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<document id="BC4874B2B03C5CBBA84B61FCD171CC02" ID-CLB-Dataset="29028" ID-DOI="10.11646/zootaxa.4459.2.1" ID-GBIF-Dataset="9b92322c-8063-439b-87f1-4c2b2d55742f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458488" ID-ZooBank="B545851E-6B10-4D3F-B17B-2643BED06EFA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539288412448" checkinUser="plazi" docAuthor="Prado, André Wanderley Do, Baptista, Renner Luiz Cerqueira &amp; Machado, Miguel" docDate="2018" docId="5C486A3CFF88FFB2CFC2FDAC7D64F8FB" docLanguage="en" docName="zootaxa.4459.2.1.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epicadinus biocellatus Mello-Leitao 1929" docType="treatment" docVersion="7" lastPageNumber="209" masterDocId="A0711244FF8EFFBACF55FF937809FF88" masterDocTitle="Taxonomic review of Epicadinus Simon, 1895 (Araneae: Thomisidae)" masterLastPageNumber="234" masterPageNumber="201" pageNumber="207" updateTime="1698578120459" updateUser="ExternalLinkService">
<mods:mods id="A258601BD162E8640E7AA51F33DE3B04" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="DFD39C53D4FD9C0AF0FE33D4EFD537D6">Taxonomic review of Epicadinus Simon, 1895 (Araneae: Thomisidae)</mods:title>
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<mods:name id="9D8896001233A253FFBF0B85B08C54EC" type="personal">
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<mods:namePart id="9AD722C183F88BB87B9523516487359E">Prado, André Wanderley Do</mods:namePart>
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<mods:namePart id="0A0F66F1D1892F8C126CA6DE25AA7460">Baptista, Renner Luiz Cerqueira</mods:namePart>
</mods:name>
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<mods:namePart id="C180CBE6E68DDE94862BCF9EA8A03598">Machado, Miguel</mods:namePart>
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<mods:typeOfResource id="88EBD49303070B0AE7C2DB50E2D3B729">text</mods:typeOfResource>
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<mods:title id="3BC60560F78CEBA4926D3C199EBC1539">Zootaxa</mods:title>
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<mods:part id="0A8BD4F6E170469E86F9C3EF314AD710">
<mods:date id="D9932BCB6B0B60506700C2B145949968">2018</mods:date>
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<mods:number id="B87F06FBA2310E49F8C59779705FCFC2">2018-08-15</mods:number>
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<mods:number id="B067DD93D0E9E7FF7A10854B9687A696">4459</mods:number>
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<mods:classification id="B2531A33DCA806D33524015F509CA59A">journal article</mods:classification>
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<mods:identifier id="C55C08415B26981390B523C1F6298E6E" type="DOI">10.11646/zootaxa.4459.2.1</mods:identifier>
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<treatment id="5C486A3CFF88FFB2CFC2FDAC7D64F8FB" ID-DOI="http://doi.org/10.5281/zenodo.6490579" ID-GBIF-Taxon="148692664" ID-Zenodo-Dep="6490579" LSID="urn:lsid:plazi:treatment:5C486A3CFF88FFB2CFC2FDAC7D64F8FB" httpUri="http://treatment.plazi.org/id/5C486A3CFF88FFB2CFC2FDAC7D64F8FB" lastPageId="8" lastPageNumber="209" pageId="6" pageNumber="207">
<subSubSection id="9CFB88A1FF88FFBCCFC2FDAC792BFDF3" pageId="6" pageNumber="207" type="nomenclature">
<paragraph id="D45EDB2AFF88FFBCCFC2FDAC7A92FDD2" blockId="6.[151,667,575,635]" box="[151,667,575,602]" pageId="6" pageNumber="207">
<heading id="8F166C46FF88FFBCCFC2FDAC7A92FDD2" bold="true" box="[151,667,575,602]" fontSize="11" level="1" pageId="6" pageNumber="207" reason="1">
<taxonomicName id="13E1A0A9FF88FFBCCFC2FDAC7A92FDD2" ID-CoL="6FRCC" authority="Mello-Leitao, 1929" authorityName="Mello-Leitao" authorityYear="1929" box="[151,667,575,602]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="biocellatus">
<emphasis id="E6950738FF88FFBCCFC2FDAC7A92FDD2" bold="true" box="[151,667,575,602]" pageId="6" pageNumber="207">
<emphasis id="E6950738FF88FFBCCFC2FDAC79A0FDD1" bold="true" box="[151,425,575,601]" italics="true" pageId="6" pageNumber="207">Epicadinus biocellatus</emphasis>
<bibRefCitation id="B070A6DBFF88FFBCCEE5FDD37A92FDD2" box="[432,667,576,602]" pageId="6" pageNumber="207" refId="ref24568" refString="Mello-Leitao, C. F. (1929) Aphantochilidas e Thomisidas do Brasil. Arquivos do Museu Nacional do Rio de Janeiro, 31, 9 - 359." type="journal article">Mello-Leitão, 1929</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D45EDB2AFF88FFBCCFC2FDF1792BFDF3" blockId="6.[151,667,575,635]" box="[151,290,610,635]" pageId="6" pageNumber="207">
<figureCitation id="4CDAC7AFFF88FFBCCFC2FDF178F2FDF3" box="[151,251,610,635]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="9.[151,250,1908,1930]" captionStartId-1="10.[151,250,1899,1921]" captionStartId-2="11.[151,250,942,964]" captionTargetBox-0="[194,1395,195,1892]" captionTargetBox-1="[197,1402,193,1885]" captionTargetBox-2="[151,1436,193,933]" captionTargetId-0="figure@9.[194,1395,193,1892]" captionTargetId-1="figure@10.[197,1402,193,1885]" captionTargetId-2="figure@11.[151,1436,193,933]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURE 2. Epicadinus biocellatus Mello-Leitão, 1929. A–G female (MPEG 31330); H female (MPEG 34862). A–C habitus (A dorsal, B lateral, C ventral); D–E, epigynum (D photo ventral, E SEM ventral); F–G vulva (F photo ventral, G dorsal); H live specimen in hunting position. Abbreviations, A, atrium; AC, anterior curve of copulatory duct; CO, copulatory opening; FD, fertilization duct; MS, median septum. Scale bars: A–C, 1 mm; D–G, 0.1 mm. Photo credits: H, César Favacho." captionText-1="FIGURE 3. Epicadinus biocellatus Mello-Leitão, 1929, male (MPEG 31328). A–C habitus (A dorsal, B lateral, C ventral); D– H left male palpus (D ventral, E–F retrolateral, G slanted retrolateral, H cymbium apex, ventral). Abbreviations, BP, basal pouch; DTA, distal tibial apophysis; E, embolus; EA, embolus apex; MF, marginal furrow of cymbium; RTA, retrolateral tibial apophysis; TF, tegular furrow; TM, tibial macrosetae. Scale bars: A–C, 1 mm; D–H, 0.1 mm." captionText-2="FIGURE 4. Epicadinus biocellatus female holotype (MHNH AR-15077, Coll. Simon 2044b) [total length: 4 mm]. A dorsal habitus; B venter of opisthosoma; C–E original labels of the splitted typical lot (C female holotype 2044b, D misidentified male of E. biocellatus 2044, E female of E. cornutus 2044a). Photo credits: Christine Rollard and MNHN Laboratory of Arachnology crew." httpUri-0="https://zenodo.org/record/1458492/files/figure.png" httpUri-1="https://zenodo.org/record/1458494/files/figure.png" httpUri-2="https://zenodo.org/record/1458496/files/figure.png" pageId="6" pageNumber="207">Figs 2–4</figureCitation>
,
<figureCitation id="4CDAC7AFFF88FFBCCE53FDF1792BFDF3" box="[262,290,610,635]" captionStart="FIGURE 15" captionStartId="31.[151,250,1300,1322]" captionTargetBox="[166,1420,208,1263]" captionTargetId="figure@31.[150,1436,193,1278]" captionTargetPageId="31" captionText="FIGURE 15. Geographic distribution records of Epicadinus, highlighting the type-locality of each of the four species (star symbols)." httpUri="https://zenodo.org/record/1458518/files/figure.png" pageId="6" pageNumber="207">15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="9CFB88A1FF88FFBCCFC2FD397B6DFD57" pageId="6" pageNumber="207" type="reference_group">
<paragraph id="D45EDB2AFF88FFBCCFC2FD397A8DFD48" blockId="6.[151,868,682,735]" box="[151,644,682,704]" pageId="6" pageNumber="207">
<treatmentCitationGroup id="F4F1FC04FF88FFBCCFC2FD397A8DFD48" box="[151,644,682,704]" pageId="6" pageNumber="207">
<taxonomicName id="13E1A0A9FF88FFBCCFC2FD397A89FD48" ID-CoL="6FRCC" authority="Mello-Leitao, 1929: 104" authorityName="Mello-Leitao" authorityPageNumber="104" authorityYear="1929" box="[151,640,682,704]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="biocellatus">
<emphasis id="E6950738FF88FFBCCFC2FD397977FD48" box="[151,382,682,704]" italics="true" pageId="6" pageNumber="207">Epicadinus biocellatus</emphasis>
<treatmentCitation id="5540FD3BFF88FFBCCED1FD397A89FD48" author="" box="[388,640,682,704]" page="104" pageId="6" pageNumber="207">
<bibRefCitation id="B070A6DBFF88FFBCCED1FD397A44FD48" box="[388,589,682,704]" pageId="6" pageNumber="207" refId="ref24568" refString="Mello-Leitao, C. F. (1929) Aphantochilidas e Thomisidas do Brasil. Arquivos do Museu Nacional do Rio de Janeiro, 31, 9 - 359." type="journal article">Mello-Leitão, 1929</bibRefCitation>
: 104
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D45EDB2AFF88FFBCCFC2FD5A7B6DFD57" blockId="6.[151,868,682,735]" box="[151,868,713,735]" pageId="6" pageNumber="207">
<treatmentCitationGroup id="F4F1FC04FF88FFBCCFC2FD5A7B6DFD57" box="[151,868,713,735]" pageId="6" pageNumber="207">
<taxonomicName id="13E1A0A9FF88FFBCCFC2FD5A7A15FD57" ID-CoL="6G3CG" authority="(Taczanowski)" authorityName="Taczanowski" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[151,540,713,735]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="trispinosus">
<emphasis id="E6950738FF88FFBCCFC2FD5A7977FD57" box="[151,382,713,735]" italics="true" pageId="6" pageNumber="207">Epicadinus trispinosus</emphasis>
(Taczanowski)
</taxonomicName>
.
