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<document id="234556CA4459A7D1450FFA811FDAAAEB" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier &amp; Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FFC16178FF02617CFEFEFADA" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pseudorphnus hiboni Paulian 1959" docType="treatment" docVersion="15" lastPageNumber="19" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="17" updateTime="1698433759527" updateUser="ExternalLinkService">
<mods:mods id="B72998BEB8407023DC9E7DBABD72D077" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2E1B9533C2C2062414230C88F9A8AC85">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
</mods:titleInfo>
<mods:name id="A2DEB1C53C4AF93C57C5D71DBC42AD0E" type="personal">
<mods:role id="42803EF94D793DDBCA63636E7B2F5341">
<mods:roleTerm id="02A9AD3582AA960A253FF7566C45E5E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09C18F1E6F4357762F611339A5115D98">Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:name id="E098FFF3F3F8FBF2474F64EA516BF527" type="personal">
<mods:role id="EBAEB169B80D4456073CEB0BBECA5DF6">
<mods:roleTerm id="8DE225CBDBA7487C8E8F1052607C8705">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1A73AA6CE75B9089C3C9495D93197A34">Montreuil, Olivier</mods:namePart>
</mods:name>
<mods:name id="C8845DB28391FCD77225AAF68FDBD0D1" type="personal">
<mods:role id="2E92EB4C906ED3107E770204B2C0A5A1">
<mods:roleTerm id="689CB04BCD4FC0688F2B317EF08DB7E8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EFDAA8415D1401FA6B6EFE1C1C21A818">Akhmetova, Lilia A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5747DB991ABE2DB305D2FAA164ADD43B">text</mods:typeOfResource>
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<mods:titleInfo id="3802D8ED969EE692AD4F4F6314415B68">
<mods:title id="FEAE786D286A34023E4D0CDBE1B0E687">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6D8EF2022E606CCB5788D1C90171415F">
<mods:date id="C375D79A85AF6D2B7438CB029BD393FF">2016</mods:date>
<mods:detail id="635C9A50CE0C7BEA9D8837005D251BAA" type="volume">
<mods:number id="00ED1AAF2FCA31D4C39D4926FD13F543">4207</mods:number>
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<mods:detail id="447B8193AF55E17EEDB219F67D09943B" type="issue">
<mods:number id="4FBAB319C1CC55098FC644570B713B35">1</mods:number>
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<mods:start id="5709ED06B99989F8F427279826FCC085">1</mods:start>
<mods:end id="031CE0CACA39EDDDBA8644808D93B40D">93</mods:end>
</mods:extent>
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<mods:classification id="CCA8E4C76D361ED64465CD7D8FAE9858">journal article</mods:classification>
<mods:identifier id="545C9118E31390508B602091AED8BCCF" type="CLB-Dataset">37549</mods:identifier>
<mods:identifier id="EA1EE6F326B283A2CD9123C3582851FD" type="DOI">10.11646/zootaxa.4207.1.1</mods:identifier>
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<treatment id="5B4FBD00FFC16178FF02617CFEFEFADA" ID-DOI="http://doi.org/10.5281/zenodo.3510784" ID-GBIF-Taxon="125107293" ID-Zenodo-Dep="3510784" LSID="urn:lsid:plazi:treatment:5B4FBD00FFC16178FF02617CFEFEFADA" httpUri="http://treatment.plazi.org/id/5B4FBD00FFC16178FF02617CFEFEFADA" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="9BFC5F9DFFC1617AFF02617CFEA6FCAE" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="D3590C16FFC1617AFF02617CFDC6FCCF" blockId="16.[151,588,800,859]" box="[151,588,800,826]" pageId="16" pageNumber="17">
<heading id="8811BB7AFFC1617AFF02617CFDC6FCCF" bold="true" box="[151,588,800,826]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="14E67795FFC1617AFF02617CFDC6FCCF" authority="Paulian, 1959" authorityName="Paulian" authorityYear="1959" box="[151,588,800,826]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hiboni">
<emphasis id="E192D004FFC1617AFF02617CFDC6FCCF" bold="true" box="[151,588,800,826]" pageId="16" pageNumber="17">
