15.05/94743</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDEDD0F8BF312AF8C1" ID-GBIF-Occurrence="4098914323" box="[189,1025,1824,1851]" collectionCode="MNCN" location="Arabic Spring" municipality="Mojacar" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94741" specimenCount="30" stateProvince="Granada">
|
||
–
|
||
<emphasis id="E6A4EAC0FFF4FFEDEDB9F8BE346FF8C0" bold="true" box="[212,324,1824,1850]" pageId="20" pageNumber="21">
|
||
<collectingRegion id="1614F830FFF4FFEDEDB9F8BE346FF8C0" box="[212,324,1824,1850]" pageId="20" pageNumber="21">Granada</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDEC0DF8BF34E2F8C1" box="[352,457,1825,1851]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
;
|
||
<collectingMunicipality id="340BACA8FFF4FFEDECBBF8BE3711F8C0" box="[470,570,1824,1850]" pageId="20" pageNumber="21">Mojácar</collectingMunicipality>
|
||
,
|
||
<location id="D10F6009FFF4FFEDEF29F8BE37C4F8C0" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDEF29F8BE37C4F8C0" box="[580,751,1824,1850]" municipality="Mojacar" name="Arabic Spring" pageId="20" pageNumber="21" stateProvince="Granada">Arabic Spring</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDEF91F8BF36DEF8C1" box="[764,1013,1824,1851]" pageId="20" pageNumber="21">MNCN 15.05/94741</specimenCode>
|
||
).
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFF4FFEDE964F8BF37B4F8A4" ID-GBIF-Occurrence="4098914376" collectionCode="MNCN" location="Sierra de la Muela" pageId="20" pageNumber="21" specimenCode="MNCN 15.05/94742" specimenCount="30" stateProvince="Murcia">
|
||
–
|
||
<emphasis id="E6A4EAC0FFF4FFEDE94DF8BE3157F8C0" bold="true" box="[1056,1148,1824,1850]" pageId="20" pageNumber="21">
|
||
<collectingRegion id="1614F830FFF4FFEDE94DF8BE3157F8C0" box="[1056,1148,1824,1850]" country="Spain" name="Murcia" pageId="20" pageNumber="21">Murcia</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="C2D6FD5BFFF4FFEDE9F5F8BF302AF8C1" box="[1176,1281,1825,1851]" pageId="20" pageNumber="21">30 specs</specimenCount>
|
||
; spring in
|
||
<location id="D10F6009FFF4FFEDEDD0F8DA34B2F8A4" LSID="urn:lsid:plazi:treatment:5C7987C4FFF4FFE0EF51FDFA314EF9B4:D10F6009FFF4FFEDEDD0F8DA34B2F8A4" box="[189,409,1860,1886]" name="Sierra de la Muela" pageId="20" pageNumber="21" stateProvince="Murcia">Sierra de la Muela</location>
|
||
;
|
||
<specimenCode id="84769EA9FFF4FFEDECC9F8DA37B0F8A4" box="[420,667,1859,1886]" pageId="20" pageNumber="21">MNCN 15.05/94742</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF4FFEDEDD0F81B36F4F865" blockId="20.[189,991,1925,1952]" box="[189,991,1925,1952]" pageId="20" pageNumber="21">Additional locality information is provided in Supp. file 1: Table S1.</paragraph>
|
||
</subSubSection>
|
||
<caption id="80AF665AFFF5FFECEDD0F8FA303FF812" ID-DOI="http://doi.org/10.5281/zenodo.7867385" ID-Zenodo-Dep="7867385" httpUri="https://zenodo.org/record/7867385/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1892,1918]" targetBox="[189,1398,274,1842]" targetPageId="21">
|
||
<paragraph id="D46F36D2FFF5FFECEDD0F8FA303FF812" blockId="21.[188,1399,1892,2025]" pageId="21" pageNumber="22">
|
||
<emphasis id="E6A4EAC0FFF5FFECEDD0F8FA343CF885" bold="true" box="[189,279,1892,1919]" pageId="21" pageNumber="22">Fig. 11.</emphasis>
|
||
Shells and operculum of
|
||
<taxonomicName id="13D04D51FFF5FFECEF53F8F836DDF87A" authority="(Paladilhe, 1869)" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[574,1014,1893,1920]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF5FFECEF53F8F83608F885" box="[574,803,1893,1919]" italics="true" pageId="21" pageNumber="22">Mercuria balearica</emphasis>
|
||
(
|
||
<bibRefCitation id="B0414B23FFF5FFECEE5CF8FB36C6F87A" author="Paladilhe A." box="[817,1005,1893,1920]" pageId="21" pageNumber="22" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="E6A4EAC0FFF5FFECE96DF8FA31A9F884" bold="true" box="[1024,1154,1892,1918]" pageId="21" pageNumber="22">A–B, G–H</emphasis>
|
||
. Stream at Sant Joan de Carbonell, Minorca, Spain.
|
||
<emphasis id="E6A4EAC0FFF5FFECEF5CF816376FF858" bold="true" box="[561,580,1928,1954]" pageId="21" pageNumber="22">C</emphasis>
|
||
. Colarsega River, near the Port of Mahon (type locality).
|
||
<emphasis id="E6A4EAC0FFF5FFECE86EF816303DF858" bold="true" box="[1283,1302,1928,1954]" pageId="21" pageNumber="22">D</emphasis>
|
||
. Fuente Árabe Spring, Mojácar, Granada, Spain (unpigmented specimen).
|
||
<emphasis id="E6A4EAC0FFF5FFECEEB0F83536DAF83F" bold="true" box="[989,1009,1963,1989]" pageId="21" pageNumber="22">E</emphasis>
|
||
. Venta El Pilar Spring, Málaga, Spain.
|
||
<emphasis id="E6A4EAC0FFF5FFECEC63F850340BF812" bold="true" box="[270,288,1998,2024]" pageId="21" pageNumber="22">F</emphasis>
|
||
. Spring, Alozaina, Cádiz, Spain.
|
||
<emphasis id="E6A4EAC0FFF5FFECEFCAF8503795F812" bold="true" box="[679,702,1998,2024]" pageId="21" pageNumber="22">G</emphasis>
|
||
. Operculum, inner side.
|
||
<emphasis id="E6A4EAC0FFF5FFECEE8CF85036D3F812" bold="true" box="[993,1016,1998,2024]" pageId="21" pageNumber="22">H</emphasis>
|
||
. Operculum, outer side.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="9CCA6559FFF6FFEEEDD0FE963705FBE1" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
|
||
<paragraph id="D46F36D2FFF6FFEFEDD0FE963465FED8" blockId="22.[189,334,264,290]" box="[189,334,264,290]" pageId="22" pageNumber="23">
|
||
<heading id="8F2781BEFFF6FFEFEDD0FE963465FED8" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
|
||
<emphasis id="E6A4EAC0FFF6FFEFEDD0FE963465FED8" bold="true" box="[189,334,264,290]" pageId="22" pageNumber="23">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF6FFEFEDD0FEA93045FDFB" blockId="22.[188,1400,311,514]" pageId="22" pageNumber="23">
|
||
<smallCapsWord id="D289A00EFFF6FFEFEDD0FEA93429FEAB" baselines="330,331" box="[189,258,311,337]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="22" pageNumber="23">SHELL</smallCapsWord>
|
||
. Ovate-conic, whorls 4–5, height
|
||
<quantity id="13289B37FFF6FFEFEFC2FEA93604FEA8" box="[687,815,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="22" pageNumber="23" unit="mm" value="2.75" valueMax="3.5" valueMin="2.0">2–3.5 mm</quantity>
|
||
, width
|
||
<quantity id="13289B37FFF6FFEFEEFCFEA9310FFEA8" box="[913,1060,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.6" metricValueMin="1.4" pageId="22" pageNumber="23" unit="mm" value="2.0" valueMax="2.6" valueMin="1.4">1.4–2.6 mm</quantity>
|
||
(
|
||
<figureCitation id="4CEB2A57FFF6FFEFE957FEA931E1FEAB" box="[1082,1226,311,337]" captionStart="Fig" captionStartId="21.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1842]" captionTargetId="figure-11@21.[189,1398,274,1842]" captionTargetPageId="21" captionText="Fig. 11. Shells and operculum of Mercuria balearica (Paladilhe, 1869). A–B, G–H. Stream at Sant Joan de Carbonell, Minorca, Spain. C. Colarsega River, near the Port of Mahon (type locality). D. Fuente Árabe Spring, Mojácar, Granada, Spain (unpigmented specimen). E. Venta El Pilar Spring, Málaga, Spain. F. Spring, Alozaina, Cádiz, Spain. G. Operculum, inner side. H. Operculum, outer side." figureDoi="http://doi.org/10.5281/zenodo.7867385" httpUri="https://zenodo.org/record/7867385/files/figure.png" pageId="22" pageNumber="23">Fig. 11A–F</figureCitation>
