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<document ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" ID-Pensoft-Pub="2511-6428-1-13" ID-Pensoft-UUID="F4C1C6A2DFD95D3BAF5AF1CE09226837" ID-ZooBank="589077DFF8BB45ABB30D7E0DE1C307FE" ModsDocID="2511-6428-74-1-13" checkinTime="1706792588976" checkinUser="pensoft" docAuthor="Irmler, Ulrich" docDate="2024" docId="5C61E69402DA58B2B358E867E0E50D80" docLanguage="en" docName="ContribEntomol 74(1): 13-34" docOrigin="Contributions to Entomology 74 (1)" docPubDate="2024-01-31" docSource="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" docTitle="Lithocharodes montanus Irmler 2024, sp. nov." docType="treatment" docUuid="4D2C1D93-B5D7-4CDE-A3CA-4075FA65E66C" docUuidSource="ZooBank" docVersion="1" id="F4C1C6A2DFD95D3BAF5AF1CE09226837" lastPageNumber="13" masterDocId="F4C1C6A2DFD95D3BAF5AF1CE09226837" masterDocTitle="Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)" masterLastPageNumber="34" masterPageNumber="13" pageNumber="13" updateTime="1706792588976" updateUser="pensoft">
<mods:mods id="F1DEE7D0A93082094F585364CF8A8476" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DE07950CE09AA520889B8A12A2C9310D">
<mods:title id="4106D7D43930AA8FF89F54911FD7E0B0">Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)</mods:title>
</mods:titleInfo>
<mods:name id="1DD16836BC1FD5342BA1635C4E2F7561" type="personal">
<mods:role id="8E82F061C1E9F9FD3B47D6FF86221DBB">
<mods:roleTerm id="3105C9AF28A84AE6E163CC505C2E0BD9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4C939881443EE1C6EB1FCD0E5149B4ED">Irmler, Ulrich</mods:namePart>
<mods:affiliation id="384E7DDEA60834EA880820695DDF290C">Institute for Ecosystem Research, University of Kiel, Olshausenstrasse 40, 24098 Kiel, Germany</mods:affiliation>
<mods:nameIdentifier id="8389C8FB2C0B24B92CAA1E42A388AC20" type="email">uirmler@ecology.uni-kiel.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="20956E1321A13ED144AD772C4FB93A0C">text</mods:typeOfResource>
<mods:relatedItem id="5B190B489E282865BCA8DA7AFFE6095C" type="host">
<mods:titleInfo id="B356EF5BB837DECF801482DEC5EF7123">
<mods:title id="CFE469859654A28C641FF96A41BCCFDB">Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="97AFA4C6E036EE239C46CDF623693260">
<mods:date id="2E5574549608CFB0C1E81B3ECF688364">2024</mods:date>
<mods:detail id="FD35C226DD495905AC4B6C38CC6CFCF5" type="pubDate">
<mods:number id="2BB3C3AEE5239FACD2C0FF7319206EEF">2024-01-31</mods:number>
</mods:detail>
<mods:detail id="E994B43B9E67F32DB6D9748885DBAE6E" type="volume">
<mods:number id="B67D97ADA587856AE0E0A6E5B4EC2BA8">74</mods:number>
</mods:detail>
<mods:detail id="61E773586738BDF54D0BD87A72243CB9" type="issue">
<mods:number id="D7127E4E3A94D6A9C4125617A8BE2544">1</mods:number>
</mods:detail>
<mods:extent id="E8E00577E5E52D017E216E865CCB4403" unit="page">
<mods:start id="398C231AC581E8D49B54ECA2B3B818A0">13</mods:start>
<mods:end id="0AACF4C99646368A713943FE45A20569">34</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="39F1ADD5B7A857F068F646E3A789C9C5">
<mods:url id="EF9DBFB493763C302E1A8BAF5C97B37D">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:url>
</mods:location>
<mods:classification id="8B99550EBBA424A13A3F9EDA22FD6E31">journal article</mods:classification>
<mods:identifier id="2E9B874338D3196B5DF5BE3ADF35299C" type="DOI">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:identifier>
<mods:identifier id="D4E209B37664963CC39C213A3EA08879" type="Pensoft-Pub">2511-6428-1-13</mods:identifier>
<mods:identifier id="585581133352E4850D35C12288BF6B21" type="ZooBank">589077DFF8BB45ABB30D7E0DE1C307FE</mods:identifier>
<mods:identifier id="D8256091E1140DF49A452BB0FC963246" type="Pensoft-UUID">F4C1C6A2DFD95D3BAF5AF1CE09226837</mods:identifier>
