180 lines
19 KiB
XML
180 lines
19 KiB
XML
<document id="FE79521441A3103EBA5E69A7F5E69E05" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FFFB6146FF0266F0FF59FE91" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madecorphnus falcatus Paulian 1992" docType="treatment" docVersion="16" lastPageNumber="45" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="43" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:mods id="7155C55E7CC6EBE1986DBF010B6587F4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AA5180AFE88A034C101154D1233C8481">
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<mods:title id="B23205CC821108FDE6CE5116E1E8A88D">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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<mods:name id="865CCCBE1CC507D94338F99B0AF6E884" type="personal">
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<mods:role id="4D005E7397FFA981B04A6CADB4821DC9">
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<mods:roleTerm id="697104DF040C83596E5EDC12047B8E4A">Author</mods:roleTerm>
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<mods:namePart id="F3C0D8D979FA749456250D17B14C0C4B">Frolov, Andrey V.</mods:namePart>
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<mods:roleTerm id="0BEB7B1DC7C0A07590A60FC6CAD69A4E">Author</mods:roleTerm>
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<mods:namePart id="A26875F06B51443F655726D98C08113C">Montreuil, Olivier</mods:namePart>
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<mods:name id="E65C799D6B24710F8474A19E4F5FBD97" type="personal">
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<mods:roleTerm id="500485DAB89D029E75BFF8EBC397DF7C">Author</mods:roleTerm>
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<mods:namePart id="437E6E3073A6B330F0D4DF9F05A58A14">Akhmetova, Lilia A.</mods:namePart>
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<mods:typeOfResource id="D5525E462EB5DCBCDD166FE695385B0A">text</mods:typeOfResource>
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<mods:title id="7BE2A9B086E8CD68EC77D39FF5888CE6">Zootaxa</mods:title>
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<mods:part id="5A20832961EE51C67FE0F76FE66F40CB">
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<mods:date id="2570D979A312C2BB276A4CCD7B935F36">2016</mods:date>
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<mods:detail id="BCCBD6AAA12BC6AC54F5F65F02459511" type="volume">
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<mods:number id="967012050825C6579F17832A99D05962">4207</mods:number>
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<mods:identifier id="690FF14FDFFBC1C820014CB2F9F28664" type="DOI">10.11646/zootaxa.4207.1.1</mods:identifier>
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<treatment id="5B4FBD00FFFB6146FF0266F0FF59FE91" ID-DOI="http://doi.org/10.5281/zenodo.6057662" ID-GBIF-Taxon="125107274" ID-Zenodo-Dep="6057662" LSID="urn:lsid:plazi:treatment:5B4FBD00FFFB6146FF0266F0FF59FE91" httpUri="http://treatment.plazi.org/id/5B4FBD00FFFB6146FF0266F0FF59FE91" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43">
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<subSubSection id="9BFC5F9DFFFB6140FF0266F0FEBCFB12" pageId="42" pageNumber="43" type="nomenclature">
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<paragraph id="D3590C16FFFB6140FF0266F0FDECFB33" blockId="42.[151,614,1196,1255]" box="[151,614,1196,1222]" pageId="42" pageNumber="43">
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<heading id="8811BB7AFFFB6140FF0266F0FDECFB33" bold="true" box="[151,614,1196,1222]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
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<taxonomicName id="14E67795FFFB6140FF0266F0FDECFB33" authority="Paulian, 1992" authorityName="Paulian" authorityYear="1992" box="[151,614,1196,1222]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="falcatus">
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<emphasis id="E192D004FFFB6140FF0266F0FDECFB33" bold="true" box="[151,614,1196,1222]" pageId="42" pageNumber="43">
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<emphasis id="E192D004FFFB6140FF0266F0FE3FFB33" bold="true" box="[151,437,1196,1222]" italics="true" pageId="42" pageNumber="43">Madecorphnus falcatus</emphasis>
