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<document id="A1DB88C3DAE5D066586C38E940D09178" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier &amp; Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FFDF617AFF026640FCD6FD38" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pseudorphnus olsoufieffi Paulian 1977" docType="treatment" docVersion="16" lastPageNumber="17" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="15" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:title id="7F650D7080EBE0629B17C928103B1CAC">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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<mods:namePart id="7F32D1C8C7FDE609C6CDCEE949A35C72">Frolov, Andrey V.</mods:namePart>
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<treatment id="5B4FBD00FFDF617AFF026640FCD6FD38" ID-DOI="http://doi.org/10.5281/zenodo.6057634" ID-GBIF-Taxon="125107268" ID-Zenodo-Dep="6057634" LSID="urn:lsid:plazi:treatment:5B4FBD00FFDF617AFF026640FCD6FD38" httpUri="http://treatment.plazi.org/id/5B4FBD00FFDF617AFF026640FCD6FD38" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
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<paragraph id="D3590C16FFDF6164FF026640FDF2FBC3" blockId="14.[151,632,1052,1111]" box="[151,632,1052,1078]" pageId="14" pageNumber="15">
<heading id="8811BB7AFFDF6164FF026640FDF2FBC3" bold="true" box="[151,632,1052,1078]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="14E67795FFDF6164FF026640FDF2FBC3" authority="Paulian, 1977" authorityName="Paulian" authorityYear="1977" box="[151,632,1052,1078]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="E192D004FFDF6164FF026640FDF2FBC3" bold="true" box="[151,632,1052,1078]" pageId="14" pageNumber="15">
<emphasis id="E192D004FFDF6164FF026640FE4DFBC3" bold="true" box="[151,455,1052,1078]" italics="true" pageId="14" pageNumber="15">Pseudorphnus olsoufieffi</emphasis>
<bibRefCitation id="B77771E7FFDF6164FE5A6640FDF2FBC3" author="Paulian" box="[463,632,1052,1078]" pageId="14" pageNumber="15" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D3590C16FFDF6164FF026662FEA6FBA2" blockId="14.[151,632,1052,1111]" box="[151,300,1086,1111]" pageId="14" pageNumber="15">
(
<figureCitation id="4BDD1093FFDF6164FF0A6662FF61FBA2" box="[159,235,1086,1111]" captionStart="FIGURE 6" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[216,1403,212,1792]" captionTargetId="figure@15.[181,1405,193,1793]" captionTargetPageId="15" captionText="FIGURE 6. Pseudorphnus olsoufieffi. Habitus (A—dorsal, B—lateral view), aedeagus in lateral view and parameres in dorsal view (C), head in dorsal view (D), head and pronotum in dorsolateral view (E), labels of holotype (F), distributional records (G)." httpUri="https://zenodo.org/record/200450/files/figure.png" pageId="14" pageNumber="15">Figs. 6</figureCitation>
AG)
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFDF6164FF0266DAFBDBFB69" box="[151,1105,1158,1180]" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="D3590C16FFDF6164FF0266DAFBDBFB69" blockId="14.[151,1105,1158,1180]" box="[151,1105,1158,1180]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="F3F62B38FFDF6164FF0266DAFBDBFB69" box="[151,1105,1158,1180]" pageId="14" pageNumber="15">
<taxonomicName id="14E67795FFDF6164FF0266DAFDE2FB69" authority="Paulian, 1977: 1205" authorityName="Paulian" authorityPageNumber="1205" authorityYear="1977" box="[151,616,1158,1180]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="E192D004FFDF6164FF0266DAFE19FB69" box="[151,403,1158,1180]" italics="true" pageId="14" pageNumber="15">Pseudorphnus olsoufieffi</emphasis>