<treatmentCitation id="5540FD3BFF88FFBCCD73FD5A7B02FD57" author="" box="[550,779,713,735]" page="99" pageId="6" pageNumber="207">
<bibRefCitation id="B070A6DBFF88FFBCCD73FD5A7AE1FD57" box="[550,744,713,735]" pageId="6" pageNumber="207" refId="ref24568" refString="Mello-Leitao, C. F. (1929) Aphantochilidas e Thomisidas do Brasil. Arquivos do Museu Nacional do Rio de Janeiro, 31, 9 - 359." type="journal article">Mello-Leitão 1929</bibRefCitation>
: 99
</treatmentCitation>
(in part)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9CFB88A1FF88FFBCCFC2FC9F7BACFA58" pageId="6" pageNumber="207" type="materials_examined">
<paragraph id="D45EDB2AFF88FFBCCFC2FC9F7A42FCC0" blockId="6.[151,1437,780,2029]" pageId="6" pageNumber="207">
<emphasis id="E6950738FF88FFBCCFC2FC9F79C3FCAD" bold="true" box="[151,458,780,805]" pageId="6" pageNumber="207">
<typeStatus id="0B5A6588FF88FFBCCFC2FC9F78DBFCAD" box="[151,210,780,805]" pageId="6" pageNumber="207">Type</typeStatus>
material:
<typeStatus id="0B5A6588FF88FFBCCE01FC9F79CCFCAD" box="[340,453,780,805]" pageId="6" pageNumber="207" type="holotype">Holotype</typeStatus>
:
</emphasis>
female,
<collectingCountry id="ACF69BBAFF88FFBCCD67FC9F7A9FFCAC" box="[562,662,780,804]" name="Brazil" pageId="6" pageNumber="207">BRASIL</collectingCountry>
or
<collectionCode id="B2F043EFFF88FFBCCD94FC9E7B05FCAC" box="[705,780,781,804]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14398" name="Universit<69> di Perugia" pageId="6" pageNumber="207">
<collectingCountry id="ACF69BBAFF88FFBCCD94FC9E7B05FCAC" box="[705,780,781,804]" name="Peru" pageId="6" pageNumber="207">PERU</collectingCountry>
</collectionCode>
, “
<collectingRegion id="162515C8FF88FFBCCC71FC9F7B51FCAC" box="[804,856,780,804]" country="Brazil" name="Para" pageId="6" pageNumber="207">Para</collectingRegion>
ou Pebas
<specimenCount id="C2E710A3FF88FFBCCC81FC9E7BE0FCAC" box="[980,1001,781,804]" pageId="6" pageNumber="207" type="female">♀</specimenCount>
<typeStatus id="0B5A6588FF88FFBCCCA6FC9F7C2EFCAC" box="[1011,1063,780,804]" pageId="6" pageNumber="207">type</typeStatus>
”, [M. de Mathan] (
<collectionCode id="B2F043EFFF88FFBCCA5FFC9F7D6BFCAC" box="[1290,1378,780,804]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="207">MNHN</collectionCode>
<collectionCode id="B2F043EFFF88FFBCCA3EFC9F7D92FCAC" box="[1387,1435,780,804]" pageId="6" pageNumber="207">AR-</collectionCode>
15077, Coll. Simon 2044b), examined.
</paragraph>
<paragraph id="D45EDB2AFF88FFBCCF92FCC77BACFA58" blockId="6.[151,1437,780,2029]" pageId="6" pageNumber="207">
<materialsCitation id="6489D177FF88FFBCCF92FCC77C6AFC18" ID-GBIF-Occurrence="1917355309" collectingDate="2011-10-21" collectionCode="MPEG" collectorName="M. A. Aguiar-Neto" country="Brazil" latitude="-3.9261973" location="Reserva Biologica do Gurupi" longLatPrecision="1" longitude="-46.772003" municipality="Bom Jardim" pageId="6" pageNumber="207" specimenCode="MPEG 31329" specimenCount="1" specimenCount-female="1" stateProvince="Maranhao">
<emphasis id="E6950738FF88FFBCCF92FCC77AAEFCE5" bold="true" box="[199,679,852,877]" pageId="6" pageNumber="207">
Additional material examined.
<collectingCountry id="ACF69BBAFF88FFBCCD69FCC77AAEFCE5" box="[572,679,852,877]" name="Brazil" pageId="6" pageNumber="207">BRAZIL</collectingCountry>
</emphasis>
:
<emphasis id="E6950738FF88FFBCCDE2FCC77B32FCE5" bold="true" box="[695,827,852,877]" italics="true" pageId="6" pageNumber="207">
<collectingRegion id="162515C8FF88FFBCCDE2FCC77B3DFCE5" box="[695,820,852,877]" country="Brazil" name="Maranhao" pageId="6" pageNumber="207">Maranhão</collectingRegion>
:
</emphasis>
<specimenCount id="C2E710A3FF88FFBCCC16FCC77BAEFCE5" box="[835,935,852,877]" pageId="6" pageNumber="207" type="female">1 female</specimenCount>
,
<collectingMunicipality id="343A4150FF88FFBCCCE6FCC67C49FCE5" box="[947,1088,852,877]" pageId="6" pageNumber="207">Bom Jardim</collectingMunicipality>
,
<location id="D13E8DF1FF88FFBCCB19FCC67D91FCE5" LSID="urn:lsid:plazi:treatment:5C486A3CFF88FFB2CFC2FDAC7D64F8FB:D13E8DF1FF88FFBCCB19FCC67D91FCE5" box="[1100,1432,852,877]" country="Brazil" latitude="-3.9261973" longLatPrecision="1" longitude="-46.772003" municipality="Bom Jardim" name="Reserva Biologica do Gurupi" pageId="6" pageNumber="207" stateProvince="Maranhao">Reserva Biológica do Gurupi</location>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCFC2FCE47925FC18" box="[151,300,887,912]" direction="south" orientation="latitude" pageId="6" pageNumber="207" precision="1" value="-3.9261973">3°55'34.31''S</geoCoordinate>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCE6EFCE479EEFC18" box="[315,487,887,912]" direction="west" orientation="longitude" pageId="6" pageNumber="207" precision="1" value="-46.772003">46°46'19.21''W</geoCoordinate>
,
<collectingDate id="B01B0402FF88FFBCCEA0FCEB7AB0FC18" box="[501,697,887,912]" pageId="6" pageNumber="207" value="2011-10-21">
<date id="A05FFDEAFF88FFBCCEA0FCEB7AB0FC18" box="[501,697,887,912]" pageId="6" pageNumber="207" value="2011-10-21">21 October 2011</date>
</collectingDate>
,
<collectorName id="7914BEFCFF88FFBCCD9DFCEB7BACFC18" box="[712,933,887,912]" pageId="6" pageNumber="207">M. A. Aguiar-Neto</collectorName>
(
<specimenCode id="84477351FF88FFBCCCE3FCEB7C52FC18" box="[950,1115,888,912]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207">MPEG 31329</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="6489D177FF88FFBCCB27FCEB7999FC50" ID-GBIF-Occurrence="1917355278" collectingDate="2012-03-15" collectionCode="MPEG" collectorName="M. A. Aguiar-Neto" country="Brazil" latitude="-3.6816916" location="Reserva Biologica do Gurupi" longLatPrecision="1" longitude="-46.754234" municipality="Centro" pageId="6" pageNumber="207" specimenCode="MPEG 31322" specimenCount="1" specimenCount-female="1" stateProvince="Maranhao">
<specimenCount id="C2E710A3FF88FFBCCB27FCEB7CD0FC18" box="[1138,1241,887,912]" pageId="6" pageNumber="207" type="female">1 female</specimenCount>
,
<collectingMunicipality id="343A4150FF88FFBCCBBDFCEB7D3DFC18" box="[1256,1332,888,912]" pageId="6" pageNumber="207">Centro</collectingMunicipality>
novo do
<collectingRegion id="162515C8FF88FFBCCFC2FC0F7905FC3D" box="[151,268,924,949]" country="Brazil" name="Maranhao" pageId="6" pageNumber="207">Maranhão</collectingRegion>
,
<location id="D13E8DF1FF88FFBCCE48FC0E7A70FC3D" LSID="urn:lsid:plazi:treatment:5C486A3CFF88FFB2CFC2FDAC7D64F8FB:D13E8DF1FF88FFBCCE48FC0E7A70FC3D" box="[285,633,924,949]" country="Brazil" latitude="-3.6816916" longLatPrecision="1" longitude="-46.754234" municipality="Centro" name="Reserva Biologica do Gurupi" pageId="6" pageNumber="207" stateProvince="Maranhao">Reserva Biológica do Gurupi</location>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCDDFFC0F7B16FC3D" box="[650,799,924,949]" direction="south" orientation="latitude" pageId="6" pageNumber="207" precision="1" value="-3.6816916">3°40'54.09''S</geoCoordinate>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCC64FC0F7BD7FC3D" box="[817,990,924,949]" direction="west" orientation="longitude" pageId="6" pageNumber="207" precision="1" value="-46.754234">46°45'15.24''W</geoCoordinate>
,
<collectingDate id="B01B0402FF88FFBCCCBAFC0F7CA0FC3C" box="[1007,1193,924,949]" pageId="6" pageNumber="207" value="2012-03-15">
<date id="A05FFDEAFF88FFBCCCBAFC0F7CA0FC3C" box="[1007,1193,924,949]" pageId="6" pageNumber="207" value="2012-03-15">15 March 2012</date>
</collectingDate>
,
<collectorName id="7914BEFCFF88FFBCCBEFFC0E7D95FC3D" box="[1210,1436,924,949]" pageId="6" pageNumber="207">M. A. Aguiar-Neto</collectorName>
(
<specimenCode id="84477351FF88FFBCCFCBFC53794AFC50" box="[158,323,960,984]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207">MPEG 31322</specimenCode>
);
<emphasis id="E6950738FF88FFBCCE0FFC2C7999FC50" box="[346,400,959,984]" italics="true" pageId="6" pageNumber="207">idem</emphasis>
</materialsCitation>
, 1 female, 3°41'7.92''S, 46°45'46.08''W,
<date id="A05FFDEAFF88FFBCCC3DFC537C31FC50" box="[872,1080,959,984]" pageId="6" pageNumber="207" value="2011-11-09">9 November 2011</date>
(MPEG 31327);
<emphasis id="E6950738FF88FFBCCA50FC2C7D32FC50" box="[1285,1339,959,984]" italics="true" pageId="6" pageNumber="207">idem</emphasis>
, 1 male (MPEG 31328);
<materialsCitation id="6489D177FF88FFBCCE37FC777BD9FBA8" ID-GBIF-Occurrence="1917355249" collectingDate="2014-05-08" collectionCode="MPEG" collectorName="M. A. Aguiar-Neto" country="Brazil" county="Centro" latitude="-1.8633556" location="Fazenda Sete Irmaos" longLatPrecision="1" longitude="-45.760838" municipality="Candido Mendes" pageId="6" pageNumber="207" specimenCode="MPEG 31324" specimenCount="1" specimenCount-male="1" stateProvince="Maranhao">
<specimenCount id="C2E710A3FF88FFBCCE37FC7779B1FC75" box="[354,440,996,1021]" pageId="6" pageNumber="207" type="male">1 male</specimenCount>
,
<collectingCounty id="3D3FA3A6FF88FFBCCE9EFC777A1EFC74" box="[459,535,996,1020]" pageId="6" pageNumber="207">Centro</collectingCounty>
novo do
<collectingRegion id="162515C8FF88FFBCCDCDFC777B05FC75" box="[664,780,996,1021]" country="Brazil" name="Maranhao" pageId="6" pageNumber="207">Maranhão</collectingRegion>
,
<collectingMunicipality id="343A4150FF88FFBCCC4AFC777BE1FC75" box="[799,1000,996,1021]" pageId="6" pageNumber="207">Cândido Mendes</collectingMunicipality>
,
<location id="D13E8DF1FF88FFBCCCAEFC777CFDFC74" LSID="urn:lsid:plazi:treatment:5C486A3CFF88FFB2CFC2FDAC7D64F8FB:D13E8DF1FF88FFBCCCAEFC777CFDFC74" box="[1019,1268,996,1021]" country="Brazil" county="Centro" latitude="-1.8633556" longLatPrecision="1" longitude="-45.760838" municipality="Candido Mendes" name="Fazenda Sete Irmaos" pageId="6" pageNumber="207" stateProvince="Maranhao">Fazenda Sete Irmãos</location>
<geoCoordinate id="B1D5BDEDFF88FFBCCA57FC777D9EFC75" box="[1282,1431,996,1021]" direction="south" orientation="latitude" pageId="6" pageNumber="207" precision="1" value="-1.8633556">1°51'48.08''S</geoCoordinate>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCFC2FB94794BFBA8" box="[151,322,1031,1056]" direction="west" orientation="longitude" pageId="6" pageNumber="207" precision="1" value="-45.760838">45°45'39.02''W</geoCoordinate>
, 0
<collectingDate id="B01B0402FF88FFBCCE0AFB9B79E5FBA8" box="[351,492,1032,1056]" pageId="6" pageNumber="207" value="2014-05-08">
<date id="A05FFDEAFF88FFBCCE0AFB9B79E5FBA8" box="[351,492,1032,1056]" pageId="6" pageNumber="207" value="2014-05-08">8 May 2014</date>
</collectingDate>
,
<collectorName id="7914BEFCFF88FFBCCEACFB9B7ADBFBA8" box="[505,722,1031,1056]" pageId="6" pageNumber="207">M. A. Aguiar-Neto</collectorName>
(
<specimenCode id="84477351FF88FFBCCDB7FB9B7B8CFBA8" box="[738,901,1032,1056]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207">MPEG 31324</specimenCode>
),
<emphasis id="E6950738FF88FFBCCCCFFB947BD9FBA8" box="[922,976,1031,1056]" italics="true" pageId="6" pageNumber="207">idem</emphasis>
</materialsCitation>
,
<materialsCitation id="6489D177FF88FFBCCCB5FB9B7943FBE0" ID-GBIF-Occurrence="1917355299" collectingDate="2001-11-19" collectingDateMax="2014-05-08" collectingDateMin="2001-11-19" collectionCode="MPEG" collectorName="L.M. Almeida-Silva &amp; Domingos" country="Brazil" latitude="-1.579911" location="Barcarena" longLatPrecision="1" longitude="-48.598866" municipality="Neto" pageId="6" pageNumber="207" specimenCode="MPEG 31330, MPEG 31332" specimenCount="3" specimenCount-female="3" stateProvince="Para">
<specimenCount id="C2E710A3FF88FFBCCCB5FB9B7C58FBA8" box="[992,1105,1031,1056]" pageId="6" pageNumber="207" type="female">2 females</specimenCount>
,
<collectingDate id="B01B0402FF88FFBCCB0BFB9B7CE5FBA8" box="[1118,1260,1032,1056]" pageId="6" pageNumber="207" value="2014-05-08">
<date id="A05FFDEAFF88FFBCCB0BFB9B7CE5FBA8" box="[1118,1260,1032,1056]" pageId="6" pageNumber="207" value="2014-05-08">8 May 2014</date>
</collectingDate>
,
<collectorName id="7914BEFCFF88FFBCCBACFB9B7D34FBA8" box="[1273,1341,1032,1056]" name="L.M. Almeida-Silva" pageId="6" pageNumber="207">M. A.</collectorName>
Aguiar-
<collectingMunicipality id="343A4150FF88FFBCCFC2FBBE78C4FBCC" box="[151,205,1069,1092]" pageId="6" pageNumber="207">Neto</collectingMunicipality>
(
<specimenCode id="84477351FF88FFBCCF8EFBBF7975FBCC" box="[219,380,1068,1092]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207">MPEG 31330</specimenCode>
).