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<bibRefCitation id="B77771E7FFFB6140FE2966F0FDECFB33" author="Paulian" box="[444,614,1196,1222]" pageId="42" pageNumber="43" refString="Paulian, R. (1992) Un nouveau genre et une nouvelle espece d'Orphnidae de Madagascar (Coleoptera, Scarabaeoidea). Revue Francaise d'Entomologie (N. S.), 14, 169 - 171." type="journal article" year="1992">Paulian, 1992</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="D3590C16FFFB6140FF026692FEBCFB12" blockId="42.[151,614,1196,1255]" box="[151,310,1230,1255]" pageId="42" pageNumber="43">(Figs. 21A–F)</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFFFB6140FF02674AFC1FFAD9" box="[151,917,1302,1324]" pageId="42" pageNumber="43" type="reference_group">
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<paragraph id="D3590C16FFFB6140FF02674AFC1FFAD9" blockId="42.[151,917,1302,1324]" box="[151,917,1302,1324]" pageId="42" pageNumber="43">
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<treatmentCitationGroup id="F3F62B38FFFB6140FF02674AFC1FFAD9" box="[151,917,1302,1324]" pageId="42" pageNumber="43">
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<taxonomicName id="14E67795FFFB6140FF02674AFDD3FAD9" authority="Paulian, 1992: 1971" authorityName="Paulian" authorityPageNumber="1971" authorityYear="1992" box="[151,601,1302,1324]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="falcatus">
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<emphasis id="E192D004FFFB6140FF02674AFE0FFAD9" box="[151,389,1302,1324]" italics="true" pageId="42" pageNumber="43">Madecorphnus falcatus</emphasis>
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<treatmentCitation id="52472A07FFFB6140FE19674AFDD3FAD9" author="Paulian" box="[396,601,1302,1324]" page="1971" pageId="42" pageNumber="43" year="1992">
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<bibRefCitation id="B77771E7FFFB6140FE19674AFD90FAD9" author="Paulian" box="[396,538,1302,1324]" pageId="42" pageNumber="43" refString="Paulian, R. (1992) Un nouveau genre et une nouvelle espece d'Orphnidae de Madagascar (Coleoptera, Scarabaeoidea). Revue Francaise d'Entomologie (N. S.), 14, 169 - 171." type="journal article" year="1992">Paulian, 1992</bibRefCitation>
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: 1971
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="52472A07FFFB6140FDF1674AFCA7FAD9" author="Frolov" box="[612,813,1302,1324]" page="1107" pageId="42" pageNumber="43" year="2010">
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<bibRefCitation id="B77771E7FFFB6140FDF1674AFD65FAD9" author="Frolov" box="[612,751,1302,1324]" pageId="42" pageNumber="43" refString="Frolov, A. V. (2010 c) Revision of the Madagascan genus Madecorphnus Paulian (Coleoptera, Scarabaeidae, Orphninae). Journal of Natural History, 44, 1095 - 1111." type="journal article" year="2010" yearSuffix="c">Frolov 2010c</bibRefCitation>
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: 1107
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</treatmentCitation>
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, 2013a: 3.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="9BFC5F9DFFFB6140FF026705FB69FA63" pageId="42" pageNumber="43" type="materials_examined">
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<paragraph id="D3590C16FFFB6140FF026705FB69FA63" blockId="42.[151,1437,1369,2006]" pageId="42" pageNumber="43">
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<materialsCitation id="638E064BFFFB6140FF026705FEAEFA63" ID-GBIF-Occurrence="1416610303" collectionCode="ANTANAMBE, PROG" country="Madagascar" location="MADAGASCAR" pageId="42" pageNumber="43" specimenCode="ANTANAMBE 25" specimenCount="90" typeStatus="holotype">
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||
<emphasis id="E192D004FFFB6140FF026705FE43FA87" bold="true" box="[151,457,1369,1394]" pageId="42" pageNumber="43">
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<typeStatus id="0C5DB2B4FFFB6140FF026705FF58FA87" box="[151,210,1369,1394]" pageId="42" pageNumber="43">Type</typeStatus>
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material examined.