</mods:mods>
<treatment id="5C61E69402DA58B2B358E867E0E50D80" LSID="urn:lsid:zoobank.org:act:4D2C1D93-B5D7-4CDE-A3CA-4075FA65E66C" httpUri="http://treatment.plazi.org/id/5C61E69402DA58B2B358E867E0E50D80" lastPageNumber="13" pageId="0" pageNumber="13">
<subSubSection id="2D7B23DBE943749B89C724F161EAC083" pageId="0" pageNumber="13" type="nomenclature">
<paragraph id="CAF037D821ED76CA46A6C121318F062F" pageId="0" pageNumber="13">
<taxonomicName id="B480A272C53BA1A1B8CF7A0B4145F431" LSID="https://zoobank.org/4D2C1D93-B5D7-4CDE-A3CA-4075FA65E66C" authority="Irmler, 2024" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Lithocharodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithocharodes montanus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="montanus" status="sp. nov.">Lithocharodes montanus</taxonomicName>
<taxonomicNameLabel id="AB8EEEC5365CCEF255A2DFF2D3CF0D9C" pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9D4B3A4F08259D15CA91B63A725A4782" pageId="0" pageNumber="13" type="description">
<paragraph id="D6474FC8FD7DD19C57D83BFDCDC131B8" pageId="0" pageNumber="13">
<figureCitation id="8DD991C72220B62BE18C2A839A51870C" captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. 5. Lithocharodes hansoni; 6. L. hibbsi; 7. L. lituratus; 8. L. montanus. a. Aedeagus; b. Paramere; c. Sternite VII; d. Tergite VII. Scale bars: 0.5 mm (a, c, d); 0.1 mm (b)." figureDoi="10.3897/contrib.entomol.74.e114543.figures5-8" httpUri="https://binary.pensoft.net/fig/975716" pageId="0" pageNumber="13">Figs 8a-d</figureCitation>
<figureCitation id="D0DC018229E77A4C085D5F9D079847E3" captionStart="Figure 20" captionStartId="F7" captionText="Figure 20. A. Fore-body of Lithocharodes lituratus; B. L. montanus; C. L. parallelus; D. L. zamorae; E. Somoleptus alajuelae; F. S. curtioculatus. Scale bars: 0.5 mm." figureDoi="10.3897/contrib.entomol.74.e114543.figure20" httpUri="https://binary.pensoft.net/fig/975721" pageId="0" pageNumber="13">, 20B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="27D7B84CBAB5174CA54550DBF2C749A3" pageId="0" pageNumber="13" type="materials_examined">
<paragraph id="E4F9310BBE9FA512558FE71F9FB9C6C4" pageId="0" pageNumber="13">Material examined.</paragraph>
<paragraph id="9CEEB46FC7E2913E99239EE7B15A8FDA" pageId="0" pageNumber="13">
<materialsCitation id="5FFB13661198754B4D8AF71EA935A249" collectingDate="1999-10-29" collectorName="Cecropia, Anderson" country="ECUADOR" elevation="1600" latitude="0.09277778" location="Maquepucuna Biological Station" longLatPrecision="21" longitude="-78.626945" specimenCount="1" stateProvince="Pichincha" typeStatus="Holotype">
<emphasis id="6A3F100285B5A81053BCD85D81320DBF" bold="true" italics="true" pageId="0" pageNumber="13">
<typeStatus id="C685E987A304918563B8460CC7374025" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="C3B93DB42CCB7931B7F3E0A09472712A" name="Ecuador">ECUADOR</collectingCountry>
male;
<collectingRegion id="8F361A5C0273513A3CE4F1CC0BC1FE78" country="Ecuador" name="Pichincha">Pichincha</collectingRegion>
;
<location id="B0CF6803D8B1637AA6EE559C986620DF" LSID="urn:lsid:plazi:treatment:5C61E69402DA58B2B358E867E0E50D80:B0CF6803D8B1637AA6EE559C986620DF" country="ECUADOR" latitude="0.09277778" longLatPrecision="21" longitude="-78.626945" name="Maquepucuna Biological Station" stateProvince="Pichincha">Maquepucuna Biological Station</location>
;
<geoCoordinate id="104AEA8DB6A324C03E29DA9CA740AF5C" degrees="0" direction="north" minutes="5" orientation="latitude" precision="15" seconds="34" value="0.09277778">0°5'34&quot;N</geoCoordinate>
,