<treatmentCitation id="52472A07FFDF6164FE0F66DAFDE2FB69" author="Paulian" box="[410,616,1158,1180]" page="1205" pageId="14" pageNumber="15" year="1977">
<bibRefCitation id="B77771E7FFDF6164FE0F66DAFDA2FB69" author="Paulian" box="[410,552,1158,1180]" pageId="14" pageNumber="15" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
: 1205
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="52472A07FFDF6164FDE766DAFC9DFB69" author="Frolov" box="[626,791,1158,1180]" page="66" pageId="14" pageNumber="15" year="2011">
<bibRefCitation id="B77771E7FFDF6164FDE766DAFD7BFB69" author="Frolov" box="[626,753,1158,1180]" pageId="14" pageNumber="15" refString="Frolov, A. V. (2011) A new species of the Madagascan genus Pseudorphnus (Coleoptera: Scarabaeidae: Orphninae). Zootaxa, 2836, 65 - 68." type="journal article" year="2011">Frolov 2011</bibRefCitation>
: 66
</treatmentCitation>
;
<treatmentCitation id="52472A07FFDF6164FCB766DAFBC7FB69" author="Frolov" box="[802,1101,1158,1180]" page="30" pageId="14" pageNumber="15" year="2006">
<bibRefCitation id="B77771E7FFDF6164FCB766DAFBADFB69" author="Frolov" box="[802,1063,1158,1180]" pageId="14" pageNumber="15" refString="Frolov, A. V. &amp; Montreuil, O. (2006) Description of the male of the rare Madagascan species Pseudorphnus hiboni with notes on the genus Pseudorphnus (Coleoptera: Scarabaeidae: Orphninae). Zootaxa, 1154, 27 - 33." type="journal article" year="2006">Frolov &amp; Montreuil 2006</bibRefCitation>
: 30
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFDF6164FF026695FAD3FA87" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="D3590C16FFDF6164FF026695FE27FADC" blockId="14.[151,1437,1225,2006]" pageId="14" pageNumber="15">
<materialsCitation id="638E064BFFDF6164FF026695FE2CFADF" ID-GBIF-Occurrence="1416610177" collectionCode="MNHN" determinerName="R. Paulian" location="Alaotra Lake" pageId="14" pageNumber="15" specimenCount="3" stateProvince="Toamasina Province" typeStatus="holotype">
<emphasis id="E192D004FFDF6164FF026695FE40FB17" bold="true" box="[151,458,1225,1250]" pageId="14" pageNumber="15">
<typeStatus id="0C5DB2B4FFDF6164FF026695FF58FB17" box="[151,210,1225,1250]" pageId="14" pageNumber="15">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="0C5DB2B4FFDF6164FE426695FDB5FB17" box="[471,575,1225,1250]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
(
<figureCitation id="4BDD1093FFDF6164FDC16695FD2DFB17" box="[596,679,1225,1250]" captionStart="FIGURE 6" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[216,1403,212,1792]" captionTargetId="figure@15.[181,1405,193,1793]" captionTargetPageId="15" captionText="FIGURE 6. Pseudorphnus olsoufieffi. Habitus (A—dorsal, B—lateral view), aedeagus in lateral view and parameres in dorsal view (C), head in dorsal view (D), head and pronotum in dorsolateral view (E), labels of holotype (F), distributional records (G)." httpUri="https://zenodo.org/record/200450/files/figure.png" pageId="14" pageNumber="15">Figs. 6</figureCitation>
AF), male, “Madacascar Antsianaka et lac Alaotra [
<collectingRegion id="1122C2F4FFDF6164FAB56695FF77FAF3" country="Madagascar" name="Toamasina" pageId="14" pageNumber="15">Toamasina Province</collectingRegion>
,
<location id="D6395ACDFFDF6164FE9A66B1FE26FAF3" LSID="urn:lsid:plazi:treatment:5B4FBD00FFDF617AFF026640FCD6FD38:D6395ACDFFDF6164FE9A66B1FE26FAF3" box="[271,428,1261,1286]" name="Alaotra Lake" pageId="14" pageNumber="15" stateProvince="Toamasina Province">Alaotra Lake</location>
] 2e Trimestre 1889 Perrot Freres /