<emphasis id="E6950738FF88FFBCCEDBFBBF79C6FBCD" bold="true" box="[398,463,1068,1093]" italics="true" pageId="6" pageNumber="207">
<collectingRegion id="162515C8FF88FFBCCEDBFBBF79C1FBCD" box="[398,456,1068,1093]" country="Brazil" name="Para" pageId="6" pageNumber="207">Pará</collectingRegion>
:
</emphasis>
<specimenCount id="C2E710A3FF88FFBCCE83FBBF7A30FBCD" box="[470,569,1068,1093]" pageId="6" pageNumber="207" type="female">1 female</specimenCount>
,
<location id="D13E8DF1FF88FFBCCD11FBBE7AB1FBCC" LSID="urn:lsid:plazi:treatment:5C486A3CFF88FFB2CFC2FDAC7D64F8FB:D13E8DF1FF88FFBCCD11FBBE7AB1FBCC" box="[580,696,1069,1092]" country="Brazil" latitude="-1.579911" longLatPrecision="1" longitude="-48.598866" municipality="Neto" name="Barcarena" pageId="6" pageNumber="207" stateProvince="Para">Barcarena</location>
, ca.
<geoCoordinate id="B1D5BDEDFF88FFBCCDBCFBBF7B76FBCD" box="[745,895,1068,1093]" direction="south" orientation="latitude" pageId="6" pageNumber="207" precision="1" value="-1.579911">1°34'47.68&quot;S</geoCoordinate>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCCDEFBBF7C31FBCD" box="[907,1080,1068,1093]" direction="west" orientation="longitude" pageId="6" pageNumber="207" precision="1" value="-48.598866">48°35'55.92&quot;W</geoCoordinate>
,
<collectingDate id="B01B0402FF88FFBCCB16FBBF7D15FBCC" box="[1091,1308,1068,1093]" pageId="6" pageNumber="207" value="2001-11-19">
<date id="A05FFDEAFF88FFBCCB16FBBF7D15FBCC" box="[1091,1308,1068,1093]" pageId="6" pageNumber="207" value="2001-11-19">19 November 2001</date>
</collectingDate>
,
<collectorName id="7914BEFCFF88FFBCCA72FBBF7D95FBCD" box="[1319,1436,1068,1093]" pageId="6" pageNumber="207">Domingos</collectorName>
(
<specimenCode id="84477351FF88FFBCCFCBFBC3794BFBE0" box="[158,322,1104,1128]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207">MPEG 31332</specimenCode>
)
</materialsCitation>
; 1 male, Belém (MNHN Coll. Simon 2044); 1 male, Belém, Reserva Mocambo, 01°26'28.07''S, 48°24'46.02''W,
<date id="A05FFDEAFF88FFBCCE02FBE77A36FB04" box="[343,575,1140,1165]" pageId="6" pageNumber="207" value="2007-12-11">11 December 2007</date>
, G. Ruiz (MPEG 31333); 1 female, Belém, Bosque Rodrigues Alves, 1°25'49.00''S, 48°27'22.03''W,
<date id="A05FFDEAFF88FFBCCEA4FB0B7A9AFB38" box="[497,659,1175,1200]" pageId="6" pageNumber="207" value="2009-04-22">22 April 2009</date>
, E. G. S. Cafofo (MPEG 31334);
<materialsCitation id="6489D177FF88FFBCCB45FB0B7D8DFB5D" ID-GBIF-Occurrence="1917355297" collectingDate="2018-02-08" collectionCode="MPEG" collectorName="P. Pantoja" country="Brazil" county="Belem" latitude="-1.4512334" location="Campus de Pesquisa" longLatPrecision="1" longitude="-48.444256" municipality="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207" specimenCode="MPEG 34862" specimenCount="2" specimenCount-female="2" stateProvince="Para">
<specimenCount id="C2E710A3FF88FFBCCB45FB0B7C89FB38" box="[1040,1152,1175,1200]" pageId="6" pageNumber="207" type="female">2 females</specimenCount>
,
<collectingCounty id="3D3FA3A6FF88FFBCCBD9FB047CD1FB38" box="[1164,1240,1175,1200]" pageId="6" pageNumber="207">Belém</collectingCounty>
,
<collectingMunicipality id="343A4150FF88FFBCCBB1FB0B7931FB5D" pageId="6" pageNumber="207">Museu Paraense Emílio Goeldi</collectingMunicipality>
,
<location id="D13E8DF1FF88FFBCCE16FB2F7A22FB5D" LSID="urn:lsid:plazi:treatment:5C486A3CFF88FFB2CFC2FDAC7D64F8FB:D13E8DF1FF88FFBCCE16FB2F7A22FB5D" box="[323,555,1212,1237]" country="Brazil" county="Belem" latitude="-1.4512334" longLatPrecision="1" longitude="-48.444256" municipality="Museu Paraense Emilio Goeldi" name="Campus de Pesquisa" pageId="6" pageNumber="207" stateProvince="Para">Campus de Pesquisa</location>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCD60FB2F7AC3FB5C" box="[565,714,1212,1236]" direction="south" orientation="latitude" pageId="6" pageNumber="207" precision="1" value="-1.4512334">1°27'04.44&quot;S</geoCoordinate>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCD81FB2F7B88FB5D" box="[724,897,1212,1237]" direction="west" orientation="longitude" pageId="6" pageNumber="207" precision="1" value="-48.444256">48°26'39.32&quot;W</geoCoordinate>
, 0
<collectingDate id="B01B0402FF88FFBCCCCEFB2F7C5CFB5C" box="[923,1109,1212,1237]" pageId="6" pageNumber="207" value="2018-02-08">
<date id="A05FFDEAFF88FFBCCCCEFB2F7C5CFB5C" box="[923,1109,1212,1237]" pageId="6" pageNumber="207" value="2018-02-08">8 February 2018</date>
</collectingDate>
,
<collectorName id="7914BEFCFF88FFBCCB0AFB2E7CC4FB5D" box="[1119,1229,1212,1237]" pageId="6" pageNumber="207">P. Pantoja</collectorName>
(
<specimenCode id="84477351FF88FFBCCB8EFB2F7D75FB5D" box="[1243,1404,1212,1237]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207">MPEG 34862</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="6489D177FF88FFBCCADBFB2F7D8BFB70" ID-GBIF-Occurrence="1917355312" collectingDate="2014-06-26" collectionCode="MPEG" collectorName="Y. Shimano" country="Brazil" latitude="-2.1075056" location="Vila de Curucambaba" longLatPrecision="1" longitude="-49.309193" municipality="Cameta" pageId="6" pageNumber="207" specimenCode="MPEG 31335" specimenCount="1" specimenCount-female="1" stateProvince="Para">
<specimenCount id="C2E710A3FF88FFBCCADBFB2F78EFFB70" pageId="6" pageNumber="207" type="female">1 female</specimenCount>
,
<collectingMunicipality id="343A4150FF88FFBCCFA4FB737943FB70" box="[241,330,1248,1272]" pageId="6" pageNumber="207">Cametá</collectingMunicipality>
,
<location id="D13E8DF1FF88FFBCCE03FB4C7A45FB70" LSID="urn:lsid:plazi:treatment:5C486A3CFF88FFB2CFC2FDAC7D64F8FB:D13E8DF1FF88FFBCCE03FB4C7A45FB70" box="[342,588,1247,1272]" country="Brazil" latitude="-2.1075056" longLatPrecision="1" longitude="-49.309193" municipality="Cameta" name="Vila de Curucambaba" pageId="6" pageNumber="207" stateProvince="Para">Vila de Curuçambaba</location>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCD02FB4C7AE5FB70" box="[599,748,1247,1272]" direction="south" orientation="latitude" pageId="6" pageNumber="207" precision="1" value="-2.1075056">2°06'27.02''S</geoCoordinate>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCDACFB737B9FFB70" box="[761,918,1248,1272]" direction="west" orientation="longitude" pageId="6" pageNumber="207" precision="1" value="-49.309193">49°18'33.1''W</geoCoordinate>
,
<collectingDate id="B01B0402FF88FFBCCCF7FB4C7C35FB70" box="[930,1084,1247,1272]" pageId="6" pageNumber="207" value="2014-06-26">
<date id="A05FFDEAFF88FFBCCCF7FB4C7C35FB70" box="[930,1084,1247,1272]" pageId="6" pageNumber="207" value="2014-06-26">26 June 2014</date>
</collectingDate>
,
<collectorName id="7914BEFCFF88FFBCCB12FB737CC3FB70" box="[1095,1226,1247,1272]" pageId="6" pageNumber="207">Y. Shimano</collectorName>
(
<specimenCode id="84477351FF88FFBCCB8DFB737D72FB70" box="[1240,1403,1247,1272]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207">MPEG 31335</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="6489D177FF88FFBCCADBFB737C21FAC8" ID-GBIF-Occurrence="1917355341" collectingDate="1994-01-26" collectingDateMax="2007-06-26" collectingDateMin="1994-01-26" collectionCode="FZBRS, MPEG" collectorName="A. D. Brescovit &amp; M. L. N. Sirotheau" country="Brazil" latitude="-2.5102668" location="Alter do Chao" longLatPrecision="1" longitude="-54.952934" municipality="Santarem" pageId="6" pageNumber="207" specimenCode="FZBRS 25051, MPEG 31336" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Para">
<specimenCount id="C2E710A3FF88FFBCCADBFB7378D9FA95" pageId="6" pageNumber="207" type="male">1 male</specimenCount>
,
<collectingMunicipality id="343A4150FF88FFBCCFB5FA977944FA94" box="[224,333,1284,1308]" pageId="6" pageNumber="207">Santarém</collectingMunicipality>
,
<location id="D13E8DF1FF88FFBCCE09FA977A0FFA95" LSID="urn:lsid:plazi:treatment:5C486A3CFF88FFB2CFC2FDAC7D64F8FB:D13E8DF1FF88FFBCCE09FA977A0FFA95" box="[348,518,1284,1309]" country="Brazil" latitude="-2.5102668" longLatPrecision="1" longitude="-54.952934" municipality="Santarem" name="Alter do Chao" pageId="6" pageNumber="207" stateProvince="Para">Alter do Chão</location>
, ca.