|
||
; Supp. file 2: Table S10); periostracum yellowish to brown; protoconch with 1.5 whorls, ca 350 µm wide, nucleus less than 200 µm wide; protoconch microsculpture granulated (
|
||
<figureCitation id="4CEB2A57FFF6FFEFEEF3FEE33106FE62" box="[926,1069,381,408]" captionStart="Fig" captionStartId="22.[189,232,1892,1918]" captionTargetBox="[189,1398,1115,1846]" captionTargetId="figure-302@22.[189,1398,1115,1846]" captionTargetPageId="22" captionText="Fig. 12. Protoconch and radula of Mercuria balearica (Paladilhe, 1869). A–C. Venta El Pilar Spring, Málaga, Spain. A. Details of the protoconch. B. Details of the protoconch nucleus. C. Protoconch microsculpture. D–F. Stream at Sant Joan de Carbonell, Minorca. D. Overview of radular teeth rows. E. Rows of lateral and inner marginal radular teeth. F. Detailed view of the outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.7867387" httpUri="https://zenodo.org/record/7867387/files/figure.png" pageId="22" pageNumber="23">Fig. 12A–C</figureCitation>
|
||
); teleoconch whorls convex, separated by a deep suture; body whorl large, convex, occupying about two-thirds of the total shell length; aperture obliquely broad ovate, complete; inner lip thicker than outer lip; aperture margin straight, inner lip touching the shell wall; umbilicus narrow, not covered by the inner lip (
|
||
<figureCitation id="4CEB2A57FFF6FFEFE9BBFE793049FDFB" box="[1238,1378,487,513]" captionStart="Fig" captionStartId="21.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1842]" captionTargetId="figure-11@21.[189,1398,274,1842]" captionTargetPageId="21" captionText="Fig. 11. Shells and operculum of Mercuria balearica (Paladilhe, 1869). A–B, G–H. Stream at Sant Joan de Carbonell, Minorca, Spain. C. Colarsega River, near the Port of Mahon (type locality). D. Fuente Árabe Spring, Mojácar, Granada, Spain (unpigmented specimen). E. Venta El Pilar Spring, Málaga, Spain. F. Spring, Alozaina, Cádiz, Spain. G. Operculum, inner side. H. Operculum, outer side." figureDoi="http://doi.org/10.5281/zenodo.7867385" httpUri="https://zenodo.org/record/7867385/files/figure.png" pageId="22" pageNumber="23">Fig. 11A–F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF6FFEFEDD0FDB73665FD9C" blockId="22.[189,1398,552,614]" pageId="22" pageNumber="23">
|
||
<smallCapsWord id="D289A00EFFF6FFEFEDD0FDB73468FDB9" baselines="572,572" box="[189,323,553,579]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Operculum" pageId="22" pageNumber="23">OPERCULUM</smallCapsWord>
|
||
. As for the genus, orange to brown, sometimes yellowish, about two whorls; muscle attachment oval, located near the nucleus (
|
||
<figureCitation id="4CEB2A57FFF6FFEFEFDCFDD2366AFD9C" box="[689,833,588,614]" captionStart="Fig" captionStartId="21.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1842]" captionTargetId="figure-11@21.[189,1398,274,1842]" captionTargetPageId="21" captionText="Fig. 11. Shells and operculum of Mercuria balearica (Paladilhe, 1869). A–B, G–H. Stream at Sant Joan de Carbonell, Minorca, Spain. C. Colarsega River, near the Port of Mahon (type locality). D. Fuente Árabe Spring, Mojácar, Granada, Spain (unpigmented specimen). E. Venta El Pilar Spring, Málaga, Spain. F. Spring, Alozaina, Cádiz, Spain. G. Operculum, inner side. H. Operculum, outer side." figureDoi="http://doi.org/10.5281/zenodo.7867385" httpUri="https://zenodo.org/record/7867385/files/figure.png" pageId="22" pageNumber="23">Fig. 11G–H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF6FFEFEDD0FD10313AFC64" blockId="22.[188,1401,653,926]" pageId="22" pageNumber="23">
|
||
<smallCapsWord id="D289A00EFFF6FFEFEDD0FD103432FD52" baselines="673,673" box="[189,281,654,680]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Radula" pageId="22" pageNumber="23">RADULA</smallCapsWord>
|
||
. Length intermediate, ca 800 µm long (35% of total shell length), containing about 65 rows of teeth. Central tooth formula (4)3-C-3(4)/1-1, central cusp V shaped, cutting edge slightly concave (
|
||
<figureCitation id="4CEB2A57FFF6FFEFEDA5FD4D341EFD14" box="[200,309,723,750]" captionStart="Fig" captionStartId="22.[189,232,1892,1918]" captionTargetBox="[189,1398,1115,1846]" captionTargetId="figure-302@22.[189,1398,1115,1846]" captionTargetPageId="22" captionText="Fig. 12. Protoconch and radula of Mercuria balearica (Paladilhe, 1869). A–C. Venta El Pilar Spring, Málaga, Spain. A. Details of the protoconch. B. Details of the protoconch nucleus. C. Protoconch microsculpture. D–F. Stream at Sant Joan de Carbonell, Minorca. D. Overview of radular teeth rows. E. Rows of lateral and inner marginal radular teeth. F. Detailed view of the outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.7867387" httpUri="https://zenodo.org/record/7867387/files/figure.png" pageId="22" pageNumber="23">Fig. 12D</figureCitation>
|
||
). Lateral tooth formula (3)4-C-4(3), central cusp V shaped and slightly longer than the central tooth one. Inner marginal teeth with 14–18 cusps; outer marginal teeth with 15–17 cusps (
|
||
<figureCitation id="4CEB2A57FFF6FFEFE9B2FD693041FCEB" box="[1247,1386,759,785]" captionStart="Fig" captionStartId="22.[189,232,1892,1918]" captionTargetBox="[189,1398,1115,1846]" captionTargetId="figure-302@22.[189,1398,1115,1846]" captionTargetPageId="22" captionText="Fig. 12. Protoconch and radula of Mercuria balearica (Paladilhe, 1869). A–C. Venta El Pilar Spring, Málaga, Spain. A. Details of the protoconch. B. Details of the protoconch nucleus. C. Protoconch microsculpture. D–F. Stream at Sant Joan de Carbonell, Minorca. D. Overview of radular teeth rows. E. Rows of lateral and inner marginal radular teeth. F. Detailed view of the outer marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.7867387" httpUri="https://zenodo.org/record/7867387/files/figure.png" pageId="22" pageNumber="23">Fig. 12E–F</figureCitation>
|
||
). Radular data were collected from the following specimens: MNCN 15.05/94745 – stream near Sant Joan de Carbonell,
|
||
<collectingCountry id="ACC77642FFF6FFEFEC32FCA334EDFCAD" box="[351,454,829,855]" name="Spain" pageId="22" pageNumber="23">Minorca</collectingCountry>
|
||
; MNCN 15.05/94747 – Barranc de Bec, Son Bou,
|
||
<collectingCountry id="ACC77642FFF6FFEFE97CFCA33153FCAD" box="[1041,1144,829,855]" name="Spain" pageId="22" pageNumber="23">Minorca</collectingCountry>
|
||
; MNCN 15.05/94738 – Venta El Pilar Spring, Málaga; MNCN 15.05/94741 – Arabic Spring, Mojácar,
|
||
<collectingCountry id="ACC77642FFF6FFEFE9CBFCFE3026FC80" box="[1190,1293,864,890]" name="Grenada" pageId="22" pageNumber="23">Granada</collectingCountry>
|
||
; MNCN 15.05/94743 – Fuente de los Cinco Caños Spring, Peal de Becerro, Jaén.
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF6FFEEEDD0FC5B31EFFE4A" blockId="22.[189,1399,965,1062]" lastBlockId="23.[188,1398,265,432]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
<smallCapsWord id="D289A00EFFF6FFEFEDD0FC5B3471FC25" baselines="984,984" box="[189,346,965,991]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pigmentation" pageId="22" pageNumber="23">PIGMENTATION</smallCapsWord>
|
||
<smallCapsWord id="D289A00EFFF6FFEFEC0EFC5534BBFC24" baselines="985" box="[355,400,971,990]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="22" pageNumber="23">AND</smallCapsWord>
|
||
<smallCapsWord id="D289A00EFFF6FFEFECF5FC5434D4FC27" baselines="984" box="[408,511,970,989]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="anatomy" pageId="22" pageNumber="23">ANATOMY</smallCapsWord>
|
||
. Animal slightly darkly pigmented, although unpigmented specimens were also found (
|
||
<figureCitation id="4CEB2A57FFF6FFEFEC22FC763495FBF8" box="[335,446,1000,1026]" captionStart="Fig" captionStartId="21.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1842]" captionTargetId="figure-11@21.[189,1398,274,1842]" captionTargetPageId="21" captionText="Fig. 11. Shells and operculum of Mercuria balearica (Paladilhe, 1869). A–B, G–H. Stream at Sant Joan de Carbonell, Minorca, Spain. C. Colarsega River, near the Port of Mahon (type locality). D. Fuente Árabe Spring, Mojácar, Granada, Spain (unpigmented specimen). E. Venta El Pilar Spring, Málaga, Spain. F. Spring, Alozaina, Cádiz, Spain. G. Operculum, inner side. H. Operculum, outer side." figureDoi="http://doi.org/10.5281/zenodo.7867385" httpUri="https://zenodo.org/record/7867385/files/figure.png" pageId="22" pageNumber="23">Fig. 11D</figureCitation>
|
||
); head and tentacles black, pigmentation lighter on eye lobes, snout and neck; snout about as long as wide, approximately parallel-sided, with medium distal lobation. Ctenidium occupying almost the total length of the pallial cavity; 21–27 gill filaments; filaments broad, triangular, fused at the base by an epithelium (
|
||
<figureCitation id="4CEB2A57FFF7FFEEEF38FEB237EAFEBC" box="[597,705,300,326]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[189,1398,1130,1885]" captionTargetId="figure-328@23.[189,1398,1130,1885]" captionTargetPageId="23" captionText="Fig. 13. Anatomy of Mercuria balearica (Paladilhe, 1869). A, D, F. Stream at Sant Joan de Carbonell, Minorca, Spain. B, C, E. Venta El Pilar Spring, Málaga, Spain. A. Female genitalia. B. Penis and head. C. Prostate gland. D. Ctenidium and osphradium. E. Stomach. F. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867389" httpUri="https://zenodo.org/record/7867389/files/figure.png" pageId="23" pageNumber="24">Fig. 13D</figureCitation>
|
||
). Osphradium elongate, more than 3 times as long as broad, positioned opposite to the middle of the ctenidium. Stomach almost as long as wide with two nearly equally sized chambers (Supp. file 2: Table S11); style sac longer than wide, with the unpigmented intestine surrounding its distal part before continuing on as a straight rectum (
|
||
<figureCitation id="4CEB2A57FFF7FFEEE921FE08319DFE4A" box="[1100,1206,406,432]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[189,1398,1130,1885]" captionTargetId="figure-328@23.[189,1398,1130,1885]" captionTargetPageId="23" captionText="Fig. 13. Anatomy of Mercuria balearica (Paladilhe, 1869). A, D, F. Stream at Sant Joan de Carbonell, Minorca, Spain. B, C, E. Venta El Pilar Spring, Málaga, Spain. A. Female genitalia. B. Penis and head. C. Prostate gland. D. Ctenidium and osphradium. E. Stomach. F. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867389" httpUri="https://zenodo.org/record/7867389/files/figure.png" pageId="23" pageNumber="24">Fig. 13E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="80AF665AFFF6FFEFEDD0F8FA3001F812" ID-DOI="http://doi.org/10.5281/zenodo.7867387" ID-Zenodo-Dep="7867387" httpUri="https://zenodo.org/record/7867387/files/figure.png" pageId="22" pageNumber="23" startId="22.[189,232,1892,1918]" targetBox="[189,1398,1115,1846]" targetPageId="22">
|
||
<paragraph id="D46F36D2FFF6FFEFEDD0F8FA3001F812" blockId="22.[189,1399,1892,2025]" pageId="22" pageNumber="23">
|
||
<emphasis id="E6A4EAC0FFF6FFEFEDD0F8FA3430F885" bold="true" box="[189,283,1892,1919]" pageId="22" pageNumber="23">Fig. 12.</emphasis>
|
||
Protoconch and radula of
|
||
<taxonomicName id="13D04D51FFF6FFEFEF30F8F83130F87A" authority="(Paladilhe, 1869)" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[605,1051,1893,1920]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF6FFEFEF30F8F8366FF885" box="[605,836,1893,1919]" italics="true" pageId="22" pageNumber="23">Mercuria balearica</emphasis>
|
||
(
|
||
<bibRefCitation id="B0414B23FFF6FFEFEE3EF8FB3139F87A" author="Paladilhe A." box="[851,1042,1893,1920]" pageId="22" pageNumber="23" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="E6A4EAC0FFF6FFEFE947F8FA314AF884" bold="true" box="[1066,1121,1892,1918]" pageId="22" pageNumber="23">A–C</emphasis>
|
||
. Venta El Pilar Spring, Málaga, Spain.
|
||
<emphasis id="E6A4EAC0FFF6FFEFEC13F81634B8F858" bold="true" box="[382,403,1928,1954]" pageId="22" pageNumber="23">A</emphasis>
|
||
. Details of the protoconch.
|
||
<emphasis id="E6A4EAC0FFF6FFEFEF9CF816362EF858" bold="true" box="[753,773,1928,1954]" pageId="22" pageNumber="23">B</emphasis>
|
||
. Details of the protoconch nucleus.
|
||
<emphasis id="E6A4EAC0FFF6FFEFE9A5F81631F6F858" bold="true" box="[1224,1245,1928,1954]" pageId="22" pageNumber="23">C</emphasis>
|
||
. Protoconch microsculpture.
|
||
<emphasis id="E6A4EAC0FFF6FFEFEC10F8353499F83F" bold="true" box="[381,434,1963,1989]" pageId="22" pageNumber="23">D–F</emphasis>
|
||
. Stream at Sant Joan de Carbonell, Minorca.
|
||
<emphasis id="E6A4EAC0FFF6FFEFEEBBF83536C1F83F" bold="true" box="[982,1002,1963,1989]" pageId="22" pageNumber="23">D</emphasis>
|
||
. Overview of radular teeth rows.
|
||
<emphasis id="E6A4EAC0FFF6FFEFEDD2F85035FAF812" bold="true" box="[191,209,1998,2024]" pageId="22" pageNumber="23">E</emphasis>
|
||
. Rows of lateral and inner marginal radular teeth.
|
||
<emphasis id="E6A4EAC0FFF6FFEFEE4DF8503619F812" bold="true" box="[800,818,1998,2024]" pageId="22" pageNumber="23">F</emphasis>
|
||
. Detailed view of the outer marginal teeth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D46F36D2FFF7FFEEEDD0FE463415FD5B" blockId="23.[189,1401,471,673]" pageId="23" pageNumber="24">
|
||
<smallCapsWord id="D289A00EFFF7FFEEEDD0FE4635D4FE08" baselines="491,491" box="[189,255,472,498]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="23" pageNumber="24">MALE</smallCapsWord>
|
||
<smallCapsWord id="D289A00EFFF7FFEEEC64FE43345CFE0A" baselines="491" box="[265,375,477,496]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="23" pageNumber="24">GENITALIA</smallCapsWord>
|
||
. Prostate gland bean-shaped, about 2 times as long as wide (Supp. file 2: Table S13), connected by the posterior vas deferens to a convoluted seminal vesicle and the testis (
|
||
<figureCitation id="4CEB2A57FFF7FFEEE9D5FE643008FDEF" box="[1208,1315,506,533]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[189,1398,1130,1885]" captionTargetId="figure-328@23.[189,1398,1130,1885]" captionTargetPageId="23" captionText="Fig. 13. Anatomy of Mercuria balearica (Paladilhe, 1869). A, D, F. Stream at Sant Joan de Carbonell, Minorca, Spain. B, C, E. Venta El Pilar Spring, Málaga, Spain. A. Female genitalia. B. Penis and head. C. Prostate gland. D. Ctenidium and osphradium. E. Stomach. F. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867389" httpUri="https://zenodo.org/record/7867389/files/figure.png" pageId="23" pageNumber="24">Fig. 13C</figureCitation>
|
||
). Penis darkly pigmented, gradually tapering, attached to the neck behind the right eye; penial appendix and distal end of the penis nearly equal in size; penial appendix unpigmented or slightly pigmented at the junction with the penis, rounded, small, base narrow, medially positioned on the inner edge of the penis (
|
||
<figureCitation id="4CEB2A57FFF7FFEEEDA8FD19341AFD5B" box="[197,305,647,673]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[189,1398,1130,1885]" captionTargetId="figure-328@23.[189,1398,1130,1885]" captionTargetPageId="23" captionText="Fig. 13. Anatomy of Mercuria balearica (Paladilhe, 1869). A, D, F. Stream at Sant Joan de Carbonell, Minorca, Spain. B, C, E. Venta El Pilar Spring, Málaga, Spain. A. Female genitalia. B. Penis and head. C. Prostate gland. D. Ctenidium and osphradium. E. Stomach. F. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867389" httpUri="https://zenodo.org/record/7867389/files/figure.png" pageId="23" pageNumber="24">Fig. 13B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF7FFEEEDD0FD5736E8FC69" blockId="23.[189,1400,712,915]" pageId="23" pageNumber="24">
|
||
<smallCapsWord id="D289A00EFFF7FFEEEDD0FD57343FFD19" baselines="732,732" box="[189,276,713,739]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="23" pageNumber="24">FEMALE</smallCapsWord>
|
||
<smallCapsWord id="D289A00EFFF7FFEEEC4DFD5034A5FD1B" baselines="732" box="[288,398,718,737]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="23" pageNumber="24">GENITALIA</smallCapsWord>
|
||
. Glandular oviduct 2.5 times as long as wide; albumen gland longer than capsule gland (
|
||
<figureCitation id="4CEB2A57FFF7FFEEEC62FD723457FCFC" box="[271,380,748,774]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[189,1398,1130,1885]" captionTargetId="figure-328@23.[189,1398,1130,1885]" captionTargetPageId="23" captionText="Fig. 13. Anatomy of Mercuria balearica (Paladilhe, 1869). A, D, F. Stream at Sant Joan de Carbonell, Minorca, Spain. B, C, E. Venta El Pilar Spring, Málaga, Spain. A. Female genitalia. B. Penis and head. C. Prostate gland. D. Ctenidium and osphradium. E. Stomach. F. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867389" httpUri="https://zenodo.org/record/7867389/files/figure.png" pageId="23" pageNumber="24">Fig. 13A</figureCitation>
|
||
); bursa copulatrix pyriform (
|
||
<figureCitation id="4CEB2A57FFF7FFEEEFBDFD723675FCFC" box="[720,862,748,774]" captionStart="Fig" captionStartId="24.[189,232,1891,1917]" captionTargetBox="[189,1398,1087,1848]" captionTargetId="figure-272@24.[189,1398,1085,1848]" captionTargetPageId="24" captionText="Fig. 14. Variation of the bursa copulatrix of Mercuria balearica (Paladilhe, 1869). A. Elongate, stream at Sant Joan de Carbonell, Minorca, Spain. B. Pyriform, Venta El Pilar Spring, Málaga, Spain. C–D. Pyriform, Fuente Valentín Spring, Alozaina, Cádiz, Spain. E–G. Detailed views of the seminal receptacle (Sr) and renal oviduct loops." figureDoi="http://doi.org/10.5281/zenodo.7867391" httpUri="https://zenodo.org/record/7867391/files/figure.png" pageId="23" pageNumber="24">Fig. 14B–D</figureCitation>
|
||
) to elongate (
|
||
<figureCitation id="4CEB2A57FFF7FFEEE96DFD723146FCFC" box="[1024,1133,748,774]" captionStart="Fig" captionStartId="24.[189,232,1891,1917]" captionTargetBox="[189,1398,1087,1848]" captionTargetId="figure-272@24.[189,1398,1085,1848]" captionTargetPageId="24" captionText="Fig. 14. Variation of the bursa copulatrix of Mercuria balearica (Paladilhe, 1869). A. Elongate, stream at Sant Joan de Carbonell, Minorca, Spain. B. Pyriform, Venta El Pilar Spring, Málaga, Spain. C–D. Pyriform, Fuente Valentín Spring, Alozaina, Cádiz, Spain. E–G. Detailed views of the seminal receptacle (Sr) and renal oviduct loops." figureDoi="http://doi.org/10.5281/zenodo.7867391" httpUri="https://zenodo.org/record/7867391/files/figure.png" pageId="23" pageNumber="24">Fig. 14A</figureCitation>
|
||
), ca 2 times as long as wide (Supp. file 2: Table S12), lying against the albumen gland; bursal duct shorter than bursa length; renal oviduct unpigmented, coiled, with three loops; coils can be wide open (
|
||
<figureCitation id="4CEB2A57FFF7FFEEE92BFCAC3185FCB6" box="[1094,1198,818,844]" captionStart="Fig" captionStartId="24.[189,232,1891,1917]" captionTargetBox="[189,1398,1087,1848]" captionTargetId="figure-272@24.[189,1398,1085,1848]" captionTargetPageId="24" captionText="Fig. 14. Variation of the bursa copulatrix of Mercuria balearica (Paladilhe, 1869). A. Elongate, stream at Sant Joan de Carbonell, Minorca, Spain. B. Pyriform, Venta El Pilar Spring, Málaga, Spain. C–D. Pyriform, Fuente Valentín Spring, Alozaina, Cádiz, Spain. E–G. Detailed views of the seminal receptacle (Sr) and renal oviduct loops." figureDoi="http://doi.org/10.5281/zenodo.7867391" httpUri="https://zenodo.org/record/7867391/files/figure.png" pageId="23" pageNumber="24">Fig. 14E</figureCitation>
|
||
) or highly coiled (
|
||
<figureCitation id="4CEB2A57FFF7FFEEEDABFCCB347EFC8A" box="[198,341,853,880]" captionStart="Fig" captionStartId="24.[189,232,1891,1917]" captionTargetBox="[189,1398,1087,1848]" captionTargetId="figure-272@24.[189,1398,1085,1848]" captionTargetPageId="24" captionText="Fig. 14. Variation of the bursa copulatrix of Mercuria balearica (Paladilhe, 1869). A. Elongate, stream at Sant Joan de Carbonell, Minorca, Spain. B. Pyriform, Venta El Pilar Spring, Málaga, Spain. C–D. Pyriform, Fuente Valentín Spring, Alozaina, Cádiz, Spain. E–G. Detailed views of the seminal receptacle (Sr) and renal oviduct loops." figureDoi="http://doi.org/10.5281/zenodo.7867391" httpUri="https://zenodo.org/record/7867391/files/figure.png" pageId="23" pageNumber="24">Fig. 14F–G</figureCitation>
|
||
); seminal receptacle elongate, with a short duct, positioned on the distal part of the renal oviduct just above the junction with the bursal duct (
|
||
<figureCitation id="4CEB2A57FFF7FFEEEE45FCE7369CFC69" box="[808,951,889,915]" captionStart="Fig" captionStartId="24.[189,232,1891,1917]" captionTargetBox="[189,1398,1087,1848]" captionTargetId="figure-272@24.[189,1398,1085,1848]" captionTargetPageId="24" captionText="Fig. 14. Variation of the bursa copulatrix of Mercuria balearica (Paladilhe, 1869). A. Elongate, stream at Sant Joan de Carbonell, Minorca, Spain. B. Pyriform, Venta El Pilar Spring, Málaga, Spain. C–D. Pyriform, Fuente Valentín Spring, Alozaina, Cádiz, Spain. E–G. Detailed views of the seminal receptacle (Sr) and renal oviduct loops." figureDoi="http://doi.org/10.5281/zenodo.7867391" httpUri="https://zenodo.org/record/7867391/files/figure.png" pageId="23" pageNumber="24">Fig. 14E–G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF7FFEEEDD0FC253705FBE1" blockId="23.[189,1400,954,1051]" pageId="23" pageNumber="24">
|
||
<smallCapsWord id="D289A00EFFF7FFEEEDD0FC253408FC2F" baselines="974,974" box="[189,291,955,981]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Nervous" pageId="23" pageNumber="24">NERVOUS</smallCapsWord>
|
||
<smallCapsWord id="D289A00EFFF7FFEEEC41FC5E3455FC29" baselines="974" box="[300,382,960,979]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="system" pageId="23" pageNumber="24">SYSTEM</smallCapsWord>
|
||
. Pigmented, elongated (mean RPG ratio = 0.53; see Supp. file 2: Table S15); cerebral ganglia approximately equal in size; pleuro-supraoesophageal connective ca 9 times as long as pleuro-suboesophageal one (
|
||
<figureCitation id="4CEB2A57FFF7FFEEECD5FB9E370BFBE1" box="[440,544,1024,1051]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[189,1398,1130,1885]" captionTargetId="figure-328@23.[189,1398,1130,1885]" captionTargetPageId="23" captionText="Fig. 13. Anatomy of Mercuria balearica (Paladilhe, 1869). A, D, F. Stream at Sant Joan de Carbonell, Minorca, Spain. B, C, E. Venta El Pilar Spring, Málaga, Spain. A. Female genitalia. B. Penis and head. C. Prostate gland. D. Ctenidium and osphradium. E. Stomach. F. Perioesophageal ring." figureDoi="http://doi.org/10.5281/zenodo.7867389" httpUri="https://zenodo.org/record/7867389/files/figure.png" pageId="23" pageNumber="24">Fig. 13F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="80AF665AFFF7FFEEEDD0F8163194F813" ID-DOI="http://doi.org/10.5281/zenodo.7867389" ID-Zenodo-Dep="7867389" httpUri="https://zenodo.org/record/7867389/files/figure.png" pageId="23" pageNumber="24" startId="23.[189,232,1928,1954]" targetBox="[189,1398,1130,1885]" targetPageId="23">
|
||
<paragraph id="D46F36D2FFF7FFEEEDD0F8163194F813" blockId="23.[189,1399,1928,2025]" pageId="23" pageNumber="24">
|
||
<emphasis id="E6A4EAC0FFF7FFEEEDD0F8163431F858" bold="true" box="[189,282,1928,1954]" pageId="23" pageNumber="24">Fig. 13.</emphasis>
|
||
Anatomy of
|
||
<taxonomicName id="13D04D51FFF7FFEEECD9F8173646F859" authority="(Paladilhe, 1869)" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[436,877,1928,1955]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF7FFEEECD9F81737B2F858" box="[436,665,1928,1954]" italics="true" pageId="23" pageNumber="24">Mercuria balearica</emphasis>
|
||
(
|
||
<bibRefCitation id="B0414B23FFF7FFEEEFC6F816364EF859" author="Paladilhe A." box="[683,869,1928,1955]" pageId="23" pageNumber="24" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="E6A4EAC0FFF7FFEEEE10F81636B9F858" bold="true" box="[893,914,1928,1954]" pageId="23" pageNumber="24">A</emphasis>
|
||
,
|
||
<emphasis id="E6A4EAC0FFF7FFEEEECCF816369DF858" bold="true" box="[929,950,1928,1954]" pageId="23" pageNumber="24">D</emphasis>
|
||
,
|
||
<emphasis id="E6A4EAC0FFF7FFEEEEA8F81636FCF858" bold="true" box="[965,983,1928,1954]" pageId="23" pageNumber="24">F</emphasis>
|
||
. Stream at Sant Joan de Carbonell, Minorca, Spain.