<taxonomicName id="14E67795FFDF6164FCF166B1FB09FAF3" box="[868,1155,1261,1286]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="olsouffiefi" status="sp. nov.">
<emphasis id="E192D004FFDF6164FCF166B1FB09FAF3" box="[868,1155,1261,1286]" italics="true" pageId="14" pageNumber="15">Pseudorphnus olsouffiefi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAA16D7FFFDF6164FB0466B2FB5EFAF0" box="[1169,1236,1262,1285]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
<determinerName id="2685E0D2FFDF6164FB7466B1FAC5FAF3" box="[1249,1359,1261,1286]" pageId="14" pageNumber="15">R.Paulian</determinerName>
det. /
<typeStatus id="0C5DB2B4FFDF6164FF02674EFEA4FADF" box="[151,302,1298,1322]" pageId="14" pageNumber="15" type="holotype">HOLOTYPE</typeStatus>
” (
<collectionCode id="B5F794D3FFDF6164FED0674EFE14FADF" box="[325,414,1298,1322]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="D3590C16FFDF6164FF526769FAD3FA87" blockId="14.[151,1437,1225,2006]" pageId="14" pageNumber="15">
<materialsCitation id="638E064BFFDF6164FF526769FADFFA87" ID-GBIF-Occurrence="1416610213" collectionCode="EST, ANKASOKA, ROUTE, LAKATO, MNHN" country="Madagascar" location="Toamasina" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Toamasina">
<emphasis id="E192D004FFDF6164FF526769FC5BFABB" bold="true" box="[199,977,1333,1358]" pageId="14" pageNumber="15">
Additional material examined.
<collectingCountry id="ABF14C86FFDF6164FDC86769FCA5FABB" box="[605,815,1333,1358]" name="Madagascar" pageId="14" pageNumber="15">MADAGASCAR</collectingCountry>
:
<collectingRegion id="1122C2F4FFDF6164FCD36769FC42FABB" box="[838,968,1333,1358]" country="Madagascar" name="Toamasina" pageId="14" pageNumber="15">Toamasina</collectingRegion>
:
</emphasis>
one female, “
<collectingCountry id="ABF14C86FFDF6164FB1B676AFAD2FABB" box="[1166,1368,1334,1358]" name="Madagascar" pageId="14" pageNumber="15">MADAGASCAR</collectingCountry>
<collectionCode id="B5F794D3FFDF6164FAFF676AFE12FA87" pageId="14" pageNumber="15">EST ANKASOKA ROUTE</collectionCode>
DE
<collectionCode id="B5F794D3FFDF6164FE596706FDB1FA87" box="[460,571,1370,1394]" pageId="14" pageNumber="15">LAKATO</collectionCode>
[Ankasoka, road to Lakato] 1-[...]-1972 R.VIOSSAT REC.” (
<collectionCode id="B5F794D3FFDF6164FB646706FAC7FA87" box="[1265,1357,1370,1394]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="15">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFDF6164FF526721FE26F9BF" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="D3590C16FFDF6164FF526721FE26F9BF" blockId="14.[151,1437,1225,2006]" pageId="14" pageNumber="15">
<emphasis id="E192D004FFDF6164FF526721FECBFA63" bold="true" box="[199,321,1405,1430]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
<taxonomicName id="14E67795FFDF6164FEDC6721FDEBFA63" box="[329,609,1405,1430]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="E192D004FFDF6164FEDC6721FDEBFA63" box="[329,609,1405,1430]" italics="true" pageId="14" pageNumber="15">Pseudorphnus olsoufieffi</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="14E67795FFDF6164FD626722FCF3FA60" box="[759,889,1406,1429]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="E192D004FFDF6164FD626722FCF3FA60" box="[759,889,1406,1429]" italics="true" pageId="14" pageNumber="15">P. carinatus</emphasis>
</taxonomicName>
in the body size, sculpture of the pronotum and elytra, and in the shape of the prothoracic ridges, but the males of the species differ in having a rather large, conical tubercle situated in the center of the frontoclypeus, as opposed to having a long transverse low ridge near the anterior margin in the later species. The two species also differ in the sculpture of the frontoclypeus (rugose in