<geoCoordinate id="A1659DE35AF2BC8BF5109A7FFE4CACE3" degrees="78" direction="west" minutes="37" orientation="longitude" precision="15" seconds="37" value="-78.626945">78°37'37&quot;W</geoCoordinate>
;
<quantity id="9005FF8FF2C33A9209A3A3ECB394C04E" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
<elevation id="8D3F79C88C2896E4B789534129DD4ECA" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 m</elevation>
</quantity>
elevation; ridgetop montane forest litter; mixed
<collectorName id="6CAF5F4C8886BAAFDA7052B45B7E418B">Cecropia</collectorName>
/cloud forest;
<collectingDate id="836644B29214D5BAE60C1D7316A62FBE" value="1999-10-29">29 Oct 1999</collectingDate>
; R,
<collectorName id="7128A2D675976917C4B16BA164FF7AD8">Anderson</collectorName>
leg.; KNHM ECU1A99 214B.
</materialsCitation>
</paragraph>
<paragraph id="D0D8972C454745895FD631073DB568BB" pageId="0" pageNumber="13">
<materialsCitation id="102D892747BE1031FD13EBDCFF9428CB" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="Cecropia &amp; Anderson" country="ECUADOR" elevation="1600" latitude="0.09277778" location="ECUADOR" longLatPrecision="21" longitude="-78.626945" specimenCount="2" specimenCount-female="2" stateProvince="Pichincha" typeStatus="Paratypes">
<emphasis id="E6BFEED166E79F1259DF0FECF49A4872" bold="true" italics="true" pageId="0" pageNumber="13">
<typeStatus id="E2FF6B311DC374BFCB49E27289BA2539" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="3879EAAEE601BE64F4521BEE0CAA7957" name="Ecuador">ECUADOR</collectingCountry>
<specimenCount id="5E9DEF57300EB5D1BEAFF83FB87027F3" count="2" type="female">2 females</specimenCount>
; same data as holotype KNHM ECU1A99 214B
</materialsCitation>
;
<materialsCitation id="5391EEF192E3BBA642FCA275CEB2FBCC" collectingDate="1999-10-27" collectingDateMax="1999-10-29" collectingDateMin="1999-10-27" collectingMethod="flight intercept trap" collectorName="Z. H. Falin" country="Ecuador" elevation="1275" latitude="0.122777775" location="Principal Trail" longLatPrecision="21" longitude="-78.65" municipality="Maquepucuna Biological Station" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha" typeStatus="paratype">
<specimenCount id="B3B9A5493A8DFFE66EE0664682006085" count="1" type="male">1 male</specimenCount>
;
<collectingRegion id="87FBBD6944E2B65AAF6B54569EAA4089" country="Ecuador" name="Pichincha">Pichincha</collectingRegion>
;
<collectingMunicipality id="73A3CD594732F73A130B099034C69FB1">Maquepucuna Biological Station</collectingMunicipality>
;
<location id="018E2184BE59A247BD55528F14FE46DB" LSID="urn:lsid:plazi:treatment:5C61E69402DA58B2B358E867E0E50D80:018E2184BE59A247BD55528F14FE46DB" country="Ecuador" latitude="0.122777775" longLatPrecision="21" longitude="-78.65" municipality="Maquepucuna Biological Station" name="Principal Trail" stateProvince="Pichincha">Principal Trail</location>
;
<geoCoordinate id="AA58BAE3EC1EE03C2D2FB84EAC372DA5" degrees="0" direction="north" minutes="7" orientation="latitude" precision="15" seconds="22" value="0.122777775">0°7'22&quot;N</geoCoordinate>
,
<geoCoordinate id="F653E7A3160B1971B86159EE1EAA5B6F" degrees="78" direction="west" minutes="39" orientation="longitude" precision="15" seconds="0" value="-78.65">78°39'0&quot;W</geoCoordinate>
;
<quantity id="EF39EBAAF2D6217CF929600E1D9BD41B" metricMagnitude="3" metricUnit="m" metricValue="1.275" unit="m" value="1275.0">
<elevation id="DFD6CB91188FCB4A254F1EBBB3910EAB" metricMagnitude="3" metricUnit="m" metricValue="1.275" unit="m" value="1275.0">1275 m</elevation>
</quantity>
elevation;
<collectingMethod id="350539E08AF45D5BE78B87EA6B79087C">flight intercept trap</collectingMethod>
;
<collectingDate id="94A6B49F389B255768D16BEC782023B6" value="1999-10-27" valueMax="1999-10-29" valueMin="1999-10-27">27-29 Oct 1999</collectingDate>
;
<collectorName id="ECF2BF20A56EAE184F4DAEF2D379E841">Z.H. Falin</collectorName>
leg; KNHM ECU1F00 048
</materialsCitation>
;
<materialsCitation id="3B6F61074E1E42517E179574E01EE190" collectingDate="1999-10-27" collectingDateMax="1999-10-29" collectingDateMin="1999-10-27" collectorName="R. Anderson" country="Ecuador" elevation="1200" latitude="0.11666667" location="Principal Trail" longLatPrecision="21" longitude="-78.634995" municipality="Maquepucuna Biological Station" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha" typeStatus="paratype">
<specimenCount id="F23A21D9A0F34BF36726EE404BA9B746" count="1" type="female">1 female</specimenCount>
, same location but;
<geoCoordinate id="F78B3449A5B3F72A03D480D8223839DA" degrees="0" direction="north" minutes="7" orientation="latitude" precision="15" seconds="0" value="0.11666667">0°7'0&quot;N</geoCoordinate>
,
<geoCoordinate id="E32CCDF0AF7410AAA53975E9BDC8755F" degrees="78" direction="west" minutes="38" orientation="longitude" precision="15" seconds="6" value="-78.634995">78°38'6&quot;W</geoCoordinate>
;
<quantity id="A09AF2E5EB316FBDCAB97949C2018DD4" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<elevation id="AF01D4166BC58571F04893DFF13505BB" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
</quantity>
elevation; montane evergreen forest litter;
<collectingDate id="F649FF74799E9D96A82226C2D2E0AAD8" value="1999-10-27">27 Oct 1999</collectingDate>
;
<collectorName id="84D64B1EEFCA4B34E6B8CCF58FCC80A3">R. Anderson</collectorName>
leg.; KNHM ECU1A99 208D
</materialsCitation>
;
<materialsCitation id="EE403D3E9120310690056D34E95947BF" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="R. Anderson" country="Ecuador" elevation="1200" latitude="0.11666667" location="Principal Trail" longLatPrecision="21" longitude="-78.634995" municipality="Maquepucuna Biological Station" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha" typeStatus="paratype">
<specimenCount id="8CA4EE0C7D14ACB0F4CD264CEEAFD3C6" count="1" type="female">1 female</specimenCount>
; same data; UIC ECU1A99 208D
</materialsCitation>
;
<materialsCitation id="24C30688AD78C0559006864C322E52FF" collectingDate="1999-10-29" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="R. Anderson" country="Ecuador" elevation="1620" latitude="0.09277778" location="Principal Trail" longLatPrecision="21" longitude="-78.626945" municipality="Maquepucuna Biological Station" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha" typeStatus="paratype">
<specimenCount id="5F510E0B7D920B9FFD8C79B790B16DC2" count="1" type="female">1 female</specimenCount>
, same location but;
<geoCoordinate id="AC5BB352AF2DFA922138D733432E92DE" degrees="0" direction="north" minutes="5" orientation="latitude" precision="15" seconds="34" value="0.09277778">0°5'34&quot;N</geoCoordinate>
,
<geoCoordinate id="5D3E4AFCCDE9827518480B416BD7709A" degrees="78" direction="west" minutes="37" orientation="longitude" precision="15" seconds="37" value="-78.626945">78°37'37&quot;W</geoCoordinate>
;
<quantity id="4E038002DD1B76501A1C74B100E33278" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" unit="m" value="1620.0">
<elevation id="4F9896AD3901D7ADB49F89988F350C57" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" unit="m" value="1620.0">1620 m</elevation>
</quantity>
elevation; ridgetop montane forest litter;
<collectingDate id="4E3247381B0007D7BC8B40685AE895DF" value="1999-10-29">29 Oct 1999</collectingDate>
;
<collectorName id="4E79B3C1144A4E8DAA6898EF4B66340B">R. Anderson</collectorName>
leg.; KNHM ECU1A99 213E
</materialsCitation>
,
<materialsCitation id="EF71B8603380E33F07A23EEA7BE296D8" collectingDate="1999-10-27" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="R. Anderson" country="Ecuador" elevation="1480" latitude="0.10694445" location="Principal Trail" longLatPrecision="21" longitude="-78.621666" municipality="Maquepucuna Biological Station" specimenCount="2" specimenCount-female="2" stateProvince="Pichincha" typeStatus="paratype">
<specimenCount id="28F89A4FD8930DE589779C55F3FE8984" count="2" type="female">2 females</specimenCount>
; same location but;
<geoCoordinate id="4AF034E568A85B41A3D24D4E2160268F" degrees="0" direction="north" minutes="6" orientation="latitude" precision="15" seconds="25" value="0.10694445">0°6'25&quot;N</geoCoordinate>
,
<geoCoordinate id="B25FF8324569A838750BD02A886BC042" degrees="78" direction="west" minutes="37" orientation="longitude" precision="15" seconds="18" value="-78.621666">78°37'18&quot;W</geoCoordinate>
; montane evergreen forest litter;
<quantity id="857DC3C987BDB35D17E006360CD7C89A" metricMagnitude="3" metricUnit="m" metricValue="1.48" unit="m" value="1480.0">
<elevation id="B81B58544F06EB6176D2BC857CFE5D7E" metricMagnitude="3" metricUnit="m" metricValue="1.48" unit="m" value="1480.0">1480 m</elevation>
</quantity>
elevation;
<collectingDate id="BFC5F82D397FD29915121537DFA22C90" value="1999-10-27">27 Oct 1999</collectingDate>
;
<collectorName id="30E2EBE3518C5869D5B0E3EAD79FB1A9">R. Anderson</collectorName>
leg.; KNHM ECU1A99 209A
</materialsCitation>
;
<materialsCitation id="2D8E894E5C74A604A1448F28AC0540A7" collectingDate="1991-12-23" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="C. Carlton, R. Leschen" country="Ecuador" elevation="1300" latitude="0.10694445" location="50 km NW Quito" longLatPrecision="21" longitude="-78.621666" municipality="Maquepucuna Biological Station" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha" typeStatus="paratype">
<specimenCount id="B3C527B54F6D1D9D6BFD8C4BE27A9845" count="1" type="female">1 female</specimenCount>
; same location but;
<location id="DF626833340A09DE3D4112F95D7B8C3C" LSID="urn:lsid:plazi:treatment:5C61E69402DA58B2B358E867E0E50D80:DF626833340A09DE3D4112F95D7B8C3C" country="Ecuador" latitude="0.10694445" longLatPrecision="21" longitude="-78.621666" name="50 km NW Quito" stateProvince="Pichincha">
<quantity id="7AD2B3456C62FACF956361C46932BA32" metricMagnitude="4" metricUnit="m" metricValue="5.0" unit="km" value="50.0">50 km</quantity>
NW Quito
</location>
;
<quantity id="2AE666E1408C2870316957FD49FE7CC0" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
<elevation id="7151E9701ADAD4CF35B518F0F6E68930" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</elevation>
</quantity>
elevation; banana duff; berlesate;
<collectingDate id="81700CAD8FA78C134B865D5BCE2427F5" value="1991-12-23">23 Dec 1991</collectingDate>
;
<collectorName id="29F5E1B816F923BC1D9063DA29CE9A1C">C. Carlton</collectorName>
,
<collectorName id="61EC0F8F0EB573953905C6FDC935712F">R. Leschen</collectorName>
leg.; KNHM #72 (KNHM)
</materialsCitation>
;
<materialsCitation id="43018D985DA1449E2816150EAE02ADC5" collectingDate="1996-06-05" collectingDateMax="1996-07-07" collectingDateMin="1996-06-05" collectingMethod="flight intercept trap" collectorName="P. Hibbs" country="Ecuador" latitude="-0.33333334" location="Bilsa" longLatPrecision="21" longitude="-79.71667" specimenCount="1" specimenCount-female="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCount id="5A8AD0DFAF85628FAD9280485289CC17" count="1" type="female">1 female</specimenCount>
;
<collectingRegion id="B7D5212FFC84BD20E4D62F2685BD8740" country="Ecuador" name="Esmeraldas">Esmeraldas</collectingRegion>
;