|
||
<emphasis id="E6A4EAC0FFF7FFEEECECF83234BEF83C" bold="true" box="[385,405,1964,1990]" pageId="23" pageNumber="24">B</emphasis>
|
||
,
|
||
<emphasis id="E6A4EAC0FFF7FFEEECCEF8353493F83F" bold="true" box="[419,440,1963,1989]" pageId="23" pageNumber="24">C</emphasis>
|
||
,
|
||
<emphasis id="E6A4EAC0FFF7FFEEECAAF83234F0F83C" bold="true" box="[455,475,1964,1990]" pageId="23" pageNumber="24">E</emphasis>
|
||
. Venta El Pilar Spring, Málaga, Spain.
|
||
<emphasis id="E6A4EAC0FFF7FFEEEEC5F8353696F83F" bold="true" box="[936,957,1963,1989]" pageId="23" pageNumber="24">A</emphasis>
|
||
. Female genitalia.
|
||
<emphasis id="E6A4EAC0FFF7FFEEE9F0F832319AF83C" bold="true" box="[1181,1201,1964,1990]" pageId="23" pageNumber="24">B</emphasis>
|
||
. Penis and head.
|
||
<emphasis id="E6A4EAC0FFF7FFEEEDD0F85035F9F812" bold="true" box="[189,210,1998,2024]" pageId="23" pageNumber="24">C</emphasis>
|
||
. Prostate gland.
|
||
<emphasis id="E6A4EAC0FFF7FFEEECFBF8513480F813" bold="true" box="[406,427,1999,2025]" pageId="23" pageNumber="24">D</emphasis>
|
||
. Ctenidium and osphradium.
|
||
<emphasis id="E6A4EAC0FFF7FFEEEE65F8513637F813" bold="true" box="[776,796,1999,2025]" pageId="23" pageNumber="24">E</emphasis>
|
||
. Stomach.
|
||
<emphasis id="E6A4EAC0FFF7FFEEEECDF8513699F813" bold="true" box="[928,946,1999,2025]" pageId="23" pageNumber="24">F</emphasis>
|
||
. Perioesophageal ring.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="9CCA6559FFF8FFE1EDD0FE9737A1FD56" pageId="24" pageNumber="25" type="biology_ecology">
|
||
<paragraph id="D46F36D2FFF8FFE1EDD0FE9734D9FED8" blockId="24.[189,498,264,291]" box="[189,498,264,291]" pageId="24" pageNumber="25">
|
||
<heading id="8F2781BEFFF8FFE1EDD0FE9734D9FED8" bold="true" box="[189,498,264,291]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
|
||
<emphasis id="E6A4EAC0FFF8FFE1EDD0FE9734D9FED8" bold="true" box="[189,498,264,291]" pageId="24" pageNumber="25">Ecology and distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF8FFE1EDD0FEA6305CFDFB" blockId="24.[188,1399,311,514]" pageId="24" pageNumber="25">
|
||
<taxonomicName id="13D04D51FFF8FFE1EDD0FEA6348DFEAB" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[189,422,311,337]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF8FFE1EDD0FEA6348DFEAB" box="[189,422,311,337]" italics="true" pageId="24" pageNumber="25">Mercuria balearica</emphasis>
|
||
</taxonomicName>
|
||
presents broad ecological plasticity, inhabiting springs, streams and ponds with conductivities ranging from 179 to 2750 µS/cm. Population densities were low. During our field surveys, we found evidence of local extinctions in the regions of
|
||
<collectingRegion id="1614F830FFF8FFE1EEC4FEE3310BFE6D" box="[937,1056,381,407]" country="Spain" name="Andalucia" pageId="24" pageNumber="25">Andalusia</collectingRegion>
|
||
and
|
||
<collectingCountry id="ACC77642FFF8FFE1E93BFEE33192FE6D" box="[1110,1209,381,407]" name="Spain" pageId="24" pageNumber="25">Minorca</collectingCountry>
|
||
(e.g., extirpated populations in the towns of Zuheros, Arriate, Alozaina, Gaucín, Ferreries), suggesting that the species was more abundant in the late twentieth century. This species occurs sympatrically with
|
||
<taxonomicName id="13D04D51FFF8FFE1E9ACFE5A34DDFDFB" authority="Altaba, 2007" authorityName="Altaba" authorityYear="2007" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="meloussensis">
|
||
<emphasis id="E6A4EAC0FFF8FFE1E9ACFE5A347DFDFB" italics="true" pageId="24" pageNumber="25">Pseudamnicola meloussensis</emphasis>
|
||
Altaba, 2007
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13D04D51FFF8FFE1EC92FE793609FDFB" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[511,802,487,513]" class="Gastropoda" family="Neritidae" genus="Theodoxus" isUncertain="true" kingdom="Animalia" order="Cycloneritida" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">
|
||
<emphasis id="E6A4EAC0FFF8FFE1EC92FE793755FDFB" box="[511,638,487,513]" italics="true" pageId="24" pageNumber="25">Theodoxus</emphasis>
|
||
aff.
|
||
<emphasis id="E6A4EAC0FFF8FFE1EFD8FE793609FDFB" box="[693,802,487,513]" italics="true" pageId="24" pageNumber="25">fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="13D04D51FFF8FFE1EE5DFE7936C4FDF8" box="[816,1007,487,514]" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="E6A4EAC0FFF8FFE1EE5DFE79369CFDFB" box="[816,951,487,513]" italics="true" pageId="24" pageNumber="25">Melanopsis</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="E6A4EAC0FFF8FFE1E94BFE76305CFDFB" box="[1062,1399,487,513]" italics="true" pageId="24" pageNumber="25">
|
||
<taxonomicName id="13D04D51FFF8FFE1E94BFE763059FDFB" baseAuthorityName="Gray" baseAuthorityYear="1843" box="[1062,1394,487,513]" class="Gastropoda" family="Tateidae" genus="Potamopyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="antipodarum">Potamopyrgus antipodarum</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF8FFE1EDD0FDB637A1FD56" blockId="24.[189,1398,552,685]" pageId="24" pageNumber="25">
|
||
The species was previously cited from springs, streams, small rivers and ditches in the southern Iberian Peninsula and on the
|
||
<collectingRegion id="1614F830FFF8FFE1ECA4FDD237A2FD9C" box="[457,649,588,614]" country="Spain" name="Illes Baleares" pageId="24" pageNumber="25">Balearic Islands</collectingRegion>
|
||
of Ibiza and
|
||
<collectingCountry id="ACC77642FFF8FFE1EE59FDD236BCFD9C" box="[820,919,588,614]" name="Spain" pageId="24" pageNumber="25">Minorca</collectingCountry>
|
||
(
|
||
<bibRefCitation id="B0414B23FFF8FFE1EEC6FDD2317BFD9C" author="Boeters H." box="[939,1104,588,614]" pageId="24" pageNumber="25" pagination="181 - 261" refId="ref28493" refString="Boeters H. 1988. Moitessieriidae und Hydrobiidae in Spanien und Portugal (Gastropoda: Prosobranchia). Archiv fur Molluskenkunde 118: 181 - 261." type="journal article" year="1988">Boeters 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B0414B23FFF8FFE1E90CFDD23012FD9C" author="Gloer P. & Boeters H. D. & Walther F." box="[1121,1337,587,614]" pageId="24" pageNumber="25" pagination="279 - 291" refId="ref29897" refString="Gloer P., Boeters H. D. & Walther F. 2015. Species of the genus Mercuria Boeters, 1971 (Caenogastropoda: Truncatelloidea: Hydrobiidae) from the European Mediterranean region, Morocco and Madeira, with descriptions of new species. Folia Malacologica 23: 279 - 291. https: // doi. org / 10.12657 / folmal. 023.024" type="journal article" year="2015">
|
||
Glöer
|
||
<emphasis id="E6A4EAC0FFF8FFE1E9C2FDD231C5FD9C" box="[1199,1262,588,614]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
). We confirmed the species in
|
||
<collectingCountry id="ACC77642FFF8FFE1ECB3FDF1376AFD73" box="[478,577,623,649]" name="Spain" pageId="24" pageNumber="25">Minorca</collectingCountry>
|
||
(Ibiza was not included as a field site) and in several other springs and streams in south-eastern
|
||
<collectingCountry id="ACC77642FFF8FFE1EC8DFD0C370FFD56" box="[480,548,658,684]" name="Georgia" pageId="24" pageNumber="25">Iberia</collectingCountry>
|
||
(
|
||
<figureCitation id="4CEB2A57FFF8FFE1EF59FD0C3752FD56" box="[564,633,658,684]" captionStart="Fig" captionStartId="3.[189,232,1847,1873]" captionTargetBox="[222,1362,1278,1786]" captionTargetId="figure-341@3.[203,1385,1259,1809]" captionTargetPageId="3" captionText="Fig. 1. Phylogenetic clustering and geographic location of the studied populations of Mercuria Boeters, 1971 throughout the western Palaearctic region. The tree topology on the left has been modified from Miller et al. (2022) to show the species clades (i.e., coloured clades) of the populations described here. One population (grey dot) represents the distribution of M. maceana (Paladilhe, 1869). Scale bar below topology: expected change per site." figureDoi="http://doi.org/10.5281/zenodo.7867361" httpUri="https://zenodo.org/record/7867361/files/figure.png" pageId="24" pageNumber="25">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CCA6559FFF8FFE0EDD0FD4A314EF9B4" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="discussion">