<geoCoordinate id="B1D5BDEDFF88FFBCCD14FA977AD1FA95" box="[577,728,1284,1309]" direction="south" orientation="latitude" pageId="6" pageNumber="207" precision="1" value="-2.5102668">2°30'36.96&quot;S</geoCoordinate>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCDBDFA977B9CFA95" box="[744,917,1284,1309]" direction="west" orientation="longitude" pageId="6" pageNumber="207" precision="1" value="-54.952934">54°57'10.56&quot;W</geoCoordinate>
,
<collectingDate id="B01B0402FF88FFBCCCF0FA977C63FA94" box="[933,1130,1284,1309]" pageId="6" pageNumber="207" value="1994-01-26">
<date id="A05FFDEAFF88FFBCCCF0FA977C63FA94" box="[933,1130,1284,1309]" pageId="6" pageNumber="207" value="1994-01-26">26 January 1994</date>
</collectingDate>
,
<collectorName id="7914BEFCFF88FFBCCB2CFA977D3BFA95" box="[1145,1330,1284,1309]" pageId="6" pageNumber="207">A. D. Brescovit</collectorName>
(
<specimenCode id="84477351FF88FFBCCA10FA9778E8FAC8" collectionCode="FZBRS" pageId="6" pageNumber="207">FZBRS 25051</specimenCode>
),
<emphasis id="E6950738FF88FFBCCFA0FAB4793AFAC8" box="[245,307,1319,1344]" italics="true" pageId="6" pageNumber="207">idem,</emphasis>
<specimenCount id="C2E710A3FF88FFBCCE6EFABB79A9FAC8" box="[315,416,1319,1344]" pageId="6" pageNumber="207" type="female">1 female</specimenCount>
,
<collectingDate id="B01B0402FF88FFBCCEF8FAB47A4EFAC8" box="[429,583,1319,1344]" pageId="6" pageNumber="207" value="2007-06-26">
<date id="A05FFDEAFF88FFBCCEF8FAB47A4EFAC8" box="[429,583,1319,1344]" pageId="6" pageNumber="207" value="2007-06-26">26 June 2007</date>
</collectingDate>
,
<collectorName id="7914BEFCFF88FFBCCD01FABA7B22FAC8" box="[596,811,1319,1344]" pageId="6" pageNumber="207">M. L. N. Sirotheau</collectorName>
(
<specimenCode id="84477351FF88FFBCCC6EFABB7BD4FAC8" box="[827,989,1319,1344]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207">MPEG 31336</specimenCode>
),
<emphasis id="E6950738FF88FFBCCCA7FAB47C21FAC8" box="[1010,1064,1319,1344]" italics="true" pageId="6" pageNumber="207">idem</emphasis>
</materialsCitation>
,
<materialsCitation id="6489D177FF88FFBCCB62FABB7CCAFAED" ID-GBIF-Occurrence="1917355292" collectingDate="2007-07-02" collectionCode="MPEG" collectorName="M. L. N. Sirotheau" country="Brazil" location="Silva" pageId="6" pageNumber="207" specimenCode="MPEG 31337, MPEG 31338" specimenCount="2" specimenCount-female="2" stateProvince="Para">
<specimenCount id="C2E710A3FF88FFBCCB62FABB7C95FAC8" box="[1079,1180,1319,1344]" pageId="6" pageNumber="207" type="female">1 female</specimenCount>
, 0
<collectingDate id="B01B0402FF88FFBCCBE2FABB7D36FAC8" box="[1207,1343,1319,1344]" pageId="6" pageNumber="207" value="2007-07-02">
<date id="A05FFDEAFF88FFBCCBE2FABB7D36FAC8" box="[1207,1343,1319,1344]" pageId="6" pageNumber="207" value="2007-07-02">2 July 2007</date>
</collectingDate>
, B. J. F.
<location id="D13E8DF1FF88FFBCCFC2FADF78D8FAED" LSID="urn:lsid:plazi:treatment:5C486A3CFF88FFB2CFC2FDAC7D64F8FB:D13E8DF1FF88FFBCCFC2FADF78D8FAED" box="[151,209,1356,1381]" country="Brazil" name="Silva" pageId="6" pageNumber="207" stateProvince="Para">Silva</location>
(
<specimenCode id="84477351FF88FFBCCFB5FADF798BFAEC" box="[224,386,1356,1380]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207">MPEG 31337</specimenCode>
),
<emphasis id="E6950738FF88FFBCCEC3FADF79DAFAED" box="[406,467,1356,1381]" italics="true" pageId="6" pageNumber="207">idem,</emphasis>
<specimenCount id="C2E710A3FF88FFBCCE89FADF7A48FAED" box="[476,577,1356,1381]" pageId="6" pageNumber="207" type="female">1 female</specimenCount>
, 0
<collectingDate id="B01B0402FF88FFBCCD09FADF7AEDFAEC" box="[604,740,1356,1381]" pageId="6" pageNumber="207" value="2007-07-02">
<date id="A05FFDEAFF88FFBCCD09FADF7AEDFAEC" box="[604,740,1356,1381]" pageId="6" pageNumber="207" value="2007-07-02">2 July 2007</date>
</collectingDate>
,
<collectorName id="7914BEFCFF88FFBCCDA5FADE7BCEFAED" box="[752,967,1356,1381]" pageId="6" pageNumber="207">M. L. N. Sirotheau</collectorName>
(
<specimenCode id="84477351FF88FFBCCC82FADF7C70FAEC" box="[983,1145,1356,1380]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207">MPEG 31338</specimenCode>
),
<emphasis id="E6950738FF88FFBCCBD8FADF7CCAFAED" box="[1165,1219,1356,1381]" italics="true" pageId="6" pageNumber="207">idem</emphasis>
</materialsCitation>
, 1 female,
<date id="A05FFDEAFF88FFBCCA10FADF78C6FA00" pageId="6" pageNumber="207" value="2007-06-30">30 June 2007</date>
(MPEG 31339),
<emphasis id="E6950738FF88FFBCCEDAFAFC79CCFA00" box="[399,453,1391,1416]" italics="true" pageId="6" pageNumber="207">idem</emphasis>
, 1 male, 0
<date id="A05FFDEAFF88FFBCCD6FFAE37AB7FA00" box="[570,702,1391,1416]" pageId="6" pageNumber="207" value="2007-07-04">4 July 2007</date>
, B. J. F. Silva (MPEG 31340);
<materialsCitation id="6489D177FF88FFBCCB42FAE37CA2FA25" ID-GBIF-Occurrence="1917355298" collectingDate="2014-03-11" collectionCode="MPEG" collectorName="M. A. Aguiar-Neto" country="Brazil" latitude="-1.5672305" location="Sao Jose do Gurupi" longLatPrecision="1" longitude="-46.27973" municipality="Viseu" pageId="6" pageNumber="207" specimenCode="MPEG 31323" specimenCount="1" specimenCount-male="1" stateProvince="Para">
<specimenCount id="C2E710A3FF88FFBCCB42FAE37C6DFA00" box="[1047,1124,1391,1416]" pageId="6" pageNumber="207" type="male">1 male</specimenCount>
,
<collectingMunicipality id="343A4150FF88FFBCCB3AFAFC7CBBFA00" box="[1135,1202,1391,1416]" pageId="6" pageNumber="207">Viseu</collectingMunicipality>
,
<location id="D13E8DF1FF88FFBCCBE9FAE37D91FA00" LSID="urn:lsid:plazi:treatment:5C486A3CFF88FFB2CFC2FDAC7D64F8FB:D13E8DF1FF88FFBCCBE9FAE37D91FA00" box="[1212,1432,1391,1416]" country="Brazil" latitude="-1.5672305" longLatPrecision="1" longitude="-46.27973" municipality="Viseu" name="Sao Jose do Gurupi" pageId="6" pageNumber="207" stateProvince="Para">São José do Gurupi</location>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCFC2FA077925FA24" box="[151,300,1428,1452]" direction="south" orientation="latitude" pageId="6" pageNumber="207" precision="1" value="-1.5672305">1°34'02.03''S</geoCoordinate>
,
<geoCoordinate id="B1D5BDEDFF88FFBCCE68FA0779E1FA25" box="[317,488,1428,1453]" direction="west" orientation="longitude" pageId="6" pageNumber="207" precision="1" value="-46.27973">46°16'47.04''W</geoCoordinate>
,
<collectingDate id="B01B0402FF88FFBCCEAFFA077AB8FA24" box="[506,689,1428,1453]" pageId="6" pageNumber="207" value="2014-03-11">
<date id="A05FFDEAFF88FFBCCEAFFA077AB8FA24" box="[506,689,1428,1453]" pageId="6" pageNumber="207" value="2014-03-11">11 March 2014</date>
</collectingDate>
,
<collectorName id="7914BEFCFF88FFBCCD97FA067BAAFA25" box="[706,931,1428,1453]" pageId="6" pageNumber="207">M. A. Aguiar-Neto</collectorName>
(
<specimenCode id="84477351FF88FFBCCCE3FA077C55FA24" box="[950,1116,1428,1452]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="6" pageNumber="207">MPEG 31323</specimenCode>
),
<emphasis id="E6950738FF88FFBCCB20FA077CA2FA25" box="[1141,1195,1428,1453]" italics="true" pageId="6" pageNumber="207">idem</emphasis>
</materialsCitation>
, 1 female, (MPEG 31326);
<emphasis id="E6950738FF88FFBCCFA0FA24793BFA58" box="[245,306,1463,1488]" italics="true" pageId="6" pageNumber="207">idem,</emphasis>
1 male, -1°33'16.09''S, 46°16'18.06''W (MPEG 31325).
</paragraph>
</subSubSection>
<subSubSection id="9CFB88A1FF88FFBDCF92FA4F790DFBF7" lastPageId="7" lastPageNumber="208" pageId="6" pageNumber="207" type="diagnosis">
<paragraph id="D45EDB2AFF88FFBCCF92FA4F7990F8D4" blockId="6.[151,1437,780,2029]" pageId="6" pageNumber="207">
<emphasis id="E6950738FF88FFBCCF92FA4F7948FA7D" bold="true" box="[199,321,1500,1525]" pageId="6" pageNumber="207">Diagnosis.</emphasis>
<taxonomicName id="13E1A0A9FF88FFBCCE19FA4E79E2FA7D" authorityName="Mello-Leitao" authorityYear="1929" box="[332,491,1500,1525]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="biocellatus">
<emphasis id="E6950738FF88FFBCCE19FA4E79E2FA7D" box="[332,491,1500,1525]" italics="true" pageId="6" pageNumber="207">E. biocellatus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="13E1A0A9FF88FFBCCD26FA4E7B18FA7C" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[627,785,1501,1524]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="trispinosus">
<emphasis id="E6950738FF88FFBCCD26FA4E7B18FA7C" box="[627,785,1501,1524]" italics="true" pageId="6" pageNumber="207">E. trispinosus</emphasis>
</taxonomicName>
, but females of
<taxonomicName id="13E1A0A9FF88FFBCCC8CFA4E7C71FA7D" authorityName="Mello-Leitao" authorityYear="1929" box="[985,1144,1500,1525]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="biocellatus">
<emphasis id="E6950738FF88FFBCCC8CFA4E7C71FA7D" box="[985,1144,1500,1525]" italics="true" pageId="6" pageNumber="207">E. biocellatus</emphasis>
</taxonomicName>
are distinguished by the carapace presenting a wide brown, median longitudinal stripe, bifurcate posteriorly, and by the opisthosoma with semicircular posterior portion, anterior margin straight and not projected towards the carapace and by the posterior conical projection about half as high as the opisthosoma (
<figureCitation id="4CDAC7AFFF88FFBCCC48F9D47B71F9E8" box="[797,888,1607,1632]" captionStart="FIGURE 2" captionStartId="9.[151,250,1908,1930]" captionTargetBox="[194,1395,195,1892]" captionTargetId="figure@9.[194,1395,193,1892]" captionTargetPageId="9" captionText="FIGURE 2. Epicadinus biocellatus Mello-Leitão, 1929. A–G female (MPEG 31330); H female (MPEG 34862). A–C habitus (A dorsal, B lateral, C ventral); D–E, epigynum (D photo ventral, E SEM ventral); F–G vulva (F photo ventral, G dorsal); H live specimen in hunting position. Abbreviations, A, atrium; AC, anterior curve of copulatory duct; CO, copulatory opening; FD, fertilization duct; MS, median septum. Scale bars: A–C, 1 mm; D–G, 0.1 mm. Photo credits: H, César Favacho." httpUri="https://zenodo.org/record/1458492/files/figure.png" pageId="6" pageNumber="207">Fig. 2A</figureCitation>
). The epigynal plate is reduced, wider than long, with an almost inconspicuous atrium, median septum and copulatory openings are located within the atrium and presenting a large posterior fold (
<figureCitation id="4CDAC7AFFF88FFBCCD44F91C7A85F920" box="[529,652,1679,1704]" captionStart="FIGURE 2" captionStartId="9.[151,250,1908,1930]" captionTargetBox="[194,1395,195,1892]" captionTargetId="figure@9.[194,1395,193,1892]" captionTargetPageId="9" captionText="FIGURE 2. Epicadinus biocellatus Mello-Leitão, 1929. A–G female (MPEG 31330); H female (MPEG 34862). A–C habitus (A dorsal, B lateral, C ventral); D–E, epigynum (D photo ventral, E SEM ventral); F–G vulva (F photo ventral, G dorsal); H live specimen in hunting position. Abbreviations, A, atrium; AC, anterior curve of copulatory duct; CO, copulatory opening; FD, fertilization duct; MS, median septum. Scale bars: A–C, 1 mm; D–G, 0.1 mm. Photo credits: H, César Favacho." httpUri="https://zenodo.org/record/1458492/files/figure.png" pageId="6" pageNumber="207">Fig. 2D, E</figureCitation>
). The copulatory ducts contour the inner margin of the spermathecae (
<figureCitation id="4CDAC7AFFF88FFBCCFCAF92778F2F944" box="[159,251,1716,1741]" captionStart="FIGURE 2" captionStartId="9.[151,250,1908,1930]" captionTargetBox="[194,1395,195,1892]" captionTargetId="figure@9.[194,1395,193,1892]" captionTargetPageId="9" captionText="FIGURE 2. Epicadinus biocellatus Mello-Leitão, 1929. A–G female (MPEG 31330); H female (MPEG 34862). A–C habitus (A dorsal, B lateral, C ventral); D–E, epigynum (D photo ventral, E SEM ventral); F–G vulva (F photo ventral, G dorsal); H live specimen in hunting position. Abbreviations, A, atrium; AC, anterior curve of copulatory duct; CO, copulatory opening; FD, fertilization duct; MS, median septum. Scale bars: A–C, 1 mm; D–G, 0.1 mm. Photo credits: H, César Favacho." httpUri="https://zenodo.org/record/1458492/files/figure.png" pageId="6" pageNumber="207">Fig. 2G</figureCitation>
). Males of
<taxonomicName id="13E1A0A9FF88FFBCCE28F9267A11F945" authorityName="Mello-Leitao" authorityYear="1929" box="[381,536,1716,1741]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="biocellatus">
<emphasis id="E6950738FF88FFBCCE28F9267A11F945" box="[381,536,1716,1741]" italics="true" pageId="6" pageNumber="207">E. biocellatus</emphasis>
</taxonomicName>
are distinguished by the yellowish-brown carapace with a dark brown, median longitudinal stripe, bifurcate posteriorly, yellowish legs, and roundish opisthosoma (
<figureCitation id="4CDAC7AFFF88FFBCCB2FF9447D09F978" box="[1146,1280,1751,1776]" captionStart="FIGURE 3" captionStartId="10.[151,250,1899,1921]" captionTargetBox="[197,1402,193,1885]" captionTargetId="figure@10.[197,1402,193,1885]" captionTargetPageId="10" captionText="FIGURE 3. Epicadinus biocellatus Mello-Leitão, 1929, male (MPEG 31328). A–C habitus (A dorsal, B lateral, C ventral); D– H left male palpus (D ventral, E–F retrolateral, G slanted retrolateral, H cymbium apex, ventral). Abbreviations, BP, basal pouch; DTA, distal tibial apophysis; E, embolus; EA, embolus apex; MF, marginal furrow of cymbium; RTA, retrolateral tibial apophysis; TF, tegular furrow; TM, tibial macrosetae. Scale bars: A–C, 1 mm; D–H, 0.1 mm." httpUri="https://zenodo.org/record/1458494/files/figure.png" pageId="6" pageNumber="207">Fig. 3A, B</figureCitation>
). The palpus presents a RTA sinuous and very long, at least
<date id="A05FFDEAFF88FFBCCDECF96F7ADCF89C" box="[697,725,1788,1812]" pageId="6" pageNumber="207">3x</date>
longer than wide, with ventral fold S-shaped, DTA short and horn-shaped and dorsum of tibia with large setiferous tubercle, directed retrolaterally, harboring a very thick macroseta (
<figureCitation id="4CDAC7AFFF88FFBCCE4DF8D77985F8D4" box="[280,396,1860,1885]" captionStart="FIGURE 3" captionStartId="10.[151,250,1899,1921]" captionTargetBox="[197,1402,193,1885]" captionTargetId="figure@10.[197,1402,193,1885]" captionTargetPageId="10" captionText="FIGURE 3. Epicadinus biocellatus Mello-Leitão, 1929, male (MPEG 31328). A–C habitus (A dorsal, B lateral, C ventral); D– H left male palpus (D ventral, E–F retrolateral, G slanted retrolateral, H cymbium apex, ventral). Abbreviations, BP, basal pouch; DTA, distal tibial apophysis; E, embolus; EA, embolus apex; MF, marginal furrow of cymbium; RTA, retrolateral tibial apophysis; TF, tegular furrow; TM, tibial macrosetae. Scale bars: A–C, 1 mm; D–H, 0.1 mm." httpUri="https://zenodo.org/record/1458494/files/figure.png" pageId="6" pageNumber="207">Fig. 3E, F</figureCitation>
).
</paragraph>
<paragraph id="D45EDB2AFF88FFBDCF92F8F47BC9FC0A" blockId="6.[151,1437,780,2029]" lastBlockId="7.[151,1437,151,2016]" lastPageId="7" lastPageNumber="208" pageId="6" pageNumber="207">
<emphasis id="E6950738FF88FFBCCF92F8F47903F808" bold="true" box="[199,266,1895,1920]" pageId="6" pageNumber="207">Notes</emphasis>
.
<bibRefCitation id="B070A6DBFF88FFBCCE4FF8F47A0BF808" box="[282,514,1895,1920]" pageId="6" pageNumber="207" refId="ref24568" refString="Mello-Leitao, C. F. (1929) Aphantochilidas e Thomisidas do Brasil. Arquivos do Museu Nacional do Rio de Janeiro, 31, 9 - 359." type="journal article">Mello-Leitão (1929)</bibRefCitation>
cited “
<collectingRegion id="162515C8FF88FFBCCD02F8FB7A84F808" box="[599,653,1896,1920]" country="Brazil" name="Para" pageId="6" pageNumber="207">Para</collectingRegion>
” as the
<typeStatus id="0B5A6588FF88FFBCCDB8F8FA7B14F809" box="[749,797,1897,1921]" pageId="6" pageNumber="207">type</typeStatus>
locality of
<taxonomicName id="13E1A0A9FF88FFBCCCF0F8FA7C4BF808" authorityName="Mello-Leitao" authorityYear="1929" box="[933,1090,1895,1920]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="biocellatus">
<emphasis id="E6950738FF88FFBCCCF0F8FA7C4BF808" box="[933,1090,1895,1920]" italics="true" pageId="6" pageNumber="207">E. biocellatus</emphasis>
</taxonomicName>
. However, the analysis of the
<typeStatus id="0B5A6588FF88FFBCCFC2F81F78F0F82D" box="[151,249,1932,1957]" pageId="6" pageNumber="207" type="holotype">holotype</typeStatus>
in Simon Collection (accession number 2044b, MNHN,
<figureCitation id="4CDAC7AFFF88FFBCCC2AF81F7BF5F82C" box="[895,1020,1932,1957]" captionStart="FIGURE 4" captionStartId="11.[151,250,942,964]" captionTargetBox="[151,1436,193,933]" captionTargetId="figure@11.[151,1436,193,933]" captionTargetPageId="11" captionText="FIGURE 4. Epicadinus biocellatus female holotype (MHNH AR-15077, Coll. Simon 2044b) [total length: 4 mm]. A dorsal habitus; B venter of opisthosoma; C–E original labels of the splitted typical lot (C female holotype 2044b, D misidentified male of E. biocellatus 2044, E female of E. cornutus 2044a). Photo credits: Christine Rollard and MNHN Laboratory of Arachnology crew." httpUri="https://zenodo.org/record/1458496/files/figure.png" pageId="6" pageNumber="207">Figs 4A–B</figureCitation>
) shows that its label, handwritten by Mello-Leitão himself, clearly states &quot;
<collectingRegion id="162515C8FF88FFBCCD67F8237A6EF840" box="[562,615,1968,1992]" country="Brazil" name="Para" pageId="6" pageNumber="207">Para</collectingRegion>
ou Pebas
<specimenCount id="C2E710A3FF88FFBCCD8DF8227AE4F840" box="[728,749,1969,1992]" pageId="6" pageNumber="207" type="female">♀</specimenCount>
<typeStatus id="0B5A6588FF88FFBCCDA6F8237B2FF840" box="[755,806,1968,1992]" pageId="6" pageNumber="207">type</typeStatus>
&quot; (
<figureCitation id="4CDAC7AFFF88FFBCCC6BF8237B9CF840" box="[830,917,1968,1993]" captionStart="FIGURE 4" captionStartId="11.[151,250,942,964]" captionTargetBox="[151,1436,193,933]" captionTargetId="figure@11.[151,1436,193,933]" captionTargetPageId="11" captionText="FIGURE 4. Epicadinus biocellatus female holotype (MHNH AR-15077, Coll. Simon 2044b) [total length: 4 mm]. A dorsal habitus; B venter of opisthosoma; C–E original labels of the splitted typical lot (C female holotype 2044b, D misidentified male of E. biocellatus 2044, E female of E. cornutus 2044a). Photo credits: Christine Rollard and MNHN Laboratory of Arachnology crew." httpUri="https://zenodo.org/record/1458496/files/figure.png" pageId="6" pageNumber="207">Fig. 4C</figureCitation>
), indicating that the specimen was collected in one of those two localities. The word &quot;
<collectingRegion id="162515C8FF88FFBCCD0FF8477A98F864" box="[602,657,2004,2028]" country="Brazil" name="Para" pageId="6" pageNumber="207">Para</collectingRegion>
&quot; or &quot;Le
<collectingRegion id="162515C8FF88FFBCCDADF8477B24F864" box="[760,813,2004,2028]" country="Brazil" name="Para" pageId="6" pageNumber="207">Para</collectingRegion>
&quot; was a reference used by Simon and other nineteenth century collectors to refer not to the Brazilian state of
<collectingRegion id="162515C8FF89FFBDCDA8FF0B7B3BFF38" box="[765,818,152,176]" country="Brazil" name="Para" pageId="7" pageNumber="208">Pará</collectingRegion>
, but to the old name of the city of Belém: Grão Belém do
<collectingRegion id="162515C8FF89FFBDCFEFFF2F78E5FF5C" box="[186,236,188,212]" country="Brazil" name="Para" pageId="7" pageNumber="208">Pará</collectingRegion>
(
<emphasis id="E6950738FF89FFBDCFA8FF2E792CFF5C" box="[253,293,189,212]" italics="true" pageId="7" pageNumber="208">e.g.</emphasis>
<bibRefCitation id="B070A6DBFF89FFBDCE78FF2F79ADFF5D" box="[301,420,188,213]" pageId="7" pageNumber="208" refId="ref24189" refString="Levi, H. W. (1964) American spiders of the genus Phoroncidia (Araneae: Theridiidae). Bulletin of the Museum of Comparative Zoology, 131, 65 - 86." type="journal article">Levi 1964</bibRefCitation>
: 72,
<bibRefCitation id="B070A6DBFF89FFBDCE8FFF2E7A8FFF5C" box="[474,646,188,212]" pageId="7" pageNumber="208" refId="ref24718" refString="Papavero, N. (1973) Essays on the History of Neotropical Dipterology, with Special Reference to Collectors (1750 - 1905). Uol. 2. Museu de Zoologia da USP, Sao Paulo, 230 pp. [pp. 217 - 446]" type="book chapter">Papavero 1973</bibRefCitation>
: 377,
<bibRefCitation id="B070A6DBFF89FFBDCD9FFF2F7B48FF5D" box="[714,833,188,213]" pageId="7" pageNumber="208" refId="ref24224" refString="Levi, H. W. (1996) The American orb weavers Hypognatha, Encyosaccus, Xylethrus, Gasteracantha, and Enacrosoma (Araneae, Araneidae). Bulletin of the Museum of Comparative Zoology, 155, 89 - 157." type="journal article">Levi 1996</bibRefCitation>
: 108, 142). Simon normally used the word “Prov.” or “Province” when referring to the current state of
<collectingRegion id="162515C8FF89FFBDCD9FFF737AF5FF70" box="[714,764,224,248]" country="Brazil" name="Para" pageId="7" pageNumber="208">Pará</collectingRegion>
(
<emphasis id="E6950738FF89FFBDCC5BFF727B3FFF70" box="[782,822,225,248]" italics="true" pageId="7" pageNumber="208">e.g.</emphasis>
<bibRefCitation id="B070A6DBFF89FFBDCC6AFF4C7BC4FF70" box="[831,973,223,248]" pageId="7" pageNumber="208" refId="ref25263" refString="Simon, E. (1895) Histoire Naturelle des Araignees. 2 Edition. Tome 1. Librairie Encyclopedique de Roret, Paris, 224 pp. [pp. 861 - 1084.]" type="book chapter">Simon 1895</bibRefCitation>
: 159). The other locality is Pebas, a city near the Amazon River, in the department of
<collectingRegion id="162515C8FF89FFBDCDECFE967B0FFE94" box="[697,774,261,284]" country="Peru" name="Loreto" pageId="7" pageNumber="208">Loreto</collectingRegion>
,
<collectingCountry id="ACF69BBAFF89FFBDCC42FE967B44FE94" box="[791,845,261,284]" name="Peru" pageId="7" pageNumber="208">Peru</collectingCountry>
, near the border with
<collectingCountry id="ACF69BBAFF89FFBDCB0EFE977CAAFE95" box="[1115,1187,260,285]" name="Brazil" pageId="7" pageNumber="208">Brazil</collectingCountry>
. This specimen most probably was collected by Marc de Mathan, a French collector who followed the Amazon River and its tributaries, from Belém to northern
<collectingCountry id="ACF69BBAFF89FFBDCEE1FEC379E3FEEF" box="[436,490,336,359]" name="Peru" pageId="7" pageNumber="208">Peru</collectingCountry>
, reaching up to
<collectingCountry id="ACF69BBAFF89FFBDCDFEFEDD7B03FEEF" box="[683,778,334,359]" name="Ecuador" pageId="7" pageNumber="208">Ecuador</collectingCountry>
, during the end of the 19th and the beginning of the 20th centuries (
<bibRefCitation id="B070A6DBFF89FFBDCE59FEE079FAFE03" box="[268,499,370,395]" pageId="7" pageNumber="208" refId="ref24662" refString="Papavero, N. (1971) Essays on the History of Neotropical Dipterology, with Special Reference to Collectors (1750 - 1905). Uol. 1. Museu de Zoologia da USP, Sao Paulo, 216 pp. https: // doi. org / 10.5962 / bhl. title. 101715" type="book">Papavero 1971: 161</bibRefCitation>
). Mathan was the sole collector cited for Pebas and he also collected in “Le
<collectingRegion id="162515C8FF89FFBDCA0BFEE07D9BFE03" box="[1374,1426,371,395]" country="Brazil" name="Para" pageId="7" pageNumber="208">Para</collectingRegion>
” (
<emphasis id="E6950738FF89FFBDCFF5FE0B78C1FE27" box="[160,200,408,431]" italics="true" pageId="7" pageNumber="208">e.g.</emphasis>
<bibRefCitation id="B070A6DBFF89FFBDCF86FE05796BFE27" box="[211,354,406,431]" pageId="7" pageNumber="208" refId="ref25222" refString="Simon, E. (1893) Descriptions d'especes et de genres nouveaux de l'ordre des Araneae. Etudes arachnologiques. 25 e Memoire. XL. Annales de la Societe Entomologique de France, 62, 299 - 330." type="journal article">Simon 1893</bibRefCitation>
: 310, 326; 1895: 135, 153). Unfortunately, spider samples deposited in MNHN, identified as belonging to a single species, often refer to more than one locality and may have been mixed in just one flask (R. Baptista, personal observation). Simon probably had identified three specimens as
<taxonomicName id="13E1A0A9FF89FFBDCB36FE737D08FE7F" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[1123,1281,480,503]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="trispinosus">
<emphasis id="E6950738FF89FFBDCB36FE737D08FE7F" box="[1123,1281,480,503]" italics="true" pageId="7" pageNumber="208">E. trispinosus</emphasis>
</taxonomicName>
, all included under accession number 2044, although they belonged to different samples from Belém and Pebas (
<figureCitation id="4CDAC7AFFF89FFBDCA64FD917D99FD93" box="[1329,1424,514,539]" captionStart="FIGURE 4" captionStartId="11.[151,250,942,964]" captionTargetBox="[151,1436,193,933]" captionTargetId="figure@11.[151,1436,193,933]" captionTargetPageId="11" captionText="FIGURE 4. Epicadinus biocellatus female holotype (MHNH AR-15077, Coll. Simon 2044b) [total length: 4 mm]. A dorsal habitus; B venter of opisthosoma; C–E original labels of the splitted typical lot (C female holotype 2044b, D misidentified male of E. biocellatus 2044, E female of E. cornutus 2044a). Photo credits: Christine Rollard and MNHN Laboratory of Arachnology crew." httpUri="https://zenodo.org/record/1458496/files/figure.png" pageId="7" pageNumber="208">Fig. 4D</figureCitation>
).
<bibRefCitation id="B070A6DBFF89FFBDCFC2FDB57976FDB7" box="[151,383,550,575]" pageId="7" pageNumber="208" refId="ref24568" refString="Mello-Leitao, C. F. (1929) Aphantochilidas e Thomisidas do Brasil. Arquivos do Museu Nacional do Rio de Janeiro, 31, 9 - 359." type="journal article">Mello-Leitão (1929)</bibRefCitation>
considered these specimens as belonging to three different species and splitted this mixed sample into three log numbers: one with a male (2044) that he also identified as
<taxonomicName id="13E1A0A9FF89FFBDCB05FDDF7CE7FDEB" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[1104,1262,588,611]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="trispinosus">
<emphasis id="E6950738FF89FFBDCB05FDDF7CE7FDEB" box="[1104,1262,588,611]" italics="true" pageId="7" pageNumber="208">E. trispinosus</emphasis>
</taxonomicName>
, agreeing with Simon (
<figureCitation id="4CDAC7AFFF89FFBDCFA5FDFD7943FD0F" box="[240,330,622,647]" captionStart="FIGURE 4" captionStartId="11.[151,250,942,964]" captionTargetBox="[151,1436,193,933]" captionTargetId="figure@11.[151,1436,193,933]" captionTargetPageId="11" captionText="FIGURE 4. Epicadinus biocellatus female holotype (MHNH AR-15077, Coll. Simon 2044b) [total length: 4 mm]. A dorsal habitus; B venter of opisthosoma; C–E original labels of the splitted typical lot (C female holotype 2044b, D misidentified male of E. biocellatus 2044, E female of E. cornutus 2044a). Photo credits: Christine Rollard and MNHN Laboratory of Arachnology crew." httpUri="https://zenodo.org/record/1458496/files/figure.png" pageId="7" pageNumber="208">Fig. 4D</figureCitation>
), another with a female of
<taxonomicName id="13E1A0A9FF89FFBDCD2FFDE37AF5FD0F" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[634,764,624,647]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="E6950738FF89FFBDCD2FFDE37AF5FD0F" box="[634,764,624,647]" italics="true" pageId="7" pageNumber="208">E. cornutus</emphasis>
</taxonomicName>
(2044a,
<figureCitation id="4CDAC7AFFF89FFBDCC35FDFD7BBFFD0F" box="[864,950,622,647]" captionStart="FIGURE 4" captionStartId="11.[151,250,942,964]" captionTargetBox="[151,1436,193,933]" captionTargetId="figure@11.[151,1436,193,933]" captionTargetPageId="11" captionText="FIGURE 4. Epicadinus biocellatus female holotype (MHNH AR-15077, Coll. Simon 2044b) [total length: 4 mm]. A dorsal habitus; B venter of opisthosoma; C–E original labels of the splitted typical lot (C female holotype 2044b, D misidentified male of E. biocellatus 2044, E female of E. cornutus 2044a). Photo credits: Christine Rollard and MNHN Laboratory of Arachnology crew." httpUri="https://zenodo.org/record/1458496/files/figure.png" pageId="7" pageNumber="208">Fig. 4E</figureCitation>
), and the last one with the female
<typeStatus id="0B5A6588FF89FFBDCA6FFDFD7D95FD0F" box="[1338,1436,622,647]" pageId="7" pageNumber="208" type="holotype">holotype</typeStatus>
of
<taxonomicName id="13E1A0A9FF89FFBDCFE3FD077958FD23" authorityName="Mello-Leitao" authorityYear="1929" box="[182,337,658,683]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="biocellatus">
<emphasis id="E6950738FF89FFBDCFE3FD077958FD23" box="[182,337,658,683]" italics="true" pageId="7" pageNumber="208">E. biocellatus</emphasis>
</taxonomicName>
(2044b,
<figureCitation id="4CDAC7AFFF89FFBDCEE2FD017A3FFD23" box="[439,566,658,683]" captionStart="FIGURE 4" captionStartId="11.[151,250,942,964]" captionTargetBox="[151,1436,193,933]" captionTargetId="figure@11.[151,1436,193,933]" captionTargetPageId="11" captionText="FIGURE 4. Epicadinus biocellatus female holotype (MHNH AR-15077, Coll. Simon 2044b) [total length: 4 mm]. A dorsal habitus; B venter of opisthosoma; C–E original labels of the splitted typical lot (C female holotype 2044b, D misidentified male of E. biocellatus 2044, E female of E. cornutus 2044a). Photo credits: Christine Rollard and MNHN Laboratory of Arachnology crew." httpUri="https://zenodo.org/record/1458496/files/figure.png" pageId="7" pageNumber="208">Figs 4A–C</figureCitation>
). The male identified as
<taxonomicName id="13E1A0A9FF89FFBDCC18FD077C32FD23" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[845,1083,659,683]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="trispinosus">
<emphasis id="E6950738FF89FFBDCC18FD077BE1FD23" box="[845,1000,660,683]" italics="true" pageId="7" pageNumber="208">E. trispinosus</emphasis>
(2044)
</taxonomicName>
by Mello-Leitão is, in fact, the then undescribed male of
<taxonomicName id="13E1A0A9FF89FFBDCE95FD2B7A55FD47" authorityName="Mello-Leitao" authorityYear="1929" box="[448,604,694,719]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="biocellatus">
<emphasis id="E6950738FF89FFBDCE95FD2B7A55FD47" box="[448,604,694,719]" italics="true" pageId="7" pageNumber="208">E. biocellatus</emphasis>
</taxonomicName>
, as many samples of the same males and females from several localities
<collectingRegion id="162515C8FF89FFBDCFC2FD4878C0FD7B" box="[151,201,731,755]" country="Brazil" name="Para" pageId="7" pageNumber="208">Pará</collectingRegion>
suggest. The other female (2044a) actually is
<taxonomicName id="13E1A0A9FF89FFBDCD99FD4F7B44FD7B" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[716,845,732,755]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="E6950738FF89FFBDCD99FD4F7B44FD7B" box="[716,845,732,755]" italics="true" pageId="7" pageNumber="208">E. cornutus</emphasis>
</taxonomicName>
, but Mello-Leitão committed a lapsus while copying the label, including only &quot;
<collectingRegion id="162515C8FF89FFBDCEEBFD6C79FEFC9F" box="[446,503,767,791]" country="Brazil" name="Para" pageId="7" pageNumber="208">Para</collectingRegion>
,&quot; without referring to &quot;Pebas&quot;. In MNHN, there is a single female specimen of
<taxonomicName id="13E1A0A9FF89FFBDCAD1FC93797DFCB3" authorityName="Mello-Leitao" authorityYear="1929" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="biocellatus">
<emphasis id="E6950738FF89FFBDCAD1FC93791BFCB3" italics="true" pageId="7" pageNumber="208">E. biocellatus</emphasis>
(2044b)
</taxonomicName>
, indicating that the species description was not based on a
<typeStatus id="0B5A6588FF89FFBDCB51FCB07C3DFCB3" box="[1028,1076,803,827]" pageId="7" pageNumber="208">type</typeStatus>
series, but on a single specimen collected either in
<collectingRegion id="162515C8FF89FFBDCE30FCD4799EFCD7" box="[357,407,839,863]" country="Brazil" name="Para" pageId="7" pageNumber="208">Pará</collectingRegion>
or Pebas. The male of
<taxonomicName id="13E1A0A9FF89FFBDCDC2FCDB7B8BFCD7" authorityName="Mello-Leitao" authorityYear="1929" box="[663,898,838,863]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="biocellatus">
<emphasis id="E6950738FF89FFBDCDC2FCDB7B38FCD7" box="[663,817,838,863]" italics="true" pageId="7" pageNumber="208">E. biocellatus</emphasis>
(2044)
</taxonomicName>
included in the same flask was not described by Mello-Leitão, since this author mistakenly attributed it to
<taxonomicName id="13E1A0A9FF89FFBDCC4BFCFF7BB0FC0B" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[798,953,876,899]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="trispinosus">
<emphasis id="E6950738FF89FFBDCC4BFCFF7BB0FC0B" box="[798,953,876,899]" italics="true" pageId="7" pageNumber="208">E. trispinosus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D45EDB2AFF89FFBDCF92FC03790DFBF7" blockId="7.[151,1437,151,2016]" pageId="7" pageNumber="208">
We assume that the true locality of the species is actually Belém, since many specimens were collected in this locality and surrounding areas of
<collectingRegion id="162515C8FF89FFBDCD7CFC207A52FC43" box="[553,603,947,971]" country="Brazil" name="Para" pageId="7" pageNumber="208">Pará</collectingRegion>
and
<collectingRegion id="162515C8FF89FFBDCDC9FC217B07FC43" box="[668,782,946,971]" country="Brazil" name="Maranhao" pageId="7" pageNumber="208">Maranhão</collectingRegion>
states, both in
<collectingCountry id="ACF69BBAFF89FFBDCC99FC217C1DFC43" box="[972,1044,946,971]" name="Brazil" pageId="7" pageNumber="208">Brazil</collectingCountry>
. In addition, no specimens of
<taxonomicName id="13E1A0A9FF89FFBDCAD1FC27791BFC67" authorityName="Mello-Leitao" authorityYear="1929" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="biocellatus">
<emphasis id="E6950738FF89FFBDCAD1FC27791BFC67" italics="true" pageId="7" pageNumber="208">E. biocellatus</emphasis>
</taxonomicName>
from western Amazon, a region where Pebas is located, were found in the collections examined. Conversely, the female of
<taxonomicName id="13E1A0A9FF89FFBDCEEEFC6F7A90FB9B" baseAuthorityName="Taczanowski" baseAuthorityYear="1872" box="[443,665,1019,1043]" class="Arachnida" family="Thomisidae" genus="Epicadinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="E6950738FF89FFBDCEEEFC6F7A32FB9B" box="[443,571,1020,1043]" italics="true" pageId="7" pageNumber="208">E. cornutus</emphasis>
(2044a)
</taxonomicName>
is most probably the specimen that was originally collected in Pebas, since no sample of this species has been collected in
<collectingRegion id="162515C8FF89FFBDCDA4FB8C7B2AFBBF" box="[753,803,1055,1079]" country="Brazil" name="Para" pageId="7" pageNumber="208">Pará</collectingRegion>
or other areas of eastern Amazon, and there are several records from western Amazon, surrounding the
<collectingCountry id="ACF69BBAFF89FFBDCDEDFBD07AE2FBD2" box="[696,747,1091,1114]" name="Peru" pageId="7" pageNumber="208">Peru</collectingCountry>
locality, i. e.,
<collectingCountry id="ACF69BBAFF89FFBDCCDAFBD17BE5FBD3" box="[911,1004,1090,1115]" name="Ecuador" pageId="7" pageNumber="208">Ecuador</collectingCountry>
and the Brazilian states of
<collectingRegion id="162515C8FF89FFBDCA73FBD07D95FBD3" box="[1318,1436,1091,1115]" country="Peru" name="Amazonas" pageId="7" pageNumber="208">Amazonas</collectingRegion>
and
<collectingRegion id="162515C8FF89FFBDCF93FBF47909FBF7" box="[198,256,1127,1151]" country="Brazil" name="Acre" pageId="7" pageNumber="208">Acre</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="9CFB88A1FF89FFB2CF92FB197B6DF8C7" lastPageId="8" lastPageNumber="209" pageId="7" pageNumber="208" type="description">
<paragraph id="D45EDB2AFF89FFBDCF92FB197BF2F9FF" blockId="7.[151,1437,151,2016]" pageId="7" pageNumber="208">
<emphasis id="E6950738FF89FFBDCF92FB197A8EFB2B" bold="true" box="[199,647,1162,1187]" pageId="7" pageNumber="208">Description. Female (MPEG 31330):</emphasis>
Total length: 4.39. Carapace: length 1.92, width 1.80, height 1.19. Clypeus: height 0.25. Chelicera: length 0.57, width 0.27. Sternum: length 0.87, width 0.69. Endites: length 0.28, width, 0.25. Labium: length 0.15, width 0.30. Eye diameters and interdistances: AME 0.07, ALE 0.08, PME 0.06, PLE 0.05, AME–AME 0.09, AME–ALE 0.07, ALE–ALE 0.23, PME–PME 0.12, PME–PLE 0.06, PLE–PLE 0.36. MOQ trapezoidal. Leg formula: 1243. Leg I – femur 1.69, patella 0.94, tibia 1.85, metatarsus 1.46, tarsus 0.69, total 6.63; II: 1.46-0.74-1.68-1.33-0.63-5.84; III: 0.66-0.51-0.73-0.60-0.40-2.90; IV: 1.08-0.48-0.86-0.70-0.45- 3.57. Paired setal tubercles on tibiae and metatarsi with length less than or equal to half the article width. Opisthosoma: length 2.47, width 2.81, height 3.28, with trapezoidal anterior half and semicircular posterior half in dorsal view, covered by abundant macrosetae of variable size. Anterior dorsal margin not projecting over posterior portion of carapace, straight in dorsal view, with anterior angles not projected. Paired conical projections smaller in median portion, directed posterolaterally. Posterior conical projection placed a bit more posteriorly, in almost central position, and dorsally directed, nearly perpendicular to venter, about twice as long as paired ones and measuring about half the height of the opisthosoma itself. In lateral view, dorsum very convex both before and after posterior conical projection. Venter with anterior portion profusely setaceous.
</paragraph>
<paragraph id="D45EDB2AFF89FFB2CF92F9117BCBFEEF" blockId="7.[151,1437,151,2016]" lastBlockId="8.[151,1437,151,1907]" lastPageId="8" lastPageNumber="209" pageId="7" pageNumber="208">
Epigynum (
<figureCitation id="4CDAC7AFFF89FFBDCE1EF91179C1F913" box="[331,456,1666,1691]" captionStart="FIGURE 2" captionStartId="9.[151,250,1908,1930]" captionTargetBox="[194,1395,195,1892]" captionTargetId="figure@9.[194,1395,193,1892]" captionTargetPageId="9" captionText="FIGURE 2. Epicadinus biocellatus Mello-Leitão, 1929. A–G female (MPEG 31330); H female (MPEG 34862). A–C habitus (A dorsal, B lateral, C ventral); D–E, epigynum (D photo ventral, E SEM ventral); F–G vulva (F photo ventral, G dorsal); H live specimen in hunting position. Abbreviations, A, atrium; AC, anterior curve of copulatory duct; CO, copulatory opening; FD, fertilization duct; MS, median septum. Scale bars: A–C, 1 mm; D–G, 0.1 mm. Photo credits: H, César Favacho." httpUri="https://zenodo.org/record/1458492/files/figure.png" pageId="7" pageNumber="208">Figs 2D, E</figureCitation>
) slightly wider than long, with septum and copulatory openings located within atrium, separating it into two parts, and a large flattened posterior area, about 1.5x longer than atrium. Epigynum small, occupying a much smaller area than the one occupied by internal genitalia. Atrium reniform, very wide, approximately
<date id="A05FFDEAFF89FFBDCE14F97C7954F88F" box="[321,349,1775,1799]" pageId="7" pageNumber="208">3x</date>
wider than long, very shallow, almost inconspicuous, delimited by a light marginal depression, strongly demarcated on anterior margin and inconspicuous on most of posterior margin. Median septum almost as wide as long, placed almost entirely inside and fused to median area of atrium, which is restricted to only two free lateral areas. Anterior base of septum entirely within atrium and slightly larger than posterior base, which is located just below atrium. Copulatory openings clearly visible, located in depressions at each side of median portion of septum. Vulva (
<figureCitation id="4CDAC7AFFF89FFBDCE1DF83179B2F833" box="[328,443,1954,1979]" captionStart="FIGURE 2" captionStartId="9.[151,250,1908,1930]" captionTargetBox="[194,1395,195,1892]" captionTargetId="figure@9.[194,1395,193,1892]" captionTargetPageId="9" captionText="FIGURE 2. Epicadinus biocellatus Mello-Leitão, 1929. A–G female (MPEG 31330); H female (MPEG 34862). A–C habitus (A dorsal, B lateral, C ventral); D–E, epigynum (D photo ventral, E SEM ventral); F–G vulva (F photo ventral, G dorsal); H live specimen in hunting position. Abbreviations, A, atrium; AC, anterior curve of copulatory duct; CO, copulatory opening; FD, fertilization duct; MS, median septum. Scale bars: A–C, 1 mm; D–G, 0.1 mm. Photo credits: H, César Favacho." httpUri="https://zenodo.org/record/1458492/files/figure.png" pageId="7" pageNumber="208">Fig. 2F, G</figureCitation>
) with short ventral ducts, connected posteriorly to copulatory openings and anteriorly to internal posterior edge of large reniform anterior curve, which is very expanded. From external posterior angle of anterior curve, a long duct originates, directed internally at its initial portion, after which there is a closed curve, directed ventrally and contouring internal margin of spermatheca, reaching down close to ventral surface. A 90o curve follows, directed posteriorly, forming a long and almost straight path, clearly visible by transparency (
<figureCitation id="4CDAC7AFFF86FFB2CA3AFF7178B5FE97" captionStart="FIGURE 2" captionStartId="9.[151,250,1908,1930]" captionTargetBox="[194,1395,195,1892]" captionTargetId="figure@9.[194,1395,193,1892]" captionTargetPageId="9" captionText="FIGURE 2. Epicadinus biocellatus Mello-Leitão, 1929. A–G female (MPEG 31330); H female (MPEG 34862). A–C habitus (A dorsal, B lateral, C ventral); D–E, epigynum (D photo ventral, E SEM ventral); F–G vulva (F photo ventral, G dorsal); H live specimen in hunting position. Abbreviations, A, atrium; AC, anterior curve of copulatory duct; CO, copulatory opening; FD, fertilization duct; MS, median septum. Scale bars: A–C, 1 mm; D–G, 0.1 mm. Photo credits: H, César Favacho." httpUri="https://zenodo.org/record/1458492/files/figure.png" pageId="8" pageNumber="209">Fig. 2D</figureCitation>
), and reaching back to near epigastric furrow, then an abrupt and long curve directed dorsally occurs, followed by another one directed anteriorly, originating a short straight path and a curve directed externally, which finally connects to internal posterior margin of large and roundish spermatheca.
</paragraph>
<paragraph id="D45EDB2AFF86FFB2CF92FEE17A0DFD0F" blockId="8.[151,1437,151,1907]" pageId="8" pageNumber="209">Coloration. Carapace yellow with a brown median longitudinal stripe, bifurcate posteriorly, not reaching margin of carapace. Light brown irregular stripes at margins. Ocular mound brown. Sternum, endites and labium mostly light yellow. Labium and sternum with light brown margins. Chelicerae light yellow with brown median stripe at each one. Legs mostly yellow with light brown irregular spots. Legs III and IV lighter than I and II and with less spots. Opisthosoma mostly yellow, lighter in folium and conical projections, where there are visible guanine crystals. Dorsally, one brown median stripe reaching up to posterior conical projection, and two black roundish spots near to base of each paired conical projections of opisthosoma. Laterally, black irregular spots, and ventrally yellow and white hues.</paragraph>
<paragraph id="D45EDB2AFF86FFB2CF92FD017CF5FBBF" blockId="8.[151,1437,151,1907]" pageId="8" pageNumber="209">
<emphasis id="E6950738FF86FFB2CF92FD017A66FD23" bold="true" box="[199,623,658,683]" pageId="8" pageNumber="209">
Description.
<emphasis id="E6950738FF86FFB2CE33FD017A66FD23" bold="true" box="[358,623,658,683]" italics="true" pageId="8" pageNumber="209">Male (MPEG 31328):</emphasis>
</emphasis>
Total length: 2.48. Carapace: length 1.27, width 1.21, height 0.71. Clypeus: height 0.15. Chelicera: length 0.37, width 0.17. Sternum: length 0.54, width 0.52. Endites: length 0.15, width, 0.18. Labium: length 0.06, width 0.17. Eye diameters and interdistances: AME 0.05, ALE 0.06, PME 0.05, PLE 0.04, AME–AME 0.01, AME–ALE 0.06, ALE–ALE 0.18, PME–PME 0.11, PME–PLE 0.07, PLE–PLE 0.31. MOQ trapezoidal. Legs 1243. Leg I: femur 1.11, patella 0.47, tibia 0.94, metatarsus 0.82, tarsus 0.46, total 3.80; II: 1.02-0.44-0.90-0.70-0.44-3.50; III: 0.50-0.28-0.43-0.33-0.27-1.81; IV: 0.62-0.32-0.53-0.39-0.30-2.13. Paired setal tubercles in tibiae and metatarsi near half to almost as long as the width of the article. Opisthosoma: length 1.21, width 1.67, height 1.28, with trapezoidal anterior half and semicircular posterior half in dorsal view, covered by abundant macrosetae of variable size. Anterior margin of dorsum straight, slightly projecting over posterior portion of carapace, with anterior angles not projected. Paired projections in median portion of opisthosoma, directed posterolaterally. Posterior conical projection placed more posteriorly than in female, and directed both dorsally and posteriorly. In lateral view, dorsum very convex both before and behind posterior conical projection.
</paragraph>
<paragraph id="D45EDB2AFF86FFB2CF92FBD17924F9DB" blockId="8.[151,1437,151,1907]" pageId="8" pageNumber="209">
Palpus (
<figureCitation id="4CDAC7AFFF86FFB2CE72FBD179B7FBD3" box="[295,446,1090,1115]" captionStart="FIGURE 3" captionStartId="10.[151,250,1899,1921]" captionTargetBox="[197,1402,193,1885]" captionTargetId="figure@10.[197,1402,193,1885]" captionTargetPageId="10" captionText="FIGURE 3. Epicadinus biocellatus Mello-Leitão, 1929, male (MPEG 31328). A–C habitus (A dorsal, B lateral, C ventral); D– H left male palpus (D ventral, E–F retrolateral, G slanted retrolateral, H cymbium apex, ventral). Abbreviations, BP, basal pouch; DTA, distal tibial apophysis; E, embolus; EA, embolus apex; MF, marginal furrow of cymbium; RTA, retrolateral tibial apophysis; TF, tegular furrow; TM, tibial macrosetae. Scale bars: A–C, 1 mm; D–H, 0.1 mm." httpUri="https://zenodo.org/record/1458494/files/figure.png" pageId="8" pageNumber="209">Figs 3 D–H</figureCitation>
). Tegulum with transversal furrow in 3 o’clock position. Embolus basis gradually separating from tegulum, initially forming a wide stripe, that tapers up to filiform apex, describing about 3/4 of a turn around lateral margin of tegulum, and with apex located on a furrow placed on prolateral margin glabrous apical area of cymbium. RTA fused to DTA, both connected directly to tibia and not constricted at base. Short Vshaped notch between RTA and DTA. RTA wide and very elongated, about
<date id="A05FFDEAFF86FFB2CCB5FB407BF5FB63" box="[992,1020,1235,1259]" pageId="8" pageNumber="209">3x</date>
longer than wide, with curved lateral margins, arranged transversally to longitudinal axis of cymbium, large median concavity, wide marginal folds, and rounded apex thinner than its median portion. RTA with longitudinal axis very slanted downward and also projecting retrolaterally, especially at its distal half. Its dorsal fold describes a long and convex curve and projects over median concavity, forming a deep marginal trough. Ventral fold describes a S-shaped curve, with concave distal portion and convex basal portion, projected over median depression only at basal area, where it extends dorsally and fuses to common base RTA + DTA, forming a deep basal pouch around DTA and dorsal fold of RTA. Basal pouch conspicuous in ventral view and with evident grooves, numerous and close to each other along its length, which give a striated aspect to surface. DTA very short, horn shaped, a little blunt, with slightly thicker base. Dorsum of tibia with large setal tubercle, surmounted by a very strong spiniform macroseta, inclined retrolaterally.
</paragraph>
<paragraph id="D45EDB2AFF86FFB2CF92F9CD7B6DF8C7" blockId="8.[151,1437,151,1907]" pageId="8" pageNumber="209">Coloration. Carapace orange yellow with a wide longitudinal dark brown median stripe, bifurcated at its posterior portion, and with irregular dark brown stripes at margins. Sternum entirely yellow; endites and labium predominantly yellow with brown margin; chelicerae light yellow with a median longitudinal brown stripe near the fang. Legs predominantly yellow with irregular light brown rings and spots. Legs III and IV lighter than I and II, and with less spots. Opisthosoma pale brown, dorsum with a trapezoidal brown folium between a white area stained by guanine. Conical projections dark yellow, also with basal guanine spots. Sides predominantly pale brown with black hues and venter pale brown with yellow hues.</paragraph>
</subSubSection>
<subSubSection id="9CFB88A1FF86FFB2CF92F8C97D64F8FB" box="[199,1389,1882,1907]" pageId="8" pageNumber="209" type="distribution">
<paragraph id="D45EDB2AFF86FFB2CF92F8C97D64F8FB" blockId="8.[151,1437,151,1907]" box="[199,1389,1882,1907]" pageId="8" pageNumber="209">
<emphasis id="E6950738FF86FFB2CF92F8C97950F8FB" bold="true" box="[199,345,1882,1907]" pageId="8" pageNumber="209">Distribution</emphasis>
. Only know from eastern Amazon, from the Brazilian states of
<collectingRegion id="162515C8FF86FFB2CB71F8C87C5FF8FB" box="[1060,1110,1883,1907]" country="Brazil" name="Para" pageId="8" pageNumber="209">Pará</collectingRegion>
and
<collectingRegion id="162515C8FF86FFB2CBD9F8C97CF4F8FB" box="[1164,1277,1882,1907]" country="Brazil" name="Maranhao" pageId="8" pageNumber="209">Maranhão</collectingRegion>
(
<figureCitation id="4CDAC7AFFF86FFB2CA58F8C97D69F8FB" box="[1293,1376,1882,1907]" captionStart="FIGURE 15" captionStartId="31.[151,250,1300,1322]" captionTargetBox="[166,1420,208,1263]" captionTargetId="figure@31.[150,1436,193,1278]" captionTargetPageId="31" captionText="FIGURE 15. Geographic distribution records of Epicadinus, highlighting the type-locality of each of the four species (star symbols)." httpUri="https://zenodo.org/record/1458518/files/figure.png" pageId="8" pageNumber="209">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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