<taxonomicName id="14E67795FFDF6164FA1267B6FE8DF9D3" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="E192D004FFDF6164FA1267B6FE8DF9D3" italics="true" pageId="14" pageNumber="15">P. olsoufieffi</emphasis>
</taxonomicName>
and smooth in
<taxonomicName id="14E67795FFDF6164FE2F6452FDB7F9D0" box="[442,573,1550,1573]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="E192D004FFDF6164FE2F6452FDB7F9D0" box="[442,573,1550,1573]" italics="true" pageId="14" pageNumber="15">P. carinatus</emphasis>
</taxonomicName>
) and in the shape of the parameres (having less acute apices not depressed laterally in
<taxonomicName id="14E67795FFDF6164FE83646EFE16F9BC" box="[278,412,1586,1609]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="E192D004FFDF6164FE83646EFE16F9BC" box="[278,412,1586,1609]" italics="true" pageId="14" pageNumber="15">P. carinatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFDF617AFF526409FB89FD70" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
<paragraph id="D3590C16FFDF6164FF526409FAEAF998" blockId="14.[151,1437,1225,2006]" box="[199,1376,1621,1646]" pageId="14" pageNumber="15">
<emphasis id="E192D004FFDF6164FF526409FE29F99B" bold="true" box="[199,419,1621,1646]" pageId="14" pageNumber="15">Description. Male.</emphasis>
Elongate, shiny (
<figureCitation id="4BDD1093FFDF6164FDF26409FD3FF99B" box="[615,693,1621,1646]" captionStart="FIGURE 6" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[216,1403,212,1792]" captionTargetId="figure@15.[181,1405,193,1793]" captionTargetPageId="15" captionText="FIGURE 6. Pseudorphnus olsoufieffi. Habitus (A—dorsal, B—lateral view), aedeagus in lateral view and parameres in dorsal view (C), head in dorsal view (D), head and pronotum in dorsolateral view (E), labels of holotype (F), distributional records (G)." httpUri="https://zenodo.org/record/200450/files/figure.png" pageId="14" pageNumber="15">Figs. 6</figureCitation>
A, B), colored uniformly brown beetle. Body length
<quantity id="141EA1F3FFDF6164FA966409FAD6F998" box="[1283,1372,1621,1646]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="14" pageNumber="15" unit="mm" value="9.5">9.5 mm</quantity>
.
</paragraph>
<paragraph id="D3590C16FFDF6164FF526425FBAFF8D7" blockId="14.[151,1437,1225,2006]" pageId="14" pageNumber="15">
Frontoclypeus slightly convex anteriorly, obtuse laterally, anterior margin setose and crenulate in dorsal view. Eyes relatively large (diameter larger than the distance between eye and gula in ventral view), incompletely divided by canthus into slightly smaller dorsal and larger ventral parts. Frontoclypeus with a short conical tubercle in center, rugosely punctate (
<figureCitation id="4BDD1093FFDF6164FE5864B9FD95F90B" box="[461,543,1765,1790]" captionStart="FIGURE 6" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[216,1403,212,1792]" captionTargetId="figure@15.[181,1405,193,1793]" captionTargetPageId="15" captionText="FIGURE 6. Pseudorphnus olsoufieffi. Habitus (A—dorsal, B—lateral view), aedeagus in lateral view and parameres in dorsal view (C), head in dorsal view (D), head and pronotum in dorsolateral view (E), labels of holotype (F), distributional records (G)." httpUri="https://zenodo.org/record/200450/files/figure.png" pageId="14" pageNumber="15">Figs. 6</figureCitation>
D, E). Labrum bilobate, slightly sinuate in the middle and relatively feebly protruding past frontoclypeus. Length in the middle is 1/8 width (in dorsal view).