|
||
<paragraph id="D46F36D2FFF8FFE1EDD0FD4A3404FD14" blockId="24.[189,303,724,750]" box="[189,303,724,750]" pageId="24" pageNumber="25">
|
||
<heading id="8F2781BEFFF8FFE1EDD0FD4A3404FD14" bold="true" box="[189,303,724,750]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
|
||
<emphasis id="E6A4EAC0FFF8FFE1EDD0FD4A3404FD14" bold="true" box="[189,303,724,750]" pageId="24" pageNumber="25">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF8FFE0EDD0FC9C368EFE0D" blockId="24.[189,1399,770,1008]" lastBlockId="25.[189,1400,265,503]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
The taxonomy of the species has undergone several revisions, with the most recent change occurring in 2019, when
|
||
<bibRefCitation id="B0414B23FFF8FFE1EC03FCBB371BFCC5" author="Quintana J." box="[366,560,805,831]" pageId="24" pageNumber="25" pagination="15 - 23" refId="ref30612" refString="Quintana J. 2019. Los moluscos del nivel B de la Cova Murada (Barranc d'Algendar, Ciutadella de Menorca, Illes Balears). Implicaciones tafonomicas. Bolleti de la Societat d'Historia Natural de les Balears 62: 15 - 23." type="journal article" year="2019">Quintana (2019)</bibRefCitation>
|
||
placed the species in the genus
|
||
<taxonomicName id="13D04D51FFF8FFE1EEC5FCBB303DFCC5" authority="Paulucci, 1878" authorityName="Paulucci" authorityYear="1878" box="[936,1302,805,831]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="E6A4EAC0FFF8FFE1EEC5FCBB3176FCC5" box="[936,1117,805,831]" italics="true" pageId="24" pageNumber="25">Pseudamnicola</emphasis>
|
||
Paulucci, 1878
|
||
</taxonomicName>
|
||
. He had found an abundant population living in Colarsega River at Port-Mahon (
|
||
<collectingCountry id="ACC77642FFF8FFE1E975FCD631F9FC99" box="[1048,1234,840,867]" name="Minorca Island" pageId="24" pageNumber="25">Minorca Island</collectingCountry>
|
||
), which is the
|
||
<typeStatus id="0B6B8870FFF8FFE1EDD3FCF235DAFC7C" box="[190,241,876,902]" pageId="24" pageNumber="25">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="13D04D51FFF8FFE1EC12FCF2373DFC7C" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[383,534,876,902]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF8FFE1EC12FCF2373DFC7C" box="[383,534,876,902]" italics="true" pageId="24" pageNumber="25">M. balearica</emphasis>
|
||
</taxonomicName>
|
||
. He sent a subsample to the MNCN in Madrid so that we could review the classification. We confirmed that all the specimens received belong to
|
||
<taxonomicName id="13D04D51FFF8FFE1E94CFC11305DFC53" authorityName="Altaba" authorityYear="2007" box="[1057,1398,911,937]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="meloussensis">
|
||
<emphasis id="E6A4EAC0FFF8FFE1E94CFC11305DFC53" box="[1057,1398,911,937]" italics="true" pageId="24" pageNumber="25">Pseudamnicola meloussensis</emphasis>
|
||
</taxonomicName>
|
||
and communicated to Quintana that more exhaustive sampling would be necessary to find specimens of
|
||
<taxonomicName id="13D04D51FFF8FFE1EDD0FC48347DFC15" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[189,342,981,1007]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF8FFE1EDD0FC48347DFC15" box="[189,342,981,1007]" italics="true" pageId="24" pageNumber="25">M. balearica</emphasis>
|
||
</taxonomicName>
|
||
. However, he thought that if all the specimens found in this locality are
|
||
<taxonomicName id="13D04D51FFF8FFE1E9D1FC4B3044FC15" authorityName="Paulucci" authorityYear="1878" box="[1212,1391,981,1007]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="E6A4EAC0FFF8FFE1E9D1FC4B3044FC15" box="[1212,1391,981,1007]" italics="true" pageId="24" pageNumber="25">Pseudamnicola</emphasis>
|
||
</taxonomicName>
|
||
, then
|
||
<taxonomicName id="13D04D51FFF9FFE0ED94FE9434B9FED9" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[249,402,265,291]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF9FFE0ED94FE9434B9FED9" box="[249,402,265,291]" italics="true" pageId="25" pageNumber="26">M. balearica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="13D04D51FFF9FFE0ECBFFE9437A1FED9" authorityName="Altaba" authorityYear="2007" box="[466,650,265,291]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="meloussensis">
|
||
<emphasis id="E6A4EAC0FFF9FFE0ECBFFE9437A1FED9" box="[466,650,265,291]" italics="true" pageId="25" pageNumber="26">P. meloussensis</emphasis>
|
||
</taxonomicName>
|
||
are probably the same species and they should be renamed as
|
||
<taxonomicName id="13D04D51FFF9FFE0EDD2FEB234C5FEBC" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[191,494,300,326]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EDD2FEB234C5FEBC" box="[191,494,300,326]" italics="true" pageId="25" pageNumber="26">Pseudamnicola balearica</emphasis>
|
||
</taxonomicName>
|
||
, the species described by
|
||
<bibRefCitation id="B0414B23FFF9FFE0EE5AFEB236D5FEBD" author="Paladilhe A." box="[823,1022,300,327]" pageId="25" pageNumber="26" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe (1869)</bibRefCitation>
|
||
(as
|
||
<taxonomicName id="13D04D51FFF9FFE0E954FEB2300EFEBC" authorityName="Paladilhe" authorityYear="1869" box="[1081,1317,300,326]" class="Gastropoda" family="Amnicolidae" genus="Amnicola" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF9FFE0E954FEB2300EFEBC" box="[1081,1317,300,326]" italics="true" pageId="25" pageNumber="26">Amnicola balearica</emphasis>
|
||
</taxonomicName>
|
||
), with
|
||
<taxonomicName id="13D04D51FFF9FFE0EDD0FECE3458FE93" authorityName="Altaba" authorityYear="2007" box="[189,371,335,361]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="meloussensis">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EDD0FECE3458FE93" box="[189,371,335,361]" italics="true" pageId="25" pageNumber="26">P. meloussensis</emphasis>
|
||
</taxonomicName>
|
||
consequently being a junior synonym of
|
||
<taxonomicName id="13D04D51FFF9FFE0EE34FECE36CDFE93" authorityName="Paladilhe" authorityYear="1869" box="[857,998,335,361]" class="Gastropoda" family="Amnicolidae" genus="Amnicola" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EE34FECE36CDFE93" box="[857,998,335,361]" italics="true" pageId="25" pageNumber="26">P. balearica</emphasis>
|
||
</taxonomicName>
|
||
. However, during a more detailed sampling of this population in 2018, we were able to collect a few specimens of
|
||
<taxonomicName id="13D04D51FFF9FFE0E9FCFEED3001FE76" authorityName="Paladilhe" authorityYear="1869" box="[1169,1322,370,396]" class="Gastropoda" family="Amnicolidae" genus="Amnicola" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF9FFE0E9FCFEED3001FE76" box="[1169,1322,370,396]" italics="true" pageId="25" pageNumber="26">M. balearica</emphasis>
|
||
</taxonomicName>
|
||
living together with
|
||
<taxonomicName id="13D04D51FFF9FFE0EC08FE093735FE4A" authorityName="Altaba" authorityYear="2007" box="[357,542,406,432]" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="meloussensis">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EC08FE093735FE4A" box="[357,542,406,432]" italics="true" pageId="25" pageNumber="26">P. meloussensis</emphasis>
|
||
</taxonomicName>
|
||
. The DNA-based sequences (
|
||
<bibRefCitation id="B0414B23FFF9FFE0EEEBFE083174FE4A" author="Miller J. P. & Delicado D. & Garcia-Guerrero F. & Ramos M. A." box="[902,1119,406,432]" pageId="25" pageNumber="26" pagination="107524" refId="ref30353" refString="Miller J. P., Delicado D., Garcia-Guerrero F. & Ramos M. A. 2022. Recurrent founder-event speciation across the Mediterranean likely shaped the species diversity and geographic distribution of the freshwater snail genus Mercuria Boeters, 1971 (Caenogastropoda: Hydrobiidae). Molecular Phylogenetics and Evolution 173: 107524. https: // doi. org / 10.1016 / j. ympev. 2022.107524" type="journal article" year="2022">