<emphasis id="E192D004FFC1617AFF02617CFE10FCCF" bold="true" box="[151,410,800,826]" italics="true" pageId="16" pageNumber="17">Pseudorphnus hiboni</emphasis>
<bibRefCitation id="B77771E7FFC1617AFE37617CFDC6FCCF" author="Paulian" box="[418,588,800,826]" pageId="16" pageNumber="17" refString="Paulian, R. (1959) Une nouvelle espece d'Orphnitae malgache (Coleoptera, Scarabaeidae). Revue Francaise d'Entomologie (N. S.), 14, 134 - 135." type="journal article" year="1959">Paulian, 1959</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D3590C16FFC1617AFF02611EFEA6FCAE" blockId="16.[151,588,800,859]" box="[151,300,834,859]" pageId="16" pageNumber="17">
(
<figureCitation id="4BDD1093FFC1617AFF0A611EFF61FCAE" box="[159,235,834,859]" captionStart="FIGURE 7" captionStartId="17.[151,250,1807,1829]" captionTargetBox="[220,1364,197,1782]" captionTargetId="figure@17.[206,1380,193,1796]" captionTargetPageId="17" captionText="FIGURE 7. Pseudorphnus hiboni. Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200451/files/figure.png" pageId="16" pageNumber="17">Figs. 7</figureCitation>
A–H)
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFC1617AFF0261D6FBC9FC55" box="[151,1091,906,928]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="D3590C16FFC1617AFF0261D6FBC9FC55" blockId="16.[151,1091,906,928]" box="[151,1091,906,928]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="F3F62B38FFC1617AFF0261D6FBC9FC55" box="[151,1091,906,928]" pageId="16" pageNumber="17">
<taxonomicName id="14E67795FFC1617AFF0261D6FDBCFC55" authority="Paulian, 1959: 143" authorityName="Paulian" authorityPageNumber="143" authorityYear="1959" box="[151,566,906,928]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hiboni">
<emphasis id="E192D004FFC1617AFF0261D6FEE5FC55" box="[151,367,906,928]" italics="true" pageId="16" pageNumber="17">Pseudorphnus hiboni</emphasis>
<treatmentCitation id="52472A07FFC1617AFEE061D6FDBCFC55" author="Paulian" box="[373,566,906,928]" page="143" pageId="16" pageNumber="17" year="1959">
<bibRefCitation id="B77771E7FFC1617AFEE061D6FD8EFC55" author="Paulian" box="[373,516,906,928]" pageId="16" pageNumber="17" refString="Paulian, R. (1959) Une nouvelle espece d'Orphnitae malgache (Coleoptera, Scarabaeidae). Revue Francaise d'Entomologie (N. S.), 14, 134 - 135." type="journal article" year="1959">Paulian, 1959</bibRefCitation>
: 143
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="52472A07FFC1617AFDD461D6FC83FC55" author="Paulian" box="[577,777,906,928]" page="1207" pageId="16" pageNumber="17" year="1977">
<bibRefCitation id="B77771E7FFC1617AFDD461D6FD43FC55" author="Paulian" box="[577,713,906,928]" pageId="16" pageNumber="17" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian 1977</bibRefCitation>
: 1207
</treatmentCitation>
;
<treatmentCitation id="52472A07FFC1617AFC8661D6FBB5FC55" ID-DOI="http://doi.org/10.5281/zenodo.3501755" ID-Zenodo-Dep="3501755" author="Frolov" box="[787,1087,906,928]" httpUri="http://treatment.plazi.org/id/BF06506D0364B87E957B500FFC2AB91D" page="28" pageId="16" pageNumber="17" year="2006">
<bibRefCitation id="B77771E7FFC1617AFC8661D6FB92FC55" author="Frolov" box="[787,1048,906,928]" pageId="16" pageNumber="17" refString="Frolov, A. V. &amp; Montreuil, O. (2006) Description of the male of the rare Madagascan species Pseudorphnus hiboni with notes on the genus Pseudorphnus (Coleoptera: Scarabaeidae: Orphninae). Zootaxa, 1154, 27 - 33." type="journal article" year="2006">Frolov &amp; Montreuil 2006</bibRefCitation>
: 28
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFC1617AFF026191FC28FABB" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="D3590C16FFC1617AFF026191FCAFFBFF" blockId="16.[151,1437,973,2006]" pageId="16" pageNumber="17">
<materialsCitation id="638E064BFFC1617AFF026191FCAFFBFF" ID-GBIF-Occurrence="1416610146" collectionCode="MNHN" collectorName="Figs. &amp; Tolanaro &amp; P. Hybon" location="Fort Dauphin" municipality="Toliary Province" pageId="16" pageNumber="17" specimenCount="4" typeStatus="holotype">