</paragraph>
<paragraph id="D3590C16FFDF6164FF526572FEEFF820" blockId="14.[151,1437,1225,2006]" pageId="14" pageNumber="15">
Pronotum 1.6 times wider than long, widest medially. Anterior margin with wide border, base with fine border. Lateral margins densely punctate, appearing crenulate in dorsal view. Disc of pronotum with deep excavation in the middle, with 2 slender longitudinal somewhat triangular ridges bordering the excavation (
<figureCitation id="4BDD1093FFDF6164FB1B6529FB57F87B" box="[1166,1245,1909,1934]" captionStart="FIGURE 6" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[216,1403,212,1792]" captionTargetId="figure@15.[181,1405,193,1793]" captionTargetPageId="15" captionText="FIGURE 6. Pseudorphnus olsoufieffi. Habitus (A—dorsal, B—lateral view), aedeagus in lateral view and parameres in dorsal view (C), head in dorsal view (D), head and pronotum in dorsolateral view (E), labels of holotype (F), distributional records (G)." httpUri="https://zenodo.org/record/200450/files/figure.png" pageId="14" pageNumber="15">Figs. 6</figureCitation>
B, E). Surface of disc between the ridges smooth, without punctures. Sides of pronotum rugose posteriorly. Lateral margins with long, brown setae.
</paragraph>
<caption id="87995C9EFFDE6165FF02654AFF4BF89F" httpUri="https://zenodo.org/record/200450/files/figure.png" pageId="15" pageNumber="16" targetBox="[216,1403,212,1792]" targetPageId="15">
<paragraph id="D3590C16FFDE6165FF02654AFF4BF89F" blockId="15.[151,1436,1814,1898]" pageId="15" pageNumber="16">
<emphasis id="E192D004FFDE6165FF02654AFE99F8D9" bold="true" box="[151,275,1814,1836]" pageId="15" pageNumber="16">FIGURE 6.</emphasis>
<emphasis id="E192D004FFDE6165FE8F654AFD94F8D9" box="[282,542,1814,1836]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="14E67795FFDE6165FE8F654AFD90F8D9" box="[282,538,1814,1836]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="olsoufieffi">Pseudorphnus olsoufieffi</taxonomicName>
.
</emphasis>
Habitus (A—dorsal, B—lateral view), aedeagus in lateral view and parameres in dorsal view (C), head in dorsal view (D), head and pronotum in dorsolateral view (E), labels of holotype (F), distributional records (G).
</paragraph>
</caption>
<paragraph id="D3590C16FFDE6165FF5265CDFB94F85F" blockId="15.[151,1436,1937,2034]" box="[199,1054,1937,1962]" pageId="15" pageNumber="16">Scutellum triangular, narrowly rounded apically, about 1/10 length of elytra.</paragraph>
<paragraph id="D3590C16FFDE617AFF5265E9FB9BFF20" blockId="15.[151,1436,1937,2034]" lastBlockId="16.[151,1436,151,717]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">Elytra convex, with marked humeral humps. Maximum width approximately at the middle. Elytral striae are feebly distinct. Elytra covered with sparse narrow and somewhat curved punctures, bearing a short, yellow seta; the setae are mostly abraded on disc. Epipleura with long, sparse, brown setae. Base of elytra bordered and densely punctate with punctures similar to those in striae but more rugose and irregular.</paragraph>
<paragraph id="D3590C16FFC1617AFF526283FE21FEE9" blockId="16.[151,1436,151,717]" pageId="16" pageNumber="17">Lateral margin of protibia basad of outer teeth not crenulated. Apices with 3 robust, spur-like setae and a number of smaller setae.</paragraph>
<paragraph id="D3590C16FFC1617AFF52637BFCE4FE59" blockId="16.[151,1436,151,717]" pageId="16" pageNumber="17">Mesothoracic and metathoracic legs are similar in shape; metafemora and metatibiae about 1/8 longer than mesofemora and mesotibiae. Tibiae somewhat triangular, with 2 apical spurs, with inner margin only slightly concave and with 1 transverse keel. Longer tibial spur as long as 2 basal tarsomeres. Claws 1/3 length of apical tarsomere. Femora almost impunctate, with 2 rows of long setae.</paragraph>
<paragraph id="D3590C16FFC1617AFF5263EBFDB0FE25" blockId="16.[151,1436,151,717]" box="[199,570,439,464]" pageId="16" pageNumber="17">The holotype lacks the abdomen.</paragraph>
<paragraph id="D3590C16FFC1617AFF526380FB56FE01" blockId="16.[151,1436,151,717]" box="[199,1244,476,501]" pageId="16" pageNumber="17">
Aedeagus with parameres having short, acute, somewhat depressed laterally apices (
<figureCitation id="4BDD1093FFC1617AFBE26380FB30FE00" box="[1143,1210,476,501]" captionStart="FIGURE 6" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[216,1403,212,1792]" captionTargetId="figure@15.[181,1405,193,1793]" captionTargetPageId="15" captionText="FIGURE 6. Pseudorphnus olsoufieffi. Habitus (A—dorsal, B—lateral view), aedeagus in lateral view and parameres in dorsal view (C), head in dorsal view (D), head and pronotum in dorsolateral view (E), labels of holotype (F), distributional records (G)." httpUri="https://zenodo.org/record/200450/files/figure.png" pageId="16" pageNumber="17">Fig. 6</figureCitation>
C).
</paragraph>
<paragraph id="D3590C16FFC1617AFF5263A3FB89FD70" blockId="16.[151,1436,151,717]" pageId="16" pageNumber="17">
<emphasis id="E192D004FFC1617AFF5263A3FEAFFDED" bold="true" box="[199,293,511,536]" pageId="16" pageNumber="17">Female.</emphasis>
The only female specimen available, putatively identified as
<taxonomicName id="14E67795FFC1617AFC46605DFBD4FDED" box="[979,1118,511,536]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="E192D004FFC1617AFC46605DFBD4FDED" box="[979,1118,511,536]" italics="true" pageId="16" pageNumber="17">P. olsoufieffi</emphasis>
</taxonomicName>
, has the body length of
<quantity id="141EA1F3FFC1617AFAFE63A3FF4CFDC9" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="16" pageNumber="17" unit="mm" value="10.5">10.5 mm</quantity>
. It differs from the male in the rugose surface of the head, absence of the frontoclypeal horn or tubercle, presence of a distinct spur on protibiae, densely punctate disc of the pronotum, and absence of the pronotal excavations and ridges. The sexes are similar in general body shape and color.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFC1617AFF5260D3FCD6FD38" pageId="16" pageNumber="17" type="distribution">
<paragraph id="D3590C16FFC1617AFF5260D3FCD6FD38" blockId="16.[151,1436,151,717]" pageId="16" pageNumber="17">
<emphasis id="E192D004FFC1617AFF5260D3FEEAFD5D" bold="true" box="[199,352,655,680]" pageId="16" pageNumber="17">Distribution.</emphasis>
This species is known from two localities in the East
<collectingCountry id="ABF14C86FFC1617AFC4960D3FBE9FD5D" box="[988,1123,655,680]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
(
<figureCitation id="4BDD1093FFC1617AFBE360D3FB36FD5D" box="[1142,1212,655,680]" captionStart="FIGURE 6" captionStartId="15.[151,250,1814,1836]" captionTargetBox="[216,1403,212,1792]" captionTargetId="figure@15.[181,1405,193,1793]" captionTargetPageId="15" captionText="FIGURE 6. Pseudorphnus olsoufieffi. Habitus (A—dorsal, B—lateral view), aedeagus in lateral view and parameres in dorsal view (C), head in dorsal view (D), head and pronotum in dorsolateral view (E), labels of holotype (F), distributional records (G)." httpUri="https://zenodo.org/record/200450/files/figure.png" pageId="16" pageNumber="17">Fig. 6</figureCitation>
G). Exact habitat is unknown but the localities are situated in the rain forest region.
</paragraph>
</subSubSection>
</treatment>
</document>