|
||
Miller
|
||
<emphasis id="E6A4EAC0FFF9FFE0EEB6FE093132FE4A" box="[987,1049,406,432]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
) and shell morphology (
|
||
<figureCitation id="4CEB2A57FFF9FFE0EDABFE273407FE29" box="[198,300,441,467]" captionStart="Fig" captionStartId="21.[189,232,1892,1918]" captionTargetBox="[189,1398,274,1842]" captionTargetId="figure-11@21.[189,1398,274,1842]" captionTargetPageId="21" captionText="Fig. 11. Shells and operculum of Mercuria balearica (Paladilhe, 1869). A–B, G–H. Stream at Sant Joan de Carbonell, Minorca, Spain. C. Colarsega River, near the Port of Mahon (type locality). D. Fuente Árabe Spring, Mojácar, Granada, Spain (unpigmented specimen). E. Venta El Pilar Spring, Málaga, Spain. F. Spring, Alozaina, Cádiz, Spain. G. Operculum, inner side. H. Operculum, outer side." figureDoi="http://doi.org/10.5281/zenodo.7867385" httpUri="https://zenodo.org/record/7867385/files/figure.png" pageId="25" pageNumber="26">Fig. 11C</figureCitation>
|
||
) of these specimens correspond to those of a member of
|
||
<taxonomicName id="13D04D51FFF9FFE0EED3FE243107FE29" authorityName="Boeters" authorityYear="1971" box="[958,1068,442,467]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EED3FE243107FE29" box="[958,1068,442,467]" italics="true" pageId="25" pageNumber="26">Mercuria</emphasis>
|
||
</taxonomicName>
|
||
, confirming their placement in this genus (and their continued presence at the
|
||
<typeStatus id="0B6B8870FFF9FFE0EE6FFE43361EFE0D" box="[770,821,477,503]" pageId="25" pageNumber="26">type</typeStatus>
|
||
locality).
|
||
</paragraph>
|
||
<caption id="80AF665AFFF8FFE1EDD0F8FD37A6F81D" ID-DOI="http://doi.org/10.5281/zenodo.7867391" ID-Zenodo-Dep="7867391" httpUri="https://zenodo.org/record/7867391/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,232,1891,1917]" targetBox="[189,1398,1087,1848]" targetPageId="24">
|
||
<paragraph id="D46F36D2FFF8FFE1EDD0F8FD37A6F81D" blockId="24.[189,1399,1891,2023]" pageId="24" pageNumber="25">
|
||
<emphasis id="E6A4EAC0FFF8FFE1EDD0F8FD340BF887" bold="true" box="[189,288,1891,1917]" pageId="24" pageNumber="25">Fig. 14.</emphasis>
|
||
Variation of the bursa copulatrix of
|
||
<taxonomicName id="13D04D51FFF8FFE1EF96F8FA31E1F884" authority="(Paladilhe, 1869)" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[763,1226,1891,1918]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF8FFE1EF96F8FA36CCF887" box="[763,999,1891,1917]" italics="true" pageId="24" pageNumber="25">Mercuria balearica</emphasis>
|
||
(
|
||
<bibRefCitation id="B0414B23FFF8FFE1EE90F8FD31EAF884" author="Paladilhe A." box="[1021,1217,1891,1918]" pageId="24" pageNumber="25" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="E6A4EAC0FFF8FFE1E9B1F8FD31DAF887" bold="true" box="[1244,1265,1891,1917]" pageId="24" pageNumber="25">A</emphasis>
|
||
. Elongate, stream at Sant Joan de Carbonell, Minorca, Spain.
|
||
<emphasis id="E6A4EAC0FFF8FFE1EE71F8193605F85B" bold="true" box="[796,814,1927,1953]" pageId="24" pageNumber="25">B</emphasis>
|
||
. Pyriform, Venta El Pilar Spring, Málaga, Spain.
|
||
<emphasis id="E6A4EAC0FFF8FFE1EDD3F83435DDF83E" bold="true" box="[190,246,1962,1988]" pageId="24" pageNumber="25">C–D</emphasis>
|
||
. Pyriform, Fuente Valentín Spring, Alozaina, Cádiz, Spain.
|
||
<emphasis id="E6A4EAC0FFF8FFE1EEA4F834312BF83E" bold="true" box="[969,1024,1962,1988]" pageId="24" pageNumber="25">E–G</emphasis>
|
||
. Detailed views of the seminal receptacle (Sr) and renal oviduct loops.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D46F36D2FFF9FFE0EDD0FD8031A1FD12" blockId="25.[188,1401,542,744]" pageId="25" pageNumber="26">
|
||
Our morphological observations (this study) and DNA results (
|
||
<bibRefCitation id="B0414B23FFF9FFE0EECCFD803158FDC2" author="Miller J. P. & Delicado D. & Garcia-Guerrero F. & Ramos M. A." box="[929,1139,542,568]" pageId="25" pageNumber="26" pagination="107524" refId="ref30353" refString="Miller J. P., Delicado D., Garcia-Guerrero F. & Ramos M. A. 2022. Recurrent founder-event speciation across the Mediterranean likely shaped the species diversity and geographic distribution of the freshwater snail genus Mercuria Boeters, 1971 (Caenogastropoda: Hydrobiidae). Molecular Phylogenetics and Evolution 173: 107524. https: // doi. org / 10.1016 / j. ympev. 2022.107524" type="journal article" year="2022">
|
||
Miller
|
||
<emphasis id="E6A4EAC0FFF9FFE0EE98FD80311BFDC2" box="[1013,1072,542,568]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
) confirm the previous assignments of other Iberian populations to
|
||
<taxonomicName id="13D04D51FFF9FFE0EFD1FDDC3678FDA1" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[700,851,577,603]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EFD1FDDC3678FDA1" box="[700,851,577,603]" italics="true" pageId="25" pageNumber="26">M. balearica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="B0414B23FFF9FFE0EE0EFDDF3128FDA1" author="Boeters H." box="[867,1027,577,603]" pageId="25" pageNumber="26" pagination="181 - 261" refId="ref28493" refString="Boeters H. 1988. Moitessieriidae und Hydrobiidae in Spanien und Portugal (Gastropoda: Prosobranchia). Archiv fur Molluskenkunde 118: 181 - 261." type="journal article" year="1988">Boeters 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B0414B23FFF9FFE0E962FDDF31F2FDA6" author="Gloer P. & Boeters H. D. & Walther F." box="[1039,1241,577,604]" pageId="25" pageNumber="26" pagination="279 - 291" refId="ref29897" refString="Gloer P., Boeters H. D. & Walther F. 2015. Species of the genus Mercuria Boeters, 1971 (Caenogastropoda: Truncatelloidea: Hydrobiidae) from the European Mediterranean region, Morocco and Madeira, with descriptions of new species. Folia Malacologica 23: 279 - 291. https: // doi. org / 10.12657 / folmal. 023.024" type="journal article" year="2015">
|
||
Glöer
|
||
<emphasis id="E6A4EAC0FFF9FFE0E935FDDC31B8FDA1" box="[1112,1171,577,603]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
). Populations from the stream near Sant Joan de Carbonell,
|
||
<collectingCountry id="ACC77642FFF9FFE0EF84FDFA3667FD84" box="[745,844,612,638]" name="Spain" pageId="25" pageNumber="26">Minorca</collectingCountry>
|
||
(MNCN 15.05/94745), Venta El Pilar Spring, Málaga (MNCN 15.05/94738) and Fuente Valentin Spring, Cádiz (MNCN 15.05/94739) accounted for most of the morphological variation observed between Minorcan and Iberian populations. Sequence divergence of COI among these populations was low (<1.2%) (
|
||
<bibRefCitation id="B0414B23FFF9FFE0EEC4FD503155FD12" author="Miller J. P. & Delicado D. & Garcia-Guerrero F. & Ramos M. A." box="[937,1150,718,744]" pageId="25" pageNumber="26" pagination="107524" refId="ref30353" refString="Miller J. P., Delicado D., Garcia-Guerrero F. & Ramos M. A. 2022. Recurrent founder-event speciation across the Mediterranean likely shaped the species diversity and geographic distribution of the freshwater snail genus Mercuria Boeters, 1971 (Caenogastropoda: Hydrobiidae). Molecular Phylogenetics and Evolution 173: 107524. https: // doi. org / 10.1016 / j. ympev. 2022.107524" type="journal article" year="2022">
|
||
Miller
|
||
<emphasis id="E6A4EAC0FFF9FFE0EE91FD50311DFD12" box="[1020,1078,718,744]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF9FFE0EDD0FC91305CFB09" blockId="25.[188,1401,783,1267]" pageId="25" pageNumber="26">
|
||
Intraspecific morphological differences for this species are observed mainly in shell, penis and female genitalia features. Specimens from
|
||
<collectingCountry id="ACC77642FFF9FFE0EF0BFCAC37E2FCB6" box="[614,713,818,844]" name="Spain" pageId="25" pageNumber="26">Minorca</collectingCountry>
|
||