<location id="88F70E5764258AA2384513FD2C4B7E2F" LSID="urn:lsid:plazi:treatment:5C61E69402DA58B2B358E867E0E50D80:88F70E5764258AA2384513FD2C4B7E2F" country="Ecuador" latitude="-0.33333334" longLatPrecision="21" longitude="-79.71667" name="Bilsa" stateProvince="Esmeraldas">Bilsa</location>
;
<geoCoordinate id="B6EA2A7729DBF86D96FC518213C74390" degrees="0" direction="south" minutes="20" orientation="latitude" precision="15" seconds="0" value="-0.33333334">0°20'0&quot;S</geoCoordinate>
,
<geoCoordinate id="27B210EDC52767EB68EF8BC6067AFA3B" degrees="79" direction="west" minutes="43" orientation="longitude" precision="15" seconds="0" value="-79.71667">79°43'0&quot;W</geoCoordinate>
;
<collectingMethod id="3213E03E254E2CC50BBBF5CE8C7CBC7D">flight intercept trap</collectingMethod>
;
<collectingDate id="58702B145F442FC570609DD88C037E4B" value="1996-06-05" valueMax="1996-07-07" valueMin="1996-06-05">5 Jun-7 Jul1996</collectingDate>
;
<collectorName id="F409DD6C523A953D490896CBBFAC951C">P. Hibbs</collectorName>
leg.; KNHM ECU1H96 014
</materialsCitation>
;
<materialsCitation id="4081F47FD91EEBDC82D05A4872FC6480" collectingDate="1996-04-28" collectingDateMax="1996-05-10" collectingDateMin="1996-04-28" collectorName="P. Hibbs" country="Ecuador" latitude="-0.33333334" location="Bilsa" longLatPrecision="21" longitude="-79.71667" specimenCount="1" specimenCount-female="1" stateProvince="Esmeraldas" typeStatus="paratype">
<specimenCount id="C3B7E472B502286A29E62562A19A9631" count="1" type="female">1 female</specimenCount>
; same location but;
<collectingDate id="6043FF4F39D550B192C4FB812219001B" value="1996-04-28" valueMax="1996-05-10" valueMin="1996-04-28">28 Apr-10 May 1996</collectingDate>
;
<collectorName id="BB8E5DFD8F140FB32C987A5E71609AA9">P. Hibbs</collectorName>
leg.; KNHM ECU1H 015
</materialsCitation>
.
</paragraph>
<caption id="2C0D9305D337A7C82CE3FEF5B39A40BC" doi="10.3897/contrib.entomol.74.e114543.figures5-8" httpUri="https://binary.pensoft.net/fig/975716" pageId="0" pageNumber="13" start="Figures 5–8" startId="F2">
<paragraph id="588AD48D8BF41C27DBE302E8688885E7" pageId="0" pageNumber="13">
<emphasis id="E9CA7CCE2BD49DF21683C543A3481727" bold="true" pageId="0" pageNumber="13">Figures 5-8.</emphasis>
<emphasis id="5D2E4EDF8E01C6ABF79C74C6BC699D64" bold="true" pageId="0" pageNumber="13">5.</emphasis>
<taxonomicName id="7DA3394EF82982BF7F513BAB4D7598ED" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Lithocharodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithocharodes hansoni" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="hansoni">
<emphasis id="C13DF189F47A4E699D11CC4E648D830F" italics="true" pageId="0" pageNumber="13">Lithocharodes hansoni</emphasis>
</taxonomicName>
;
<emphasis id="F49DB8FD78C138983D4453AFCCDD1A13" bold="true" pageId="0" pageNumber="13">6.</emphasis>
<taxonomicName id="F0162868EAA1E66503D90D839C578D66" lsidName="L. hibbsi" pageId="0" pageNumber="13" rank="species" species="hibbsi">
<emphasis id="3C6E59E06C4FFECD9460F91FD20A61C0" italics="true" pageId="0" pageNumber="13">L. hibbsi</emphasis>
</taxonomicName>
;
<emphasis id="03098A70511C2918EDA0FAF4D0FE3E7D" bold="true" pageId="0" pageNumber="13">7.</emphasis>
<taxonomicName id="AC487AE64F33A8A784F168350A09C3E9" lsidName="L. lituratus" pageId="0" pageNumber="13" rank="species" species="lituratus">
<emphasis id="23AAF8C58E88380BCF7DA22B76C17369" italics="true" pageId="0" pageNumber="13">L. lituratus</emphasis>
</taxonomicName>
;
<emphasis id="00D10D8AFF02B2D5BFBECCA6F2D6DAE1" bold="true" pageId="0" pageNumber="13">8.</emphasis>
<taxonomicName id="44A010701FD4E3E07B7DF7A37F650239" lsidName="L. montanus" pageId="0" pageNumber="13" rank="species" species="montanus">
<emphasis id="08D9DFD4465C8A3AAE13DFA16DF76BBB" italics="true" pageId="0" pageNumber="13">L. montanus</emphasis>
</taxonomicName>
.
<emphasis id="6CAEBD946B7DB4A0397A0DC2651BBD9C" bold="true" pageId="0" pageNumber="13">a.</emphasis>
Aedeagus;
<emphasis id="2927A3CEA2614F9000829239C86B9BFA" bold="true" pageId="0" pageNumber="13">b.</emphasis>
Paramere;
<emphasis id="F83D288F9CEBBC4F6C105CAC568715D7" bold="true" pageId="0" pageNumber="13">c.</emphasis>
Sternite VII;
<emphasis id="EE0B4642CF3AEED63C59E40E829C76BC" bold="true" pageId="0" pageNumber="13">d.</emphasis>
Tergite VII. Scale bars:
<quantity id="F0F3774B72A234F773678F19B58A05D2" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
(
<emphasis id="7D1BB9F6A48113E633B57830AFCCB60D" bold="true" pageId="0" pageNumber="13">a, c, d</emphasis>
);
<quantity id="DBD4F61C6E5C8AB885D2FD9D93DCC93A" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1 mm</quantity>
(
<emphasis id="9AFD8725597BF763125D9F34E5C8B780" bold="true" pageId="0" pageNumber="13">b</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9FAD27EC04297D75D616F514E7F4B10C" pageId="0" pageNumber="13" type="diagnosis">
<paragraph id="CAA2AA55A665B199D2B5FD7693BE021C" pageId="0" pageNumber="13">Diagnosis.</paragraph>
<paragraph id="2E6DB47489246A0D329A654B8F27B702" pageId="0" pageNumber="13">
The structure of the aedeagus characterises
<taxonomicName id="3EF3BBA9D84C725B665EE0CC9DB87A80" lsidName="L. montanus" pageId="0" pageNumber="13" rank="species" species="montanus">
<emphasis id="22460A93466921497083A6F2BC9F8B04" italics="true" pageId="0" pageNumber="13">L. montanus</emphasis>
</taxonomicName>
as a member of the
<taxonomicName id="1BC19E9F3BF9659B068B418D73A2882B" lsidName="L. simillima" pageId="0" pageNumber="13" rank="species" species="simillima">
<emphasis id="CE9DD30B0F81A1BE994CC7C60B7C0816" italics="true" pageId="0" pageNumber="13">L. simillima</emphasis>
</taxonomicName>
-group by the elongate endophallus with lobes. It differs from the other species of the group by the elongate head with short eyes. Eyes are less than one fifth as wide as postocular space. The other species of the group have mostly a triangular head with distinctly larger eyes.