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</emphasis>
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<typeStatus id="0C5DB2B4FFFB6140FE406705FDB7FA87" box="[469,573,1369,1394]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
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(Figs. 21A–E), male, “
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<collectingCountry id="ABF14C86FFFB6140FCC06706FB95FA87" box="[853,1055,1370,1394]" name="Madagascar" pageId="42" pageNumber="43">MADAGASCAR</collectingCountry>
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<specimenCode id="8340A46DFFFB6140FBBF6706FA88FA87" box="[1066,1282,1369,1394]" collectionCode="ANTANAMBE" pageId="42" pageNumber="43">
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<collectionCode id="B5F794D3FFFB6140FBBF6706FB51FA87" box="[1066,1243,1370,1394]" pageId="42" pageNumber="43">ANTANAMBE</collectionCode>
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<date id="A7582AD6FFFB6140FB726705FA88FA87" box="[1255,1282,1369,1394]" pageId="42" pageNumber="43" value="1990-01-25">25</date>
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</specimenCode>
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/01/
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<specimenCount id="C5E0C79FFFFB6140FABB6705FAE7FA87" box="[1326,1389,1369,1394]" pageId="42" pageNumber="43" type="generic">90 P.</specimenCount>
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E.C. Foret
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<collectionCode id="B5F794D3FFFB6140FF4C6722FEAEFA63" box="[217,292,1406,1430]" pageId="42" pageNumber="43">PROG</collectionCode>
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</materialsCitation>
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. MAG188-007 /
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<taxonomicName id="14E67795FFFB6140FE766721FD45FA63" box="[483,719,1405,1430]" class="Insecta" family="Scarabaeidae" genus="Sissantobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov.">
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<emphasis id="E192D004FFFB6140FE766721FD45FA63" box="[483,719,1405,1430]" italics="true" pageId="42" pageNumber="43">Sissantobius falcatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FAA16D7FFFFB6140FD406722FC80FA60" box="[725,778,1406,1429]" pageId="42" pageNumber="43" rank="species">n.sp.</taxonomicNameLabel>
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R. Paulian det. / HOLOTYPE” (MNHN).
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFFFB6140FF5267FDFF7DF9D3" pageId="42" pageNumber="43" type="diagnosis">
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<paragraph id="D3590C16FFFB6140FF5267FDFF7DF9D3" blockId="42.[151,1437,1369,2006]" pageId="42" pageNumber="43">
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<emphasis id="E192D004FFFB6140FF5267FDFECBFA4F" bold="true" box="[199,321,1441,1466]" pageId="42" pageNumber="43">Diagnosis.</emphasis>
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This is the largest known species of
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<taxonomicName id="14E67795FFFB6140FD7F67FDFC18FA4F" box="[746,914,1441,1466]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
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<emphasis id="E192D004FFFB6140FD7F67FDFC18FA4F" box="[746,914,1441,1466]" italics="true" pageId="42" pageNumber="43">Madecorphnus</emphasis>
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</taxonomicName>
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with the body length reaching
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<quantity id="141EA1F3FFFB6140FB6367FEFAB6FA4C" box="[1270,1340,1442,1465]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="42" pageNumber="43" unit="mm" value="7.0">7 mm</quantity>
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. Also, it differs from other
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<taxonomicName id="14E67795FFFB6140FEF16799FD86FA2B" box="[356,524,1477,1502]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
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<emphasis id="E192D004FFFB6140FEF16799FD86FA2B" box="[356,524,1477,1502]" italics="true" pageId="42" pageNumber="43">Madecorphnus</emphasis>
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</taxonomicName>
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species in having a distinctive shape of the parameres (Fig. 21B) and internal sac armature consisting of 2 small, bifurcate sclerites, 1 long spur-like sclerite, and numerous small spinules (Figs. 21D–E).