<emphasis id="E192D004FFC1617AFF026191FE35FC13" bold="true" box="[151,447,973,998]" pageId="16" pageNumber="17">
<typeStatus id="0C5DB2B4FFC1617AFF026191FF58FC13" box="[151,210,973,998]" pageId="16" pageNumber="17">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="0C5DB2B4FFC1617AFE536191FDA4FC13" box="[454,558,973,998]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
(
<figureCitation id="4BDD1093FFC1617AFDAB6191FD06FC13" box="[574,652,973,998]" captionStart="FIGURE 7" captionStartId="17.[151,250,1807,1829]" captionTargetBox="[220,1364,197,1782]" captionTargetId="figure@17.[206,1380,193,1796]" captionTargetPageId="17" captionText="FIGURE 7. Pseudorphnus hiboni. Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200451/files/figure.png" pageId="16" pageNumber="17">
<collectorName id="7E1369C0FFC1617AFDAB6191FDFDFC13" box="[574,631,973,998]" pageId="16" pageNumber="17">Figs.</collectorName>
7
</figureCitation>
F–G), female, “
<location id="D6395ACDFFC1617AFCAF6192FC59FC13" LSID="urn:lsid:plazi:treatment:5B4FBD00FFC16178FF02617CFEFEFADA:D6395ACDFFC1617AFCAF6192FC59FC13" box="[826,979,973,998]" municipality="Toliary Province" name="Fort Dauphin" pageId="16" pageNumber="17">Fort Dauphin</location>
[
<collectingMunicipality id="333D966CFFC1617AFC756191FB28FC13" box="[992,1186,973,998]" pageId="16" pageNumber="17">Toliary Province</collectingMunicipality>
,
<collectorName id="7E1369C0FFC1617AFB396191FA9EFC13" box="[1196,1300,973,998]" pageId="16" pageNumber="17">Tôlanaro</collectorName>
] (
<collectorName id="7E1369C0FFC1617AFABF6192FA1EFC13" box="[1322,1428,973,998]" pageId="16" pageNumber="17">P. Hybon</collectorName>
) 1943 /
<taxonomicName id="14E67795FFC1617AFF7161ADFE59FBFF" box="[228,467,1009,1034]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hiboni" status="sp. nov.">
<emphasis id="E192D004FFC1617AFF7161ADFE59FBFF" box="[228,467,1009,1034]" italics="true" pageId="16" pageNumber="17">Pseudorphnus hiboni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAA16D7FFFC1617AFE4F61AEFD9CFBFC" box="[474,534,1010,1033]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
<typeStatus id="0C5DB2B4FFC1617AFD8861AEFDDFFBFF" box="[541,597,1010,1034]" pageId="16" pageNumber="17">Type</typeStatus>
/
<typeStatus id="0C5DB2B4FFC1617AFDFE61AEFD2FFBFF" box="[619,677,1010,1034]" pageId="16" pageNumber="17">Type</typeStatus>
” (
<collectionCode id="B5F794D3FFC1617AFD2861AEFC93FBFF" box="[701,793,1010,1034]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="D3590C16FFC1617AFF526649FC28FABB" blockId="16.[151,1437,973,2006]" pageId="16" pageNumber="17">
<materialsCitation id="638E064BFFC1617AFF526649FEE6FB17" ID-GBIF-Occurrence="1416610155" collectingDate="2001-01-04" collectingDateMax="2002-02-19" collectingDateMin="2001-01-04" collectionCode="UHHF, ZIN, CASC" collectorName="Madagascar Ranomafana NP &amp; I. Hanski &amp; Kavanaugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas" country="Madagascar" county="Vatoharana" elevation="900" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Fianarantsoa">
<emphasis id="E192D004FFC1617AFF526649FC40FBDB" bold="true" box="[199,970,1045,1070]" pageId="16" pageNumber="17">
Additional material examined.
<collectingCountry id="ABF14C86FFC1617AFDD76649FC9EFBDB" box="[578,788,1045,1070]" name="Madagascar" pageId="16" pageNumber="17">MADAGASCAR</collectingCountry>
:
<collectingRegion id="1122C2F4FFC1617AFCB76649FC4FFBDB" box="[802,965,1045,1070]" country="Madagascar" name="Fianarantsoa" pageId="16" pageNumber="17">Fianarantsoa</collectingRegion>
:
</emphasis>
two females (
<collectionCode id="B5F794D3FFC1617AFBE5664AFB4AFBDB" box="[1136,1216,1046,1070]" pageId="16" pageNumber="17">UHHF</collectionCode>
) and three females (
<collectionCode id="B5F794D3FFC1617AFF0A6666FF59FBA7" box="[159,211,1082,1106]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="16" pageNumber="17">ZIN</collectionCode>
), “
<collectorName id="7E1369C0FFC1617AFF626665FDC7FBA4" box="[247,589,1081,1106]" pageId="16" pageNumber="17">Madagascar Ranomafana NP</collectorName>
,
<collectingCounty id="3A38749AFFC1617AFDCB6665FD56FBA7" box="[606,732,1081,1106]" pageId="16" pageNumber="17">Vatoharana</collectingCounty>
wet forest,
<collectingDate id="B71CD33EFFC1617AFCE16666FC6DFBA7" box="[884,999,1082,1106]" pageId="16" pageNumber="17" value="2002-02-19">
<date id="A7582AD6FFC1617AFCE16666FC6DFBA7" box="[884,999,1082,1106]" pageId="16" pageNumber="17" value="2002-02-19">19.2.2002</date>
</collectingDate>
,
<collectingMethod id="0AA77401FFC1617AFC6C6665FB73FBA4" box="[1017,1273,1081,1106]" pageId="16" pageNumber="17">fish bated pitfall trap</collectingMethod>
, [
<geoCoordinate id="B6D26AD1FFC1617AFA876665FA1DFBA7" box="[1298,1431,1081,1106]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="15" value="-21.3">21°18′00″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FFC1617AFF026601FE96FB83" box="[151,284,1117,1142]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="15" value="47.5">47°30′00″E</geoCoordinate>
]
<collectorName id="7E1369C0FFC1617AFEBF6602FE1BFB83" box="[298,401,1117,1142]" pageId="16" pageNumber="17">I. Hanski</collectorName>
leg.”; five males, five females, &quot;
<collectingCountry id="ABF14C86FFC1617AFC906601FC1BFB83" box="[773,913,1117,1142]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
, Province
<collectingRegion id="1122C2F4FFC1617AFB9D6601FB10FB83" box="[1032,1178,1117,1142]" country="Madagascar" name="Fianarantsoa" pageId="16" pageNumber="17">Fianarantsoa</collectingRegion>
,
<collectingMunicipality id="333D966CFFC1617AFB336601FF47FB6F" pageId="16" pageNumber="17">Ranomafana National Park</collectingMunicipality>
,
<location id="D6395ACDFFC1617AFF7C66DDFE2CFB6C" LSID="urn:lsid:plazi:treatment:5B4FBD00FFC16178FF02617CFEFEFADA:D6395ACDFFC1617AFF7C66DDFE2CFB6C" box="[233,422,1153,1178]" country="Madagascar" county="Vatoharana" latitude="-21.25041" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" name="Talatakely area" pageId="16" pageNumber="17" stateProvince="Fianarantsoa">Talatakely area</location>
,
<elevation id="58CBEB25FFC1617AFE5666DEFD97FB6C" box="[451,541,1154,1178]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="17" unit="m" value="900.0">
<quantity id="141EA1F3FFC1617AFE5666DEFD97FB6C" box="[451,541,1154,1178]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="17" unit="m" value="900.0">900 m</quantity>
</elevation>
, mixed tropical forest,
<collectingDate id="B71CD33EFFC1617AFCF666DDFBD0FB6F" box="[867,1114,1153,1178]" pageId="16" pageNumber="17" value="2001-01-04" valueMax="2001-01-16" valueMin="2001-01-04">
<date id="A7582AD6FFC1617AFCF666DDFBD0FB6F" box="[867,1114,1153,1178]" pageId="16" pageNumber="17" value="2001-01-04" valueMax="2001-01-16" valueMin="2001-01-04">4–16 January 2001</date>
</collectingDate>
/
<geoCoordinate id="B6D26AD1FFC1617AFB0766DDFA80FB6F" box="[1170,1290,1153,1178]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="1" value="-21.25041">21.25041S</geoCoordinate>
<geoCoordinate id="B6D26AD1FFC1617AFAB766DDFA16FB6F" box="[1314,1436,1153,1178]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="47.41945">47.41945E</geoCoordinate>
D.H.&amp;K.M.
<collectorName id="7E1369C0FFC1617AFE8366F9FE2AFB4B" box="[278,416,1189,1214]" pageId="16" pageNumber="17">Kavanaugh</collectorName>
,
<collectorName id="7E1369C0FFC1617AFE3A66FAFD91FB48" box="[431,539,1190,1213]" pageId="16" pageNumber="17">R.L.Brett</collectorName>
,
<collectorName id="7E1369C0FFC1617AFDBF66F9FD07FB4B" box="[554,653,1189,1214]" pageId="16" pageNumber="17">E.Elsom</collectorName>
, and
<collectorName id="7E1369C0FFC1617AFD5B66FAFCA5FB4B" box="[718,815,1190,1214]" pageId="16" pageNumber="17">F.Vargas</collectorName>
colls.
<collectingMethod id="0AA77401FFC1617AFCE566F9FC7AFB48" box="[880,1008,1189,1214]" pageId="16" pageNumber="17">pitfal traps</collectingMethod>
COL-DHK-2001-001TN / CASENT 8006675&quot; (
<collectionCode id="B5F794D3FFC1617AFE806696FEEEFB17" box="[277,356,1226,1250]" pageId="16" pageNumber="17">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFFC1617AFEEC6695FD65FAF3" ID-GBIF-Occurrence="1416610161" collectingDate="2005-02-02" collectionCode="ZIN" collectorName="H. Viljanen" country="Madagascar" latitude="-21.3" location="Vatoharana" longLatPrecision="21" longitude="47.5" pageId="16" pageNumber="17" specimenCount="1">
three males, “
<collectingCountry id="ABF14C86FFC1617AFD816695FD14FB17" box="[532,670,1225,1250]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
Ranomafana NP,
<location id="D6395ACDFFC1617AFCF96695FC60FB17" LSID="urn:lsid:plazi:treatment:5B4FBD00FFC16178FF02617CFEFEFADA:D6395ACDFFC1617AFCF96695FC60FB17" box="[876,1002,1225,1250]" country="Madagascar" latitude="-21.3" longLatPrecision="21" longitude="47.5" name="Vatoharana" pageId="16" pageNumber="17">Vatoharana</location>
wet forest,
<collectingDate id="B71CD33EFFC1617AFBE76695FB5DFB17" box="[1138,1239,1225,1250]" pageId="16" pageNumber="17" value="2005-02-02">
<date id="A7582AD6FFC1617AFBE76695FB5DFB17" box="[1138,1239,1225,1250]" pageId="16" pageNumber="17" value="2005-02-02">2.2.2005</date>
</collectingDate>
, chicken int. trap [
<geoCoordinate id="B6D26AD1FFC1617AFF0B66B1FEA9FAF3" box="[158,291,1261,1286]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="15" value="-21.3">21°18′00″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FFC1617AFEA066B1FE36FAF3" box="[309,444,1261,1286]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="15" value="47.5">47°30′00″E</geoCoordinate>
],
<collectorName id="7E1369C0FFC1617AFE4166B2FDD1FAF3" box="[468,603,1261,1286]" pageId="16" pageNumber="17">H. Viljanen</collectorName>
leg.” (
<collectionCode id="B5F794D3FFC1617AFD2666B2FD6DFAF3" box="[691,743,1262,1286]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="16" pageNumber="17">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFFC1617AFC9466B2FC14FABB" ID-GBIF-Occurrence="1416610129" collectingDate="2001-01-05" collectingDateMax="2003-12-02" collectingDateMin="2001-01-05" collectionCode="MNHN, ZIN, ABCB" collectorName="Ilkka Hanski &amp; S. Murzin" country="Madagascar" elevation="900" location="Fianarantsoa" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Fianarantsoa">
one male (
<collectionCode id="B5F794D3FFC1617AFC1466B2FC50FAF3" box="[897,986,1262,1286]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="17">MNHN</collectionCode>
) and one female (
<collectionCode id="B5F794D3FFC1617AFB2E66B2FB65FAF3" box="[1211,1263,1262,1286]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="16" pageNumber="17">ZIN</collectionCode>
), “
<collectingCountry id="ABF14C86FFC1617AFA8766B1FA16FAF3" box="[1298,1436,1261,1286]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
Ranomafana NP, wet forest,
<collectingDate id="B71CD33EFFC1617AFE4D674EFDC1FADF" box="[472,587,1298,1322]" pageId="16" pageNumber="17" value="2003-12-02">
<date id="A7582AD6FFC1617AFE4D674EFDC1FADF" box="[472,587,1298,1322]" pageId="16" pageNumber="17" value="2003-12-02">12.2.2003</date>
</collectingDate>
,
<collectingMethod id="0AA77401FFC1617AFDC2674DFD4EFADC" box="[599,708,1297,1322]" pageId="16" pageNumber="17">dung trap</collectingMethod>
,
<collectorName id="7E1369C0FFC1617AFD45674DFCEAFADF" box="[720,864,1297,1322]" pageId="16" pageNumber="17">Ilkka Hanski</collectorName>
leg.”; one male and one female, “
<collectingCountry id="ABF14C86FFC1617AFB48674DFAEDFADF" box="[1245,1383,1297,1322]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
env.
<collectingRegion id="1122C2F4FFC1617AFF026769FEACFABB" box="[151,294,1333,1358]" country="Madagascar" name="Fianarantsoa" pageId="16" pageNumber="17">Fianarantsoa</collectingRegion>
Ranomafana
<elevation id="58CBEB25FFC1617AFE54676AFD8BFABB" box="[449,513,1334,1358]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="17" unit="m" value="900.0">
<quantity id="141EA1F3FFC1617AFE54676AFD8BFABB" box="[449,513,1334,1358]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="17" unit="m" value="900.0">900m</quantity>
</elevation>
<collectingDate id="B71CD33EFFC1617AFD9D6769FD05FABB" box="[520,655,1333,1358]" pageId="16" pageNumber="17" value="2001-01-05" valueMax="2001-01-15" valueMin="2001-01-05">
<date id="A7582AD6FFC1617AFD9D6769FD05FABB" box="[520,655,1333,1358]" pageId="16" pageNumber="17" value="2001-01-05" valueMax="2001-01-15" valueMin="2001-01-05">5–15.I.2001</date>
</collectingDate>
<collectorName id="7E1369C0FFC1617AFD036769FC8AFABB" box="[662,768,1333,1358]" pageId="16" pageNumber="17">S.Murzin</collectorName>
leg” (
<collectionCode id="B5F794D3FFC1617AFCD1676AFC1CFABB" box="[836,918,1334,1358]" pageId="16" pageNumber="17">ABCB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFC1617AFF526705FDCBF9F4" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="D3590C16FFC1617AFF526705FDCBF9F4" blockId="16.[151,1437,973,2006]" pageId="16" pageNumber="17">
<emphasis id="E192D004FFC1617AFF526705FECBFA87" bold="true" box="[199,321,1369,1394]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
<taxonomicName id="14E67795FFC1617AFED96705FDCAFA87" box="[332,576,1369,1394]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hiboni">
<emphasis id="E192D004FFC1617AFED96705FDCAFA87" box="[332,576,1369,1394]" italics="true" pageId="16" pageNumber="17">Pseudorphnus hiboni</emphasis>
</taxonomicName>
is easily recognized by its large size: the body is longer than 11.0 mm, whereas the specimens of the other species are mostly shorter than
<quantity id="141EA1F3FFC1617AFC1F6721FC7BFA60" box="[906,1009,1405,1430]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="16" pageNumber="17" unit="mm" value="10.5">10.5 mm</quantity>
. It also differs in having the relatively slender horns in either sides of the excavation near the anterior margin of the pronotum in males, finger-shaped apices of the parameres, and the base of the elytra bordered and densely punctate with punctures similar to those in the striae and forming a “tile” pattern.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFC16178FF526451FBD9FBE5" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="description">
<paragraph id="D3590C16FFC1617AFF526451FECBF9BF" blockId="16.[151,1437,973,2006]" pageId="16" pageNumber="17">
<emphasis id="E192D004FFC1617AFF526451FE2EF9D3" bold="true" box="[199,420,1549,1574]" pageId="16" pageNumber="17">Description. Male.</emphasis>
Body elongated, strongly shiny (
<figureCitation id="4BDD1093FFC1617AFC8E6451FCE0F9D3" box="[795,874,1549,1574]" captionStart="FIGURE 7" captionStartId="17.[151,250,1807,1829]" captionTargetBox="[220,1364,197,1782]" captionTargetId="figure@17.[206,1380,193,1796]" captionTargetPageId="17" captionText="FIGURE 7. Pseudorphnus hiboni. Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200451/files/figure.png" pageId="16" pageNumber="17">Figs. 7</figureCitation>
A, F). Color brown, elytra, and underside of body slightly lighter.
</paragraph>
<paragraph id="D3590C16FFC1617AFF476409FBAFF90B" blockId="16.[151,1437,973,2006]" pageId="16" pageNumber="17">Frontoclypeus slightly convex anteriorly, rounded laterally, anterior margin setose and crenulate in dorsal view. Eyes relatively large (diameter larger than the distance between eye and gula in ventral view), incompletely divided by canthus into smaller dorsal and larger ventral parts. Frontoclypeus with relatively long horn rounded apically. Dorsal surface of head impunctate. Labrum bilobate, slightly sinuate in the middle and relatively feebly protruding past frontoclypeus. Length in the middle is 1/8 width (in dorsal view).</paragraph>
<paragraph id="D3590C16FFC1617AFF526556FEEFF820" blockId="16.[151,1437,973,2006]" pageId="16" pageNumber="17">
Pronotum 1.5 times wider than long; widest medially. Anterior margin with wide border, base with fine border. Lateral margins densely punctate, appearing crenulate in dorsal view. Disc of pronotum with deep, somewhat transverse excavation in the middle, with 2 horns bordering excavation near anterior margin. Pronotal horns as long as frontoclypeal horn but more slender, directed upward and forward (
<figureCitation id="4BDD1093FFC1617AFC256529FC79F87B" box="[944,1011,1909,1934]" captionStart="FIGURE 7" captionStartId="17.[151,250,1807,1829]" captionTargetBox="[220,1364,197,1782]" captionTargetId="figure@17.[206,1380,193,1796]" captionTargetPageId="17" captionText="FIGURE 7. Pseudorphnus hiboni. Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200451/files/figure.png" pageId="16" pageNumber="17">Fig. 7</figureCitation>
B). Surface of disc between the horns smooth, without punctures. Sides of pronotum rugose, somewhat tuberculate posteriorly. Lateral margins with long, brown setae.
</paragraph>
<caption id="87995C9EFFC0617BFF026553FDC3F897" httpUri="https://zenodo.org/record/200451/files/figure.png" pageId="17" pageNumber="18" targetBox="[220,1364,197,1782]" targetPageId="17">
<paragraph id="D3590C16FFC0617BFF026553FDC3F897" blockId="17.[151,1436,1807,1890]" pageId="17" pageNumber="18">
<emphasis id="E192D004FFC0617BFF026553FE98F8D0" bold="true" box="[151,274,1807,1829]" pageId="17" pageNumber="18">FIGURE 7.</emphasis>
<emphasis id="E192D004FFC0617BFE8C6553FE72F8D0" box="[281,504,1807,1829]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="14E67795FFC0617BFE8C6553FE7EF8D0" box="[281,500,1807,1829]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="hiboni">Pseudorphnus hiboni</taxonomicName>
.
</emphasis>
Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H).
</paragraph>
</caption>
<paragraph id="D3590C16FFC0617BFF5265D3FB94F85D" blockId="17.[151,1437,1935,2032]" box="[199,1054,1935,1960]" pageId="17" pageNumber="18">Scutellum triangular, narrowly rounded apically, about 1/10 length of elytra.</paragraph>
<paragraph id="D3590C16FFC06178FF5265EFFB78FF02" blockId="17.[151,1437,1935,2032]" lastBlockId="18.[151,1437,151,1328]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
Elytra convex, with feebly marked humeral humps. Maximum width approximately at the middle. Elytra with 7 distinct striae on disc. Striae with characteristic, narrow, semicircular punctures. Each puncture with a short, yellow seta. Only sutural stria reaches elytral apex. Striae 2–5 about 2/3 length of elytron, striae 6–7 about 1/3–1/4 length of elytron and do not reach elytral base. Epipleura with long, sparse, brown setae. Base of elytra bordered and densely punctate with punctures similar to those in striae and forming a “tile” pattern (
<figureCitation id="4BDD1093FFC36178FB1B6283FB5AFF0D" box="[1166,1232,223,248]" captionStart="FIGURE 7" captionStartId="17.[151,250,1807,1829]" captionTargetBox="[220,1364,197,1782]" captionTargetId="figure@17.[206,1380,193,1796]" captionTargetPageId="17" captionText="FIGURE 7. Pseudorphnus hiboni. Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200451/files/figure.png" pageId="18" pageNumber="19">Fig. 7</figureCitation>
E).
</paragraph>
<paragraph id="D3590C16FFC36178FF526358FEDDFE25" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="19">
Protibiae of typical shape of
<taxonomicName id="14E67795FFC36178FD826358FD3DFEE8" box="[535,695,260,285]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="E192D004FFC36178FD826358FD3DFEE8" box="[535,695,260,285]" italics="true" pageId="18" pageNumber="19">Pseudorphnus</emphasis>
</taxonomicName>
, with 2 strong outer teeth and a small 1 located basally. Lateral margin basad of outer teeth not crenulate. Apices with 3 robust, spur-like setae and a number of smaller setae. Protarsi well developed, about 4/5 length of protibiae. Claws 1/3 length of apical tarsomere. Apical protarsomere as long as tarsomeres 3 and 4 combined, as thick as other tarsomeres. Ventral surface of protibiae smooth with 2 rows of setae along sides and sparse longer setae in the middle. Ventral surface of femora sparsely punctate, with 1 raised longitudinal line.
</paragraph>
<paragraph id="D3590C16FFC36178FF526380FBB2FDAA" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="19">Mesothoracic and metathoracic legs similar in shape; metafemora and metatibiae about 1/8 longer than mesofemora and mesotibiae. Mesotibiae and metatibiae somewhat triangular, with 2 apical spurs, with inner margin only slightly concave and with 1 transverse keel. Longer tibial spur as long as 2 basal tarsomeres. Claws 1/ 3 length of apical tarsomere. Femora almost impunctate, with 2 rows of long setae.</paragraph>
<paragraph id="D3590C16FFC36178FF526030FE28FD5D" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="19">Abdominal sternites irregularly punctate, pubescent with dense, long setae. Sternite 8 medially as long as sternites 4–5 combined.</paragraph>
<paragraph id="D3590C16FFC36178FF5260E8FC0CFD38" blockId="18.[151,1437,151,1328]" box="[199,902,692,717]" pageId="18" pageNumber="19">Pygidium transverse, irregularly punctate, hidden under elytra.</paragraph>
<paragraph id="D3590C16FFC36178FF52608BFB51FD05" blockId="18.[151,1437,151,1328]" box="[199,1243,727,752]" pageId="18" pageNumber="19">
Aedeagus (
<figureCitation id="4BDD1093FFC36178FED0608BFE0DFD05" box="[325,391,727,752]" captionStart="FIGURE 7" captionStartId="17.[151,250,1807,1829]" captionTargetBox="[220,1364,197,1782]" captionTargetId="figure@17.[206,1380,193,1796]" captionTargetPageId="17" captionText="FIGURE 7. Pseudorphnus hiboni. Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200451/files/figure.png" pageId="18" pageNumber="19">Fig. 7</figureCitation>
C) with slender, finger-shaped apices of parameres, feebly tapering apically.
</paragraph>
<paragraph id="D3590C16FFC36178FF5260A0FE06FC75" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="19">
<emphasis id="E192D004FFC36178FF5260A0FEAFFCE0" bold="true" box="[199,293,764,789]" pageId="18" pageNumber="19">Female.</emphasis>
The female (
<figureCitation id="4BDD1093FFC36178FE5560A0FD8EFCE1" box="[448,516,764,789]" captionStart="FIGURE 7" captionStartId="17.[151,250,1807,1829]" captionTargetBox="[220,1364,197,1782]" captionTargetId="figure@17.[206,1380,193,1796]" captionTargetPageId="17" captionText="FIGURE 7. Pseudorphnus hiboni. Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200451/files/figure.png" pageId="18" pageNumber="19">Fig. 7</figureCitation>
F) differs from the male in having rugose surface of the head, pronotum slightly impressed anteriorly with sparse punctures in the middle, absence of the frontoclypeal horn or tubercle, presence of the distinct spur of protibiae, and absence of the pronotal excavations and ridges. The sexes are similar in general body shape and color.
</paragraph>
<paragraph id="D3590C16FFC36178FF5261D0FBD9FBE5" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="19">
<emphasis id="E192D004FFC36178FF5261D0FEB5FC50" bold="true" box="[199,319,908,933]" pageId="18" pageNumber="19">Variation.</emphasis>
All examined male specimens had well-developed prothoracic ridges and frontoclypeal horns, thus showing a relatively low allometric variability. Sculpture of the elytra varies with some specimens having no distinct rows of the punctures on the elytra, except for the first (sutural) stria. Body length of the examined specimens varied from 11.0 to
<quantity id="141EA1F3FFC36178FE6461ABFDDFFBFA" box="[497,597,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="18" pageNumber="19" unit="mm" value="12.5">12.5 mm</quantity>
(males) and from 11.0 to 13.0 mm (females).
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFC36178FF526640FEFEFADA" pageId="18" pageNumber="19" type="distribution">
<paragraph id="D3590C16FFC36178FF526640FEFEFADA" blockId="18.[151,1437,151,1328]" pageId="18" pageNumber="19">
<emphasis id="E192D004FFC36178FF526640FE73FBC0" bold="true" box="[199,505,1052,1077]" pageId="18" pageNumber="19">Distribution and habitat.</emphasis>
The species was described from Tôlanaro (formerly Fort Dauphin) in southern
<collectingCountry id="ABF14C86FFC36178FF026663FEABFBAD" box="[151,289,1087,1112]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
. All the other findings were made in a few localities in the Ranomafana National Park (
<figureCitation id="4BDD1093FFC36178FA946663FACFFBAD" box="[1281,1349,1087,1112]" captionStart="FIGURE 7" captionStartId="17.[151,250,1807,1829]" captionTargetBox="[220,1364,197,1782]" captionTargetId="figure@17.[206,1380,193,1796]" captionTargetPageId="17" captionText="FIGURE 7. Pseudorphnus hiboni. Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200451/files/figure.png" pageId="18" pageNumber="19">Fig. 7</figureCitation>
H). The park is situated in
<collectingRegion id="1122C2F4FFC36178FEF06638FDE9FB88" box="[357,611,1124,1149]" country="Madagascar" name="Fianarantsoa" pageId="18" pageNumber="19">Fianarantsoa Province</collectingRegion>
, about
<quantity id="141EA1F3FFC36178FD266638FC82FB88" box="[691,776,1124,1149]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="km" value="100.0">100 km</quantity>
west of the Indian Ocean on the east-facing escarpment of Madagascar's central high plateau and covers about 43 0 0 0 ha of tropical wet forest with elevations varying from
<quantity id="141EA1F3FFC36178FF0266F0FE97FB31" box="[151,285,1196,1220]" metricMagnitude="2" metricUnit="m" metricValue="3.87" metricValueMax="4.0" metricValueMin="3.7399999999999998" pageId="18" pageNumber="19" unit="m" value="387.0" valueMax="400.0" valueMin="374.0">
374–
<specimenCount id="C5E0C79FFFC36178FF5B66F0FE97FB31" box="[206,285,1196,1220]" pageId="18" pageNumber="19" type="generic">400 m</specimenCount>
</quantity>
. Some of the examined specimens of
<taxonomicName id="14E67795FFC36178FD4B66F1FCCCFB30" box="[734,838,1196,1221]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hiboni">
<emphasis id="E192D004FFC36178FD4B66F1FCCCFB30" box="[734,838,1196,1221]" italics="true" pageId="18" pageNumber="19">P. hiboni</emphasis>
</taxonomicName>
were captured by pitfall traps baited with fish and chicken intestine or dung and exposed for approximately 45 hours including two nights (Heidi Viljanen, personal communication). Short-time exposures of the traps suggest that the beetles were attracted to the carrion rather than accidental captures.
</paragraph>
</subSubSection>
</treatment>
</document>
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