<taxonomicName id="D7C9CE768CB00050E1DBD40AD323BD93" authorityName="Irmler" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Lithocharodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithocharodes somoleptoides" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="somoleptoides">
<emphasis id="B195CFAC537901B92A9D66FB60C9362A" italics="true" pageId="0" pageNumber="13">Lithocharodes somoleptoides</emphasis>
</taxonomicName>
Irmler, 2021 has a similar shape of the head, but the endophallus has distinct torsions, which are absent in
<taxonomicName id="8280673B3F61E0379108AB2866FDDA94" lsidName="L. montanus" pageId="0" pageNumber="13" rank="species" species="montanus">
<emphasis id="C23AE41E0DB9E1973B87E6C6323E9A1A" italics="true" pageId="0" pageNumber="13">L. montanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A992C7EA8BAC18FE2EE5E14295522ABA" pageId="0" pageNumber="13" type="description">
<paragraph id="B3DF53FC6DFA0ADFDF2B103AAFF8B727" pageId="0" pageNumber="13">Description.</paragraph>
<paragraph id="F632E3598A0DB7D7E4D1AE893FE1C7A2" pageId="0" pageNumber="13">Length: 6.0 mm; Colouration: completely blackish; legs and antennae yellow brown.</paragraph>
<paragraph id="504BD1BCE4CDB43DA21CE86168D8298B" pageId="0" pageNumber="13">Head: 1.02 mm long, 0.78 mm wide; eyes short; PS:E ratio nearly 5.2; postocular sides parallel; without postocular angles; posterior part rounded; nearly semi-circular; inter-antennal furrows indistinct; indistinct depression between eyes; setiferous punctation moderately deep and dense; on average, interstices between punctures 1-1.5 times as wide as diameter of punctures; surface without microsculpture; polished; antennae with first antennomere elongate; half as long as head; second and third antennomere triangular; longer than wide; combined longer than half-length of first antennomere; following antennomeres transverse; on average, twice as wide as long; width increasing to apex. Pronotum: 1.21 mm long, 0.72 mm wide; sides parallel; anterior and posterior part nearly semi-circular; without distinct angles; setiferous punctation deep and dense; on average, interstices between punctures less wide than diameter of punctures; wide mid-line impunctate. Elytra: 1.22 mm long, 0.98 mm wide; humeral angles obtuse; posteriorly, sides slightly divergent; posterior margin retreated to suture; setiferous punctation deep and dense, but slightly finer and sparser than on pronotum; on average, interstices between punctures as wide as to slightly less wide as diameter of punctures; surface without microsculpture; polished. Abdomen finely and sparsely punctate; surface without microsculpture; polished; sternite VII of male with posterior margin sinuate with distinct prominent centre; posterior margin of tergite VII of male less sinuate as sternite; meso-tibia with 4, meta-tibia with 3 ctenidia. Aedeagus oval with sub-rectangular anterior margin; dorsal plate nearly total covering central lobe; endophallus elongate with one basal and one apical torsion; covered by short lobes in central part, teeth-like structures in basal part and smaller lobes in apical part; paramere lunulate; on inner edge with long setae at base and shorter setae near apex.</paragraph>
</subSubSection>
<subSubSection id="F8229AD5731513D17F06686D18AF5258" pageId="0" pageNumber="13" type="etymology">
<paragraph id="3022546196003DD1A79A083ADA932C30" pageId="0" pageNumber="13">Etymology.</paragraph>
<paragraph id="983F9ACA12BD71F0BB58DF5BBD372DB5" pageId="0" pageNumber="13">
The species name is derived from the Latin word
<emphasis id="50CD281429C16A94D2BA4CC41AE84609" italics="true" pageId="0" pageNumber="13">mons</emphasis>
and refers to the high montane area, where it was collected in Ecuador.
</paragraph>
</subSubSection>
<subSubSection id="446DAE38595DEC100247CB1F3EA57D50" pageId="0" pageNumber="13" type="geography">
<paragraph id="9491D74408179BBA4CD1FE0555B52947" pageId="0" pageNumber="13">Geography.</paragraph>
<paragraph id="52487EE3CB512D0AC8BE6B5DD9743CAA" pageId="0" pageNumber="13">Western Ecuador.</paragraph>
</subSubSection>
<subSubSection id="6372521A3F0CCE319686C163A1398780" pageId="0" pageNumber="13" type="ecology">
<paragraph id="B27A035C6BE8C0CA28BC5F051897B58D" pageId="0" pageNumber="13">Ecology.</paragraph>
<paragraph id="8D3A2A4F889E3695484E1190AA607781" pageId="0" pageNumber="13">Floor of montane and cloud forests or similar agricultural systems between 1300 and 1600 m elevation from October to May.</paragraph>
</subSubSection>
</treatment>
</document>
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