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFFFB6146FF52646DFE28FEE8" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43" type="description">
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<paragraph id="D3590C16FFFB6140FF52646DFB91F9BF" blockId="42.[151,1437,1369,2006]" box="[199,1051,1585,1610]" pageId="42" pageNumber="43">
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<emphasis id="E192D004FFFB6140FF52646DFE29F9BF" bold="true" box="[199,419,1585,1610]" pageId="42" pageNumber="43">Description. Male.</emphasis>
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Body length
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<quantity id="141EA1F3FFFB6140FDAE646DFD4EF9BC" box="[571,708,1585,1610]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.1" metricValueMin="6.9" pageId="42" pageNumber="43" unit="mm" value="7.0" valueMax="7.1" valueMin="6.9">6.9–7.1 mm</quantity>
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. Color dark brown (Fig. 21A).
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</paragraph>
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<paragraph id="D3590C16FFFB6140FF526409FE94F967" blockId="42.[151,1437,1369,2006]" pageId="42" pageNumber="43">Right mandible 2 times longer than left, without tooth behind apex. Labrum trapezoidal, feebly visible in dorsal view.</paragraph>
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<paragraph id="D3590C16FFFB6140FF5264C1FC44F92F" blockId="42.[151,1437,1369,2006]" pageId="42" pageNumber="43">Frontoclypeus asymmetrical, slightly convex anteriorly, obtuse, with 2 long and a number of smaller setae. Frontoclypeus slightly depressed in the middle anteriorly, almost smooth.</paragraph>
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<paragraph id="D3590C16FFFB6140FF5264BAFCD9F8D4" blockId="42.[151,1437,1369,2006]" pageId="42" pageNumber="43">Pronotum 1.6 times wider than long, widest medially. Margins with relatively wide border, lateral margins with 4 long setae. Pronotum evenly convex, with minute punctures.</paragraph>
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<paragraph id="D3590C16FFFB6140FF526571FD89F847" blockId="42.[151,1437,1369,2006]" pageId="42" pageNumber="43">Elytra convex, with distinct humeral and apical tubercules. Maximum width approximately at basal 1/3. First stria distinct and reaching the apex of elytron, other striae indistinct. Epipleura with long, sparse, brown setae. Base of elytra with border from scutellum to humeral humps. Elytra sparsely punctate with relatively large punctures and minute punctures between them.</paragraph>
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<paragraph id="D3590C16FFFB6141FF5265E1FD2CFF0D" blockId="42.[151,1437,1369,2006]" lastBlockId="43.[151,1437,151,248]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex with robust, spur-like seta and a few smaller setae basally. Mesothoracic and metathoracic legs similar in shape. Longer tibial spur as long as 2 basal tarsomeres in mesothoracic legs and shorter than 3 basal tarsomeres in metathoracic legs. Metafemora with a few pectinate setae on the inner side apically.</paragraph>
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<paragraph id="D3590C16FFFA6141FF0265CCFE11F816" pageId="43" pageNumber="44">
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<table id="A1E6FEB6FFFA9E95FF0265CCFA16F816" box="[151,1436,1936,2019]" gridcols="3" gridrows="3" pageId="43" pageNumber="44">
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||
<tr id="6DD60E54FFFA9E95FF0265CCFA16F853" box="[151,1436,1936,1958]" gridrow="0" pageId="43" pageNumber="44">
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||
<th id="2E076728FFFA9E95FF0265CCFD09F853" box="[151,643,1936,1958]" gridcol="0" gridrow="0" pageId="43" pageNumber="44">
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<emphasis id="E192D004FFFA6141FF0265CCFEAAF853" bold="true" box="[151,288,1936,1958]" pageId="43" pageNumber="44">FIGURE 21.</emphasis>
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<taxonomicName id="14E67795FFFA6141FEB265CCFD9CF853" box="[295,534,1936,1958]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="falcatus">
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<emphasis id="E192D004FFFA6141FEB265CCFD9CF853" box="[295,534,1936,1958]" italics="true" pageId="43" pageNumber="44">Madecorphnus falcatus</emphasis>
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</taxonomicName>
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, holotype.