are generally larger in size and present globose to high-spired shells, whereas the Iberian specimens have only globose shells (Supp. file 2: Table S10). The specimens from Los Granados Trough in Málaga (MNCN 15.05/94735) have the smallest shells. All the specimens have a pigmented epithelium, though some specimens from Arabic Spring in Mojácar (MNCN 15.05/94741) are unpigmented. Variation in the male reproductive organs is as previously described for the genus: the penis is either slightly longer or shorter than the appendix, and the appendix is usually rounded and unpigmented (only the population in Peal del Becerro, Jaén presents a slightly pigmented appendix). The bursa copulatrix varies from pyriform in most of the populations to elongate in the specimens from Venta El Pilar Spring, Málaga (MNCN 15.05/94734). Populations from
|
||
<collectingCountry id="ACC77642FFF9FFE0E87EFBD2305DFB9C" box="[1299,1398,1100,1126]" name="Spain" pageId="25" pageNumber="26">Minorca</collectingCountry>
|
||
have both morphotypes. Females from Venta El Pilar Spring (MNCN 15.05/94734), the stream in Montecorto (MNCN 15.05/94736), Fuente Valentín Spring (MNCN 15.05/94739) and Arabic Spring (MNCN 15.05/94741) have a bursa copulatrix that surrounds the albumen gland, which is located beneath it (see
|
||
<figureCitation id="4CEB2A57FFF9FFE0EC06FB4734F9FB09" box="[363,466,1241,1267]" captionStart="Fig" captionStartId="24.[189,232,1891,1917]" captionTargetBox="[189,1398,1087,1848]" captionTargetId="figure-272@24.[189,1398,1085,1848]" captionTargetPageId="24" captionText="Fig. 14. Variation of the bursa copulatrix of Mercuria balearica (Paladilhe, 1869). A. Elongate, stream at Sant Joan de Carbonell, Minorca, Spain. B. Pyriform, Venta El Pilar Spring, Málaga, Spain. C–D. Pyriform, Fuente Valentín Spring, Alozaina, Cádiz, Spain. E–G. Detailed views of the seminal receptacle (Sr) and renal oviduct loops." figureDoi="http://doi.org/10.5281/zenodo.7867391" httpUri="https://zenodo.org/record/7867391/files/figure.png" pageId="25" pageNumber="26">Fig. 14C</figureCitation>
|
||
) rather than parallel to it (the typical orientation in other members of the genus).
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFF9FFE0EDD0FA85314EF9B4" blockId="25.[188,1400,1306,1615]" pageId="25" pageNumber="26">
|
||
<taxonomicName id="13D04D51FFF9FFE0EDD0FA85348AFACE" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[189,417,1306,1332]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EDD0FA85348AFACE" box="[189,417,1306,1332]" italics="true" pageId="25" pageNumber="26">Mercuria balearica</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="13D04D51FFF9FFE0EF8BFA853670FACE" baseAuthorityName="Draparnaud" baseAuthorityYear="1805" box="[742,859,1306,1332]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="similis">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EF8BFA853670FACE" box="[742,859,1306,1332]" italics="true" pageId="25" pageNumber="26">M. similis</emphasis>
|
||
</taxonomicName>
|
||
by its smaller, less globose shell, less convex body whorl, shorter tapering triangular penis with a rounded unpigmented penial appendix that is slightly larger than or equal in size to the penis, pyriform bursa copulatrix and higher number of cusps on the central and lateral teeth of the radula.
|
||
<taxonomicName id="13D04D51FFF9FFE0EF02FA1A367AFA64" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[623,849,1412,1438]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EF02FA1A367AFA64" box="[623,849,1412,1438]" italics="true" pageId="25" pageNumber="26">Mercuria balearica</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="13D04D51FFF9FFE0EEBFFA1A3154FA64" authority="Miller & Delicado & García-Guerrero & Khalloufi & Ramos, 2023" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[978,1151,1412,1438]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="carrillorum" status="sp. nov.">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EEBFFA1A3154FA64" box="[978,1151,1412,1438]" italics="true" pageId="25" pageNumber="26">M. carrillorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD9757BBFFF9FFE0E9EAFA1A31CBFA64" box="[1159,1248,1412,1438]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="13D04D51FFF9FFE0E87BFA1A305DFA64" authority="Miller & Delicado & García-Guerrero & Khalloufi & Ramos, 2023" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[1302,1398,1412,1438]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="felixi" status="sp. nov.">
|
||
<emphasis id="E6A4EAC0FFF9FFE0E87BFA1A305DFA64" box="[1302,1398,1412,1438]" italics="true" pageId="25" pageNumber="26">M. felixi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD9757BBFFF9FFE0EDD0FA393432FA3B" box="[189,281,1447,1473]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in shell shape (ovate-conic) and colour but differs from these congeners by having a shorter, more pigmented penis with a larger rounded penial appendix. Also, their distribution areas do not overlap. Sequence divergences for COI indicated that
|
||
<taxonomicName id="13D04D51FFF9FFE0EFBFFA703643F9FD" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[722,872,1517,1543]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="balearica">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EFBFFA703643F9FD" box="[722,872,1517,1543]" italics="true" pageId="25" pageNumber="26">M. balearica</emphasis>
|
||
</taxonomicName>
|
||
is most closely related to
|
||
<taxonomicName id="13D04D51FFF9FFE0E9F0FA70306EF9FD" authorityName="Boulaassafer, Ghamizi & Delicado" authorityYear="2018" box="[1181,1349,1517,1543]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="tensiftensis">
|
||
<emphasis id="E6A4EAC0FFF9FFE0E9F0FA70306EF9FD" box="[1181,1349,1517,1543]" italics="true" pageId="25" pageNumber="26">M. tensiftensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="13D04D51FFF9FFE0EDD3F98F3436F9D0" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[190,285,1552,1578]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="felixi">
|
||
<emphasis id="E6A4EAC0FFF9FFE0EDD3F98F3436F9D0" box="[190,285,1552,1578]" italics="true" pageId="25" pageNumber="26">M. felixi</emphasis>
|
||
</taxonomicName>
|
||
(5.2% mean divergence with both species) and most distantly to
|
||
<taxonomicName id="13D04D51FFF9FFE0E96EF98F318CF9D0" authorityName="Boulaassafer, Ghamizi & Delicado" authorityYear="2018" box="[1027,1191,1552,1578]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="midarensis">
|
||
<emphasis id="E6A4EAC0FFF9FFE0E96EF98F318CF9D0" box="[1027,1191,1552,1578]" italics="true" pageId="25" pageNumber="26">M. midarensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="13D04D51FFF9FFE0E9B1F98F34AFF9B5" authority="(Paladilhe, 1869)" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="melitensis">
|
||
<emphasis id="E6A4EAC0FFF9FFE0E9B1F98F305EF9D0" box="[1244,1397,1552,1578]" italics="true" pageId="25" pageNumber="26">M. melitensis</emphasis>
|
||
(
|
||
<bibRefCitation id="B0414B23FFF9FFE0EDA8F9AA3456F9B5" author="Paladilhe A." box="[197,381,1588,1615]" pageId="25" pageNumber="26" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(8.6% mean divergence with both species) (
|
||
<bibRefCitation id="B0414B23FFF9FFE0EEEAF9AA3173F9B4" author="Miller J. P. & Delicado D. & Garcia-Guerrero F. & Ramos M. A." box="[903,1112,1588,1614]" pageId="25" pageNumber="26" pagination="107524" refId="ref30353" refString="Miller J. P., Delicado D., Garcia-Guerrero F. & Ramos M. A. 2022. Recurrent founder-event speciation across the Mediterranean likely shaped the species diversity and geographic distribution of the freshwater snail genus Mercuria Boeters, 1971 (Caenogastropoda: Hydrobiidae). Molecular Phylogenetics and Evolution 173: 107524. https: // doi. org / 10.1016 / j. ympev. 2022.107524" type="journal article" year="2022">
|
||
Miller
|
||
<emphasis id="E6A4EAC0FFF9FFE0EEB5F9AA3139F9B4" box="[984,1042,1588,1614]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |