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<document id="0FFB24868CB3044FA23DF76F2FF15B32" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier &amp; Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FF9C6125FF02673CFBC6FD95" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Triodontus hildebrandtii Fairmaire 1883" docType="treatment" docVersion="16" lastPageNumber="80" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="78" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:title id="D0F00EFDD0750F861E8626723685174A">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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<mods:namePart id="9E05AA3D1D7C8F3B80148FD645CF63F3">Frolov, Andrey V.</mods:namePart>
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<mods:namePart id="06F7D33CC8C053AE3C8E2D78BEC2F214">Montreuil, Olivier</mods:namePart>
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<mods:namePart id="EE9E0CC64967E3D3A6E64599A91AD7C9">Akhmetova, Lilia A.</mods:namePart>
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<treatment id="5B4FBD00FF9C6125FF02673CFBC6FD95" ID-DOI="http://doi.org/10.5281/zenodo.6057699" ID-GBIF-Taxon="125107295" ID-Zenodo-Dep="6057699" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9C6125FF02673CFBC6FD95" httpUri="http://treatment.plazi.org/id/5B4FBD00FF9C6125FF02673CFBC6FD95" lastPageId="79" lastPageNumber="80" pageId="77" pageNumber="78">
<subSubSection id="9BFC5F9DFF9C6127FF02673CFEBDFA6E" pageId="77" pageNumber="78" type="nomenclature">
<paragraph id="D3590C16FF9C6127FF02673CFD2AFA8F" blockId="77.[151,672,1376,1435]" box="[151,672,1376,1402]" pageId="77" pageNumber="78">
<heading id="8811BB7AFF9C6127FF02673CFD2AFA8F" bold="true" box="[151,672,1376,1402]" fontSize="11" level="1" pageId="77" pageNumber="78" reason="1">
<taxonomicName id="14E67795FF9C6127FF02673CFD2AFA8F" authority="Fairmaire, 1883" authorityName="Fairmaire" authorityYear="1883" box="[151,672,1376,1402]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="hildebrandtii">
<emphasis id="E192D004FF9C6127FF02673CFD2AFA8F" bold="true" box="[151,672,1376,1402]" pageId="77" pageNumber="78">
<emphasis id="E192D004FF9C6127FF02673CFE35FA8F" bold="true" box="[151,447,1376,1402]" italics="true" pageId="77" pageNumber="78">Triodontus hildebrandtii</emphasis>
(
<bibRefCitation id="B77771E7FF9C6127FE5B673CFD13FA8F" author="Fairmaire" box="[462,665,1376,1402]" pageId="77" pageNumber="78" refString="Fairmaire, L. (1883) Diagnoses de Coleopteres nouveaux de Madagascar. Le Naturaliste, 5, 364 - 365." type="journal article" year="1883">Fairmaire, 1883</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D3590C16FF9C6127FF0267DEFEBDFA6E" blockId="77.[151,672,1376,1435]" box="[151,311,1410,1435]" pageId="77" pageNumber="78">
(
<figureCitation id="4BDD1093FF9C6127FF0A67DEFF71FA6E" box="[159,251,1410,1435]" captionStart="FIGURE 40" captionStartId="78.[151,250,1404,1426]" captionTargetBox="[184,1405,231,1377]" captionTargetId="figure@78.[148,1437,170,1386]" captionTargetPageId="78" captionText="FIGURE 40. Triodontus hildebrandtii, holotype male. Habitus in dorsal (A) and lateral view (B), internal sac of aedeagus (C), labels of holotype (D), aedeagus in lateral view and parameres in dorsal and ventral view (E)." httpUri="https://zenodo.org/record/200483/files/figure.png" pageId="77" pageNumber="78">Figs. 40</figureCitation>
AE)
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9C6127FF026796FD5EF9EB" pageId="77" pageNumber="78" type="reference_group">
<paragraph id="D3590C16FF9C6127FF026796FDA3FA0A" blockId="77.[151,724,1482,1566]" pageId="77" pageNumber="78">
<treatmentCitationGroup id="F3F62B38FF9C6127FF026796FDA3FA0A" pageId="77" pageNumber="78">
<taxonomicName id="14E67795FF9C6127FF026796FD41FA15" authority="Fairmaire, 1883: 365" authorityName="Fairmaire" authorityPageNumber="365" authorityYear="1883" box="[151,715,1482,1504]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="hildebrandtii">
<emphasis id="E192D004FF9C6127FF026796FEF0FA15" box="[151,378,1482,1504]" italics="true" pageId="77" pageNumber="78">Orphnus hildebrandtii</emphasis>
<treatmentCitation id="52472A07FF9C6127FE156796FDDDFA15" author="Fairmaire" box="[384,599,1482,1504]" page="365" pageId="77" pageNumber="78" year="1883">
<bibRefCitation id="B77771E7FF9C6127FE156796FDAFFA15" author="Fairmaire" box="[384,549,1482,1504]" pageId="77" pageNumber="78" refString="Fairmaire, L. (1883) Diagnoses de Coleopteres nouveaux de Madagascar. Le Naturaliste, 5, 364 - 365." type="journal article" year="1883">Fairmaire, 1883</bibRefCitation>
: 365
</treatmentCitation>
, 1884: 131
</taxonomicName>
.
<taxonomicName id="14E67795FF9C6127FF0267B5FDAFFA0A" authority="Arrow 1912: 29" authorityName="Arrow" authorityPageNumber="29" authorityYear="1912" box="[151,549,1513,1535]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="hildebrandti">
<emphasis id="E192D004FF9C6127FF0267B5FEF9FA0A" box="[151,371,1513,1535]" italics="true" pageId="77" pageNumber="78">Orphnus hildebrandti</emphasis>
:
<treatmentCitation id="52472A07FF9C6127FE1567B6FDAFFA0A" author="Arrow" box="[384,549,1513,1535]" page="29" pageId="77" pageNumber="78" year="1912">
<bibRefCitation id="B77771E7FF9C6127FE1567B6FE75FA0A" author="Arrow" box="[384,511,1513,1535]" pageId="77" pageNumber="78" refString="Arrow, G. J. (1912) Pachypodinae, Pleocominae, Aclopinae, Glaphyrinae, Ochodaeinae, Orphninae, Idiostominae, Hybosorinae, Dynamopinae, Acanthocerinae, Troginae. In: Junk, W. (Ed.), Coleopterorum Catalogus, 43. W. Junk, Berlin, Germany, pp. 1 - 66." type="journal article" year="1912">Arrow 1912</bibRefCitation>
: 29
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D3590C16FF9C6127FF026454FD5EF9EB" blockId="77.[151,724,1482,1566]" box="[151,724,1544,1566]" pageId="77" pageNumber="78">
<treatmentCitationGroup id="F3F62B38FF9C6127FF026454FD5EF9EB" box="[151,724,1544,1566]" pageId="77" pageNumber="78">
<taxonomicName id="14E67795FF9C6127FF026454FD5AF9EB" authority="Paulian 1937: 142" authorityName="Paulian" authorityPageNumber="142" authorityYear="1937" box="[151,720,1544,1566]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="hildebrandti">
<emphasis id="E192D004FF9C6127FF026454FE02F9EB" box="[151,392,1544,1566]" italics="true" pageId="77" pageNumber="78">Triodontus hildebrandti</emphasis>
:
<treatmentCitation id="52472A07FF9C6127FE006454FDDAF9EB" author="Paulian" box="[405,592,1544,1566]" page="142" pageId="77" pageNumber="78" year="1937">
<bibRefCitation id="B77771E7FF9C6127FE006454FD94F9EB" author="Paulian" box="[405,542,1544,1566]" pageId="77" pageNumber="78" refString="Paulian, R. (1937) Faune entomologique de Madagascar. Coleoptera Lamellicornia, Scarabaeidae Acanthocerini, Trogini, Aulonocnemis, Hybosorini, Orphnini et Ochodaeini. Bulletin de l'Academie malgache (nouvelle serie), 19, 129 - 143." type="journal article" year="1937">Paulian 1937</bibRefCitation>
: 142
</treatmentCitation>
, 1977: 1213
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9C6127FF026416FC5CF95E" pageId="77" pageNumber="78" type="materials_examined">
<paragraph id="D3590C16FF9C6127FF026416FD21F972" blockId="77.[151,1437,1610,2031]" pageId="77" pageNumber="78">
<emphasis id="E192D004FF9C6127FF026416FE4CF996" bold="true" box="[151,454,1610,1635]" pageId="77" pageNumber="78">Type material examined.</emphasis>
Holotype (
<figureCitation id="4BDD1093FF9C6127FDDE6416FD21F996" box="[587,683,1610,1635]" captionStart="FIGURE 40" captionStartId="78.[151,250,1404,1426]" captionTargetBox="[184,1405,231,1377]" captionTargetId="figure@78.[148,1437,170,1386]" captionTargetPageId="78" captionText="FIGURE 40. Triodontus hildebrandtii, holotype male. Habitus in dorsal (A) and lateral view (B), internal sac of aedeagus (C), labels of holotype (D), aedeagus in lateral view and parameres in dorsal and ventral view (E)." httpUri="https://zenodo.org/record/200483/files/figure.png" pageId="77" pageNumber="78">Figs. 40</figureCitation>
AE), male, “MUSÉUM PARIS 1906 Coll. Leon FAIRMAIRE /
<taxonomicName id="14E67795FF9C6127FF026432FE6DF972" authority="Hildebrandti Waterh" authorityName="Hildebrandti Waterh" box="[151,487,1646,1671]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="E192D004FF9C6127FF026432FE1BF972" box="[151,401,1646,1671]" italics="true" pageId="77" pageNumber="78">Orphnus Hildebrandti</emphasis>
Waterh
</taxonomicName>
/ 434” (MNHN).
</paragraph>
<paragraph id="D3590C16FF9C6127FF5264CEFC5CF95E" blockId="77.[151,1437,1610,2031]" box="[199,982,1682,1707]" pageId="77" pageNumber="78">
<materialsCitation id="638E064BFF9C6127FF5264CEFC5CF95E" ID-GBIF-Occurrence="1416610131" box="[199,982,1682,1707]" collectionCode="IRSNB" country="Madagascar" location="Two" pageId="77" pageNumber="78" specimenCount="1">
<emphasis id="E192D004FF9C6127FF5264CEFDBEF95E" bold="true" box="[199,564,1682,1707]" pageId="77" pageNumber="78">Additional material examined.</emphasis>
<location id="D6395ACDFF9C6127FDAE64CFFDE6F95F" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9C6125FF02673CFBC6FD95:D6395ACDFF9C6127FDAE64CFFDE6F95F" box="[571,620,1683,1706]" country="Madagascar" name="Two" pageId="77" pageNumber="78">Two</location>
males, “
<collectingCountry id="ABF14C86FF9C6127FD5964CEFCD2F95E" box="[716,856,1682,1707]" name="Madagascar" pageId="77" pageNumber="78">Madagascar</collectingCountry>
” (
<collectionCode id="B5F794D3FF9C6127FCE464CFFC40F95E" box="[881,970,1683,1707]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="77" pageNumber="78">IRSNB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9C6127FF5264EAFB52F907" pageId="77" pageNumber="78" type="diagnosis">
<paragraph id="D3590C16FF9C6127FF5264EAFB52F907" blockId="77.[151,1437,1610,2031]" pageId="77" pageNumber="78">
<emphasis id="E192D004FF9C6127FF5264EAFECBF93A" bold="true" box="[199,321,1718,1743]" pageId="77" pageNumber="78">Diagnosis.</emphasis>
<taxonomicName id="14E67795FF9C6127FEDE64EAFDEAF93A" box="[331,608,1718,1743]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="hildebrandtii">
<emphasis id="E192D004FF9C6127FEDE64EAFDEAF93A" box="[331,608,1718,1743]" italics="true" pageId="77" pageNumber="78">Triodontus hildebrandtii</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="14E67795FF9C6127FCAA64EBFC28F93B" box="[831,930,1719,1742]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="viettei">
<emphasis id="E192D004FF9C6127FCAA64EBFC28F93B" box="[831,930,1719,1742]" italics="true" pageId="77" pageNumber="78">T. viettei</emphasis>
</taxonomicName>
but differs from it in having outer lobes of parameres with wide lateral notches, as opposed to having slit-shaped lateral notches in
<taxonomicName id="14E67795FF9C6127FBE46480FB5AF906" box="[1137,1232,1756,1779]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="viettei">
<emphasis id="E192D004FF9C6127FBE46480FB5AF906" box="[1137,1232,1756,1779]" italics="true" pageId="77" pageNumber="78">T. viettei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9C6125FF5264A2FCA5FF02" lastPageId="79" lastPageNumber="80" pageId="77" pageNumber="78" type="description">
<paragraph id="D3590C16FF9C6127FF5264A2FD93F8CE" blockId="77.[151,1437,1610,2031]" pageId="77" pageNumber="78">
<emphasis id="E192D004FF9C6127FF5264A2FE2EF8E2" bold="true" box="[199,420,1790,1815]" pageId="77" pageNumber="78">Description. Male.</emphasis>
Body length 15.0 mm. Color uniformly dark brown, legs, antennae, and underside of the body slightly paler (
<figureCitation id="4BDD1093FF9C6127FEED657EFE5FF8CE" box="[376,469,1826,1851]" captionStart="FIGURE 40" captionStartId="78.[151,250,1404,1426]" captionTargetBox="[184,1405,231,1377]" captionTargetId="figure@78.[148,1437,170,1386]" captionTargetPageId="78" captionText="FIGURE 40. Triodontus hildebrandtii, holotype male. Habitus in dorsal (A) and lateral view (B), internal sac of aedeagus (C), labels of holotype (D), aedeagus in lateral view and parameres in dorsal and ventral view (E)." httpUri="https://zenodo.org/record/200483/files/figure.png" pageId="77" pageNumber="78">Figs. 40</figureCitation>
AB).
</paragraph>
<paragraph id="D3590C16FF9C6127FF52651AFB8EF81A" blockId="77.[151,1437,1610,2031]" pageId="77" pageNumber="78">Frontoclypeus slightly convex anteriorly, rounded laterally, anterior margin crenulate and setose in dorsal view. Eyes relatively small (diameter slightly smaller than the distance between eye and gula in ventral view), incompletely divided by canthus into small, dorsal and large, ventral parts. Frontoclypeus with tubercles mediad of each eye and with long horn in the center of the frontoclypeus. The horn is longer than width of the head, acutely rounded apically, somewhat curved caudally, slightly rugose on posterior side.</paragraph>
<caption id="87995C9EFF9F6124FF026720FBC0FA44" httpUri="https://zenodo.org/record/200483/files/figure.png" pageId="78" pageNumber="79" targetBox="[184,1405,231,1377]" targetPageId="78">
<paragraph id="D3590C16FF9F6124FF026720FBC0FA44" blockId="78.[151,1435,1404,1457]" pageId="78" pageNumber="79">
<emphasis id="E192D004FF9F6124FF026720FEAAFA67" bold="true" box="[151,288,1404,1426]" pageId="78" pageNumber="79">FIGURE 40.</emphasis>
<taxonomicName id="14E67795FF9F6124FEB06720FD94FA67" box="[293,542,1404,1426]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="hildebrandtii">
<emphasis id="E192D004FF9F6124FEB06720FD94FA67" box="[293,542,1404,1426]" italics="true" pageId="78" pageNumber="79">Triodontus hildebrandtii</emphasis>
</taxonomicName>
, holotype male. Habitus in dorsal (A) and lateral view (B), internal sac of aedeagus (C), labels of holotype (D), aedeagus in lateral view and parameres in dorsal and ventral view (E).
</paragraph>
</caption>
<paragraph id="D3590C16FF9F6124FF526783FD21F95F" blockId="78.[151,1437,1501,2030]" pageId="78" pageNumber="79">Pronotum with a bulge medially, with large excavations each side of bulge, and with 2 ridge-shaped tubercles laterally of each excavation. Median bulge is rounded and somewhat depressed on top, with 2 distinct tubercles and a depression, receiving frontoclypeal horn. Lateral margins with wide border appearing somewhat crenulate (in dorsal view), with a row of brown setae. Anterior margin with wide, smooth border. Posterior margin with fine border; not crenulate; punctate with small, narrow, longitudinal punctures. Surface of most pronotum almost smooth, with minute, feebly-visible punctation.</paragraph>
<paragraph id="D3590C16FF9F6124FF5264E9FC5CF93B" blockId="78.[151,1437,1501,2030]" box="[199,982,1717,1742]" pageId="78" pageNumber="79">Scutellum rounded apically, visible part is about 1/13 length of elytra.</paragraph>
<paragraph id="D3590C16FF9F6124FF526485FD59F8AB" blockId="78.[151,1437,1501,2030]" pageId="78" pageNumber="79">Elytra convex, with distinct humeral humps. Maximum width approximately at basal 1/3. Elytra with 10 feebly visible striae on disc and with oblique line from base of 6th stria to approximately middle of 8th stria. Stria with small punctures. Intervals with sparse, minute punctures, almost smooth. Base of elytra with an irregular row of coarse punctures each bearing a small, yellow seta.</paragraph>
<paragraph id="D3590C16FF9F6124FF526535FE7DF853" blockId="78.[151,1437,1501,2030]" pageId="78" pageNumber="79">Sternite 8 medially as wide as sternites 25 combined, without tubercle or concavity in the middle, with apical border deeply sinuate medially.</paragraph>
<paragraph id="D3590C16FF9F6124FF5265EEFAE6F81B" blockId="78.[151,1437,1501,2030]" pageId="78" pageNumber="79">
Parameres with outer lobes being relatively wide (in dorsal view) and having slit-shaped lateral notches (
<figureCitation id="4BDD1093FF9F6124FAFB65EDFF3EF81B" captionStart="FIGURE 40" captionStartId="78.[151,250,1404,1426]" captionTargetBox="[184,1405,231,1377]" captionTargetId="figure@78.[148,1437,170,1386]" captionTargetPageId="78" captionText="FIGURE 40. Triodontus hildebrandtii, holotype male. Habitus in dorsal (A) and lateral view (B), internal sac of aedeagus (C), labels of holotype (D), aedeagus in lateral view and parameres in dorsal and ventral view (E)." httpUri="https://zenodo.org/record/200483/files/figure.png" pageId="78" pageNumber="79">Fig. 40</figureCitation>
E). Internal sac of aedeagus with 1 strongly sclerotized sclerite and 2 additional smaller sclerites (
<figureCitation id="4BDD1093FF9F6124FB6C6589FAC6F81B" box="[1273,1356,2005,2030]" captionStart="FIGURE 40" captionStartId="78.[151,250,1404,1426]" captionTargetBox="[184,1405,231,1377]" captionTargetId="figure@78.[148,1437,170,1386]" captionTargetPageId="78" captionText="FIGURE 40. Triodontus hildebrandtii, holotype male. Habitus in dorsal (A) and lateral view (B), internal sac of aedeagus (C), labels of holotype (D), aedeagus in lateral view and parameres in dorsal and ventral view (E)." httpUri="https://zenodo.org/record/200483/files/figure.png" pageId="78" pageNumber="79">Fig. 40</figureCitation>
C).
</paragraph>
<paragraph id="D3590C16FF9E6125FF5262CBFE28FF45" blockId="79.[151,1437,151,608]" box="[199,418,151,176]" pageId="79" pageNumber="80">
<emphasis id="E192D004FF9E6125FF5262CBFEAFFF45" bold="true" box="[199,293,151,176]" pageId="79" pageNumber="80">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="D3590C16FF9E6125FF5262E0FCA5FF02" blockId="79.[151,1437,151,608]" pageId="79" pageNumber="80">
<emphasis id="E192D004FF9E6125FF5262E0FEB6FF20" bold="true" box="[199,316,188,213]" pageId="79" pageNumber="80">Variation.</emphasis>
The non-type specimens differ from the holotype in slightly less-developed frontoclypeal and pronotal armature. Body length varies from 14.0
<quantity id="141EA1F3FF9E6125FD506283FCA1FF02" box="[709,811,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="79" pageNumber="80" unit="mm" value="14.5">14.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9E6125FF526358FEA2FEB5" pageId="79" pageNumber="80" type="distribution">
<paragraph id="D3590C16FF9E6125FF526358FEA2FEB5" blockId="79.[151,1437,151,608]" pageId="79" pageNumber="80">
<emphasis id="E192D004FF9E6125FF526358FEEAFEE8" bold="true" box="[199,352,260,285]" pageId="79" pageNumber="80">Distribution.</emphasis>
The distribution of this species is yet unknown since all three known specimens lack precise locality data.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9E6125FF526310FBC6FD95" pageId="79" pageNumber="80" type="discussion">
<paragraph id="D3590C16FF9E6125FF526310FD8FFE00" blockId="79.[151,1437,151,608]" pageId="79" pageNumber="80">
<materialsCitation id="638E064BFF9E6125FF526310FD8FFE00" ID-GBIF-Occurrence="1416610137" collectionCode="MNHN" collectorName="Alluaud &amp; This &amp; We" county="There" location="Tanala forest" municipality="Paulian" pageId="79" pageNumber="80" specimenCount="1" typeStatus="holotype">
<emphasis id="E192D004FF9E6125FF526310FEB1FE90" bold="true" box="[199,315,332,357]" pageId="79" pageNumber="80">Remarks.</emphasis>
Paulian wrote that the
<typeStatus id="0C5DB2B4FF9E6125FDAB6311FDE4FE90" box="[574,622,333,357]" pageId="79" pageNumber="80">type</typeStatus>
of
<taxonomicName id="14E67795FF9E6125FD016311FCC9FE90" box="[660,835,332,357]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="hildebrandtii">
<emphasis id="E192D004FF9E6125FD016311FCC9FE90" box="[660,835,332,357]" italics="true" pageId="79" pageNumber="80">T. hildebrandtii</emphasis>
</taxonomicName>
was unknown to him, but he treated it a valid species and mentioned some material from
<location id="D6395ACDFF9E6125FDD46333FD52FE7D" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9C6125FF02673CFBC6FD95:D6395ACDFF9E6125FDD46333FD52FE7D" box="[577,728,367,392]" county="There" municipality="Paulian" name="Tanala forest" pageId="79" pageNumber="80">Tanala forest</location>
(collection of
<location id="D6395ACDFF9E6125FC196333FC62FE7D" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9C6125FF02673CFBC6FD95:D6395ACDFF9E6125FC196333FC62FE7D" box="[908,1000,367,392]" county="There" municipality="Paulian" name="Alluaud" pageId="79" pageNumber="80">Alluaud</location>
) (
<bibRefCitation id="B77771E7FF9E6125FB916333FB2BFE7D" author="Paulian" box="[1028,1185,367,392]" pageId="79" pageNumber="80" refString="Paulian, R. (1937) Faune entomologique de Madagascar. Coleoptera Lamellicornia, Scarabaeidae Acanthocerini, Trogini, Aulonocnemis, Hybosorini, Orphnini et Ochodaeini. Bulletin de l'Academie malgache (nouvelle serie), 19, 129 - 143." type="journal article" year="1937">
<collectingMunicipality id="333D966CFF9E6125FB916333FBD0FE7D" box="[1028,1114,367,392]" pageId="79" pageNumber="80">Paulian</collectingMunicipality>
1937
</bibRefCitation>
,
<bibRefCitation id="B77771E7FF9E6125FB27632CFB64FE7D" author="Paulian" box="[1202,1262,368,392]" pageId="79" pageNumber="80" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">1977</bibRefCitation>
).
<collectingCounty id="3A38749AFF9E6125FA926333FAC2FE7D" box="[1287,1352,367,392]" pageId="79" pageNumber="80">There</collectingCounty>
is one
<taxonomicName id="14E67795FF9E6125FF0263C8FE9AFE58" box="[151,272,404,429]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="E192D004FF9E6125FF0263C8FE9AFE58" box="[151,272,404,429]" italics="true" pageId="79" pageNumber="80">Triodontus</emphasis>
</taxonomicName>
specimen in the collection of
<collectionCode id="B5F794D3FF9E6125FD1763C9FD50FE59" box="[642,730,405,428]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
, which bears the label “Forêt Tanala (coll.
<collectorName id="7E1369C0FF9E6125FB7263C8FAC2FE58" box="[1255,1352,404,429]" pageId="79" pageNumber="80">Alluaud</collectorName>
)”.
<collectorName id="7E1369C0FF9E6125FAFE63C8FA17FE58" box="[1387,1437,404,429]" pageId="79" pageNumber="80">This</collectorName>
specimen, in our opinion, belongs to
<taxonomicName id="14E67795FF9E6125FDA963E5FD19FE25" box="[572,659,441,464]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="owas">
<emphasis id="E192D004FF9E6125FDA963E5FD19FE25" box="[572,659,441,464]" italics="true" pageId="79" pageNumber="80">T. owas</emphasis>
</taxonomicName>
.
<collectorName id="7E1369C0FF9E6125FD3763E4FD4DFE3A" box="[674,711,440,463]" pageId="79" pageNumber="80">We</collectorName>
think that the application of the name
<taxonomicName id="14E67795FF9E6125FBEA63E5FAA4FE25" box="[1151,1326,439,464]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="hildebrandtii">
<emphasis id="E192D004FF9E6125FBEA63E5FAA4FE25" box="[1151,1326,439,464]" italics="true" pageId="79" pageNumber="80">T. hildebrandtii</emphasis>
</taxonomicName>
to this or other specimens was unjustified.
</materialsCitation>
</paragraph>
<paragraph id="D3590C16FF9E6125FF52605CFBC6FD95" blockId="79.[151,1437,151,608]" pageId="79" pageNumber="80">
In the collection of MNHN we found a specimen with a label “
<taxonomicName id="14E67795FF9E6125FC0D63A3FB78FDED" authority="Hildebrandti Waterh" authorityName="Hildebrandti Waterh" box="[920,1266,511,536]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="E192D004FF9E6125FC0D63A3FB19FDED" box="[920,1171,511,536]" italics="true" pageId="79" pageNumber="80">Orphnus Hildebrandti</emphasis>
Waterh
</taxonomicName>
”. Although the label is not handwritten by Fairmaire, the specimen originated from Fairmaires collection and agrees with the original description of
<taxonomicName id="14E67795FF9E6125FE02601BFD18FD95" box="[407,658,583,608]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="hildebrandtii">
<emphasis id="E192D004FF9E6125FE02601BFD18FD95" box="[407,658,583,608]" italics="true" pageId="79" pageNumber="80">Orphnus hildebrandtii</emphasis>
</taxonomicName>
. Therefore we consider it the holotype.
</paragraph>
</subSubSection>
</treatment>
</document>