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</th>
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<th id="2E076728FFFA9E95FD1E65CCFBE2F853" box="[651,1128,1936,1958]" gridcol="1" gridrow="0" pageId="43" pageNumber="44">Habitus in dorsal view (A), aedeagus in lateral</th>
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<th id="2E076728FFFA9E95FBFA65CCFA16F853" box="[1135,1436,1936,1958]" gridcol="2" gridrow="0" pageId="43" pageNumber="44">view and parameres in dorsal</th>
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||
</tr>
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<tr id="6DD60E54FFFA9E95FF0265F2FA16F831" box="[151,1436,1966,1988]" gridrow="1" pageId="43" pageNumber="44">
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<th id="2E076728FFFA9E95FF0265F2FD09F831" box="[151,643,1966,1988]" gridcol="0" gridrow="1" pageId="43" pageNumber="44">and apical view (B), labels (C), internal sac</th>
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<td id="2E076728FFFA9E95FD1E65F2FBE2F831" box="[651,1128,1966,1988]" gridcol="1" gridrow="1" pageId="43" pageNumber="44">of aedeagus (D—photograph, E—schematic</td>
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<td id="2E076728FFFA9E95FBFA65F2FA16F831" box="[1135,1436,1966,1988]" gridcol="2" gridrow="1" pageId="43" pageNumber="44">representation of armature),</td>
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</tr>
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<tr id="6DD60E54FFFA9E95FF026591FA16F816" box="[151,1436,1997,2019]" gridrow="2" pageId="43" pageNumber="44" rowspan-1="1" rowspan-2="1">
|
||
<th id="2E076728FFFA9E95FF026591FD09F816" box="[151,643,1997,2019]" gridcol="0" gridrow="2" pageId="43" pageNumber="44">distributional records (F).</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="D3590C16FFFD6146FF5262C4FD8CFF0D" blockId="44.[151,1437,151,356]" pageId="44" pageNumber="45">Parameres relatively long, their apices curved downwards (in lateral view), with small teeth laterally visible in apical view (Fig. 21B). Internal sac with 1 long, spur-shaped sclerite, 2 smaller, double sclerites, and a field small adjacent spinules (Figs. 21D–E).</paragraph>
|
||
<paragraph id="D3590C16FFFD6146FF526358FE28FEE8" blockId="44.[151,1437,151,356]" box="[199,418,260,285]" pageId="44" pageNumber="45">
|
||
<emphasis id="E192D004FFFD6146FF526358FEAFFEE8" bold="true" box="[199,293,260,285]" pageId="44" pageNumber="45">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BFC5F9DFFFD6146FF52637BFF59FE91" pageId="44" pageNumber="45" type="distribution">
|
||
<paragraph id="D3590C16FFFD6146FF52637BFF59FE91" blockId="44.[151,1437,151,356]" pageId="44" pageNumber="45">
|
||
<emphasis id="E192D004FFFD6146FF52637BFE7FFEB5" bold="true" box="[199,501,295,320]" pageId="44" pageNumber="45">Distribution and habitat.</emphasis>
|
||
<taxonomicName id="14E67795FFFD6146FE68637BFC8DFEB5" box="[509,775,295,320]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="falcatus">
|
||
<emphasis id="E192D004FFFD6146FE68637BFC8DFEB5" box="[509,775,295,320]" italics="true" pageId="44" pageNumber="45">Madecorphnus falcatus</emphasis>
|
||
</taxonomicName>
|
||
is known from two localities in eastern
|
||
<collectingCountry id="ABF14C86FFFD6146FB43637BFAD7FEB5" box="[1238,1373,295,320]" name="Madagascar" pageId="44" pageNumber="45">Madagascar</collectingCountry>
|
||
(Fig. 21F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |