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<document id="B5A35A29217A0DEE0321ADCDAF0D97E4" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier &amp; Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FF84613DFF026380FA8AFEE9" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Triodontus copridoides Paulian 1977" docType="treatment" docVersion="16" lastPageNumber="88" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="86" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:title id="54D6F5028044E86C785D51AC5E8095E2">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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<mods:namePart id="0654D549E50CDA3404B326576535D94D">Frolov, Andrey V.</mods:namePart>
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<mods:namePart id="5E9ED7771DD972A73FFAD05FCE5323B8">Montreuil, Olivier</mods:namePart>
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<mods:namePart id="252CECBBFAE17923E5446515E8DB6C92">Akhmetova, Lilia A.</mods:namePart>
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<treatment id="5B4FBD00FF84613DFF026380FA8AFEE9" ID-DOI="http://doi.org/10.5281/zenodo.6057710" ID-GBIF-Taxon="125107305" ID-Zenodo-Dep="6057710" LSID="urn:lsid:plazi:treatment:5B4FBD00FF84613DFF026380FA8AFEE9" httpUri="http://treatment.plazi.org/id/5B4FBD00FF84613DFF026380FA8AFEE9" lastPageId="87" lastPageNumber="88" pageId="85" pageNumber="86">
<subSubSection id="9BFC5F9DFF84613FFF026380FEBCFDE2" pageId="85" pageNumber="86" type="nomenclature">
<paragraph id="D3590C16FF84613FFF026380FDD4FE03" blockId="85.[151,606,476,535]" box="[151,606,476,502]" pageId="85" pageNumber="86">
<heading id="8811BB7AFF84613FFF026380FDD4FE03" bold="true" box="[151,606,476,502]" fontSize="11" level="1" pageId="85" pageNumber="86" reason="1">
<taxonomicName id="14E67795FF84613FFF026380FDD4FE03" authority="Paulian, 1977" authorityName="Paulian" authorityYear="1977" box="[151,606,476,502]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="copridoides">
<emphasis id="E192D004FF84613FFF026380FDD4FE03" bold="true" box="[151,606,476,502]" pageId="85" pageNumber="86">
<emphasis id="E192D004FF84613FFF026380FE27FE03" bold="true" box="[151,429,476,502]" italics="true" pageId="85" pageNumber="86">Triodontus copridoides</emphasis>
<bibRefCitation id="B77771E7FF84613FFE216380FDD4FE03" author="Paulian" box="[436,606,476,502]" pageId="85" pageNumber="86" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="D3590C16FF84613FFF0263A2FEBCFDE2" blockId="85.[151,606,476,535]" box="[151,310,510,535]" pageId="85" pageNumber="86">
(
<figureCitation id="4BDD1093FF84613FFF0A63A2FF70FDE2" box="[159,250,510,535]" captionStart="FIGURE 44" captionStartId="86.[151,250,1871,1893]" captionTargetBox="[185,1396,298,1847]" captionTargetId="figure@86.[184,1403,229,1850]" captionTargetPageId="86" captionText="FIGURE 44. Triodontus copridoides, holotype male (A, B, D, E), male (C). Habitus in dorsal view (A, C), labels (D), aedeagus in lateral view and parameres in dorsal and ventral view (D), abdomen (E), distributional records (F)." httpUri="https://zenodo.org/record/200487/files/figure.png" pageId="85" pageNumber="86">Figs. 44</figureCitation>
AF)
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF84613FFF02601AFDD0FDA9" box="[151,602,582,604]" pageId="85" pageNumber="86" type="reference_group">
<paragraph id="D3590C16FF84613FFF02601AFDD0FDA9" blockId="85.[151,602,582,604]" box="[151,602,582,604]" pageId="85" pageNumber="86">
<treatmentCitationGroup id="F3F62B38FF84613FFF02601AFDD0FDA9" box="[151,602,582,604]" pageId="85" pageNumber="86">
<taxonomicName id="14E67795FF84613FFF02601AFDDCFDA9" authority="Paulian, 1977: 1211" authorityName="Paulian" authorityPageNumber="1211" authorityYear="1977" box="[151,598,582,604]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="copridoides">
<emphasis id="E192D004FF84613FFF02601AFE09FDA9" box="[151,387,582,604]" italics="true" pageId="85" pageNumber="86">Triodontus copridoides</emphasis>
<treatmentCitation id="52472A07FF84613FFE1C601AFDDCFDA9" author="Paulian" box="[393,598,582,604]" page="1211" pageId="85" pageNumber="86" year="1977">
<bibRefCitation id="B77771E7FF84613FFE1C601AFD9DFDA9" author="Paulian" box="[393,535,582,604]" pageId="85" pageNumber="86" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
: 1211
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF84613FFF0260D5FE29FCC7" pageId="85" pageNumber="86" type="materials_examined">
<paragraph id="D3590C16FF84613FFF0260D5FCE3FD1F" blockId="85.[151,1437,649,2006]" pageId="85" pageNumber="86">
<materialsCitation id="638E064BFF84613FFF0260D5FCEFFD1F" ID-GBIF-Occurrence="1416610087" collectionCode="MUSEUM, PARIS, COLL, OBERTHUR, TYPE, MNHN" collectorName="Ex Musaeo D." country="Madagascar" determinerName="R. Paulian" location="Madagascar" pageId="85" pageNumber="86" specimenCode="PARIS 1952" specimenCount="3" typeStatus="holotype">
<emphasis id="E192D004FF84613FFF0260D5FE4CFD57" bold="true" box="[151,454,649,674]" pageId="85" pageNumber="86">
<typeStatus id="0C5DB2B4FF84613FFF0260D5FF58FD57" box="[151,210,649,674]" pageId="85" pageNumber="86">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="0C5DB2B4FF84613FFE4460D5FDB3FD57" box="[465,569,649,674]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
(
<figureCitation id="4BDD1093FF84613FFDD960D5FD26FD57" box="[588,684,649,674]" captionStart="FIGURE 44" captionStartId="86.[151,250,1871,1893]" captionTargetBox="[185,1396,298,1847]" captionTargetId="figure@86.[184,1403,229,1850]" captionTargetPageId="86" captionText="FIGURE 44. Triodontus copridoides, holotype male (A, B, D, E), male (C). Habitus in dorsal view (A, C), labels (D), aedeagus in lateral view and parameres in dorsal and ventral view (D), abdomen (E), distributional records (F)." httpUri="https://zenodo.org/record/200487/files/figure.png" pageId="85" pageNumber="86">Figs. 44</figureCitation>
AB, DE), male, “
<collectingCountry id="ABF14C86FF84613FFC0860D5FBADFD57" box="[925,1063,649,674]" name="Madagascar" pageId="85" pageNumber="86">Madagascar</collectingCountry>
Cowans [= Ankafina Tsarafidy, according to
<bibRefCitation id="B77771E7FF84613FFEA060F1FE68FD33" author="Paulian" box="[309,482,685,710]" pageId="85" pageNumber="86" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian (1977)</bibRefCitation>
] /
<collectorName id="7E1369C0FF84613FFD9F60F2FD24FD33" box="[522,686,685,710]" pageId="85" pageNumber="86">Ex Musaeo D.</collectorName>
Sharp 1890 /
<collectionCode id="B5F794D3FF84613FFCC260F1FC58FD33" box="[855,978,685,710]" pageId="85" pageNumber="86">MUSÉUM</collectionCode>
<specimenCode id="8340A46DFF84613FFC4A60F2FBFAFD33" box="[991,1136,685,710]" collectionCode="PARIS" pageId="85" pageNumber="86">
<collectionCode id="B5F794D3FF84613FFC4A60F2FBA6FD33" box="[991,1068,686,710]" pageId="85" pageNumber="86">PARIS</collectionCode>
1952
</specimenCode>
<collectionCode id="B5F794D3FF84613FFBE860F2FB4CFD33" box="[1149,1222,686,710]" pageId="85" pageNumber="86">COLL</collectionCode>
R
<collectionCode id="B5F794D3FF84613FFB6760F2FA0DFD33" box="[1266,1415,686,710]" pageId="85" pageNumber="86">OBERTHUR</collectionCode>
/
<taxonomicName id="14E67795FF84613FFF02608DFE11FD1F" box="[151,411,721,746]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="copridoides" status="sp. nov.">
<emphasis id="E192D004FF84613FFF02608DFE11FD1F" box="[151,411,721,746]" italics="true" pageId="85" pageNumber="86">Triodontus copridoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAA16D7FFF84613FFE37608EFE54FD1C" box="[418,478,722,745]" pageId="85" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
<determinerName id="2685E0D2FF84613FFE70608EFDD0FD1F" box="[485,602,721,746]" pageId="85" pageNumber="86">R. Paulian</determinerName>
det. /
<collectionCode id="B5F794D3FF84613FFD35608EFD62FD1F" box="[672,744,722,746]" pageId="85" pageNumber="86">
<typeStatus id="0C5DB2B4FF84613FFD35608EFD62FD1F" box="[672,744,722,746]" pageId="85" pageNumber="86">TYPE</typeStatus>
</collectionCode>
” (
<collectionCode id="B5F794D3FF84613FFC94608EFCD7FD1F" box="[769,861,722,746]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="85" pageNumber="86">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="D3590C16FF84613FFF5260A9FE29FCC7" blockId="85.[151,1437,649,2006]" pageId="85" pageNumber="86">
<materialsCitation id="638E064BFF84613FFF5260A9FE29FCC7" ID-GBIF-Occurrence="1416610085" collectionCode="SMTFD" country="Madagascar" location="Amber Mountains" municipality="Amber" pageId="85" pageNumber="86" specimenCount="1" stateProvince="Antsiranana">
<emphasis id="E192D004FF84613FFF5260A9FC62FCFB" bold="true" box="[199,1000,757,782]" pageId="85" pageNumber="86">
Additional material examined.
<collectingCountry id="ABF14C86FF84613FFDF560A9FCB8FCFB" box="[608,818,757,782]" name="Madagascar" pageId="85" pageNumber="86">MADAGASCAR</collectingCountry>
:
<collectingRegion id="1122C2F4FF84613FFCDF60A9FC69FCFB" box="[842,995,757,782]" country="Madagascar" name="Antsiranana" pageId="85" pageNumber="86">Antsiranana</collectingRegion>
:
</emphasis>
two males, “
<collectingMunicipality id="333D966CFF84613FFB0B60A9FB64FCFB" box="[1182,1262,757,782]" pageId="85" pageNumber="86">Amber</collectingMunicipality>
geb. [
<location id="D6395ACDFF84613FFADE60A9FE9EFCC7" LSID="urn:lsid:plazi:treatment:5B4FBD00FF84613DFF026380FA8AFEE9:D6395ACDFF84613FFADE60A9FE9EFCC7" country="Madagascar" municipality="Amber" name="Amber Mountains" pageId="85" pageNumber="86" stateProvince="Antsiranana">Amber Mountains</location>
]” (
<collectionCode id="B5F794D3FF84613FFEA66146FE1DFCC7" box="[307,407,794,818]" pageId="85" pageNumber="86">SMTFD</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF84613FFF526161FDE8FC34" pageId="85" pageNumber="86" type="diagnosis">
<paragraph id="D3590C16FF84613FFF526161FDE8FC34" blockId="85.[151,1437,649,2006]" pageId="85" pageNumber="86">
<emphasis id="E192D004FF84613FFF526161FECBFCA3" bold="true" box="[199,321,829,854]" pageId="85" pageNumber="86">Diagnosis.</emphasis>
<taxonomicName id="14E67795FF84613FFEDC6161FDC5FCA3" box="[329,591,829,854]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="copridoides">
<emphasis id="E192D004FF84613FFEDC6161FDC5FCA3" box="[329,591,829,854]" italics="true" pageId="85" pageNumber="86">Triodontus copridoides</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="14E67795FF84613FFCB06162FC18FCA0" box="[805,914,830,853]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="itremoi">
<emphasis id="E192D004FF84613FFCB06162FC18FCA0" box="[805,914,830,853]" italics="true" pageId="85" pageNumber="86">T. itremoi</emphasis>
</taxonomicName>
in general appearance and in the shape of the parameres but it can be separated from it by having the abdominal sternite 8 with a shallow but distinct, somewhat transversal fossa, larger tubercles laterad of each eye, and somewhat shorter and less oblique notches on the lateral sides of the outer lobes of the parameres.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF84613DFF526191FEB9FF20" lastPageId="87" lastPageNumber="88" pageId="85" pageNumber="86" type="description">
<paragraph id="D3590C16FF84613FFF526191FD8FFBFF" blockId="85.[151,1437,649,2006]" pageId="85" pageNumber="86">
<emphasis id="E192D004FF84613FFF526191FE29FC13" bold="true" box="[199,419,973,998]" pageId="85" pageNumber="86">Description. Male.</emphasis>
Color of head, pronotum, and elytra dark brown; legs, antennae, and underside of the body paler, castaneous (
<figureCitation id="4BDD1093FF84613FFEF161ADFE48FBFF" box="[356,450,1009,1034]" captionStart="FIGURE 44" captionStartId="86.[151,250,1871,1893]" captionTargetBox="[185,1396,298,1847]" captionTargetId="figure@86.[184,1403,229,1850]" captionTargetPageId="86" captionText="FIGURE 44. Triodontus copridoides, holotype male (A, B, D, E), male (C). Habitus in dorsal view (A, C), labels (D), aedeagus in lateral view and parameres in dorsal and ventral view (D), abdomen (E), distributional records (F)." httpUri="https://zenodo.org/record/200487/files/figure.png" pageId="85" pageNumber="86">Figs. 44</figureCitation>
A, C).
</paragraph>
<paragraph id="D3590C16FF84613FFF476649FC2BFB4B" blockId="85.[151,1437,649,2006]" pageId="85" pageNumber="86">Frontoclypeus slightly convex anteriorly, rounded laterally, anterior margin crenulate and setose in dorsal view. Eyes relatively small (diameter slightly smaller than the distance between eye and gula in ventral view), incompletely divided by canthus into small, dorsal and large, ventral parts. Frontoclypeus with tubercles mediad of each eye and with long horn in center of frontoclypeus. The horn is longer than width of the head, acutely rounded apically, somewhat curved caudally, slightly rugose on posterior side.</paragraph>
<paragraph id="D3590C16FF84613FFF526696FAF4FA63" blockId="85.[151,1437,649,2006]" pageId="85" pageNumber="86">Pronotum with a bulge medially in distal 1/3, with large excavations aside the bulge, and with 2 ridge-shaped tubercles laterally of each excavation. The median bulge is rounded and somewhat depressed on the top, without distinct tubercles but with a longitudinal depression receiving frontoclypeal horn. Lateral margins with wide border appearing somewhat crenulate on the hind angles (in dorsal view), with a row of brown setae. Anterior margin with wide, smooth border. Posterior margin with fine border; not crenulate; punctate with small, narrow, longitudinal punctures. Surface of pronotum mainly smooth, anterior and posterior angles with sparse and coarse punctation.</paragraph>
<paragraph id="D3590C16FF84613FFF5267FDFC5CFA4F" blockId="85.[151,1437,649,2006]" box="[199,982,1441,1466]" pageId="85" pageNumber="86">Scutellum rounded apically, visible part is about 1/15 length of elytra.</paragraph>
<paragraph id="D3590C16FF84613FFF526799FD35F9BC" blockId="85.[151,1437,649,2006]" pageId="85" pageNumber="86">Elytra convex, with distinct humeral humps. Maximum width approximately at the middle. Elytra with 10 feebly visible striae on disc and with oblique line from base of 6th stria to approximately middle of 8th stria. Stria with small punctures. Intervals with minute punctures, almost smooth. Base of elytra with an irregular row of coarse punctures each bearing small, yellow seta.</paragraph>
<paragraph id="D3590C16FF84613FFF526409FF47F92F" blockId="85.[151,1437,649,2006]" pageId="85" pageNumber="86">Protibiae with 3 outer teeth, not crenulate basad of the teeth. Apices with 3 robust, spur-like setae and a few small setae. Anterior tarsi about 3/4 length of protibiae. Claws 1/3 length of apical tarsomere. Apical protarsomere as long as tarsomere 3 and 4 combined, as thick as other tarsomeres. Ventral surface of protibiae with a longitudinal keel.</paragraph>
<paragraph id="D3590C16FF84613FFF5264B9FBC1F89C" blockId="85.[151,1437,649,2006]" pageId="85" pageNumber="86">Mesothoracic and metathoracic legs similar in shape; metafemora and metatibiae slightly longer than mesofemora and mesotibiae, respectively. Tibiae somewhat triangular, with 2 apical spurs, with inner margin only slightly concave. Longer tibial spur as long as or slightly longer than 2 basal tarsomeres. Claws about 1/3 length of apical tarsomere. Ventral sides of femora almost impunctate; with sparse, long setae.</paragraph>
<paragraph id="D3590C16FF84613FFF526529FE9CF847" blockId="85.[151,1437,649,2006]" pageId="85" pageNumber="86">Sternite 8 medially about 1.5 as short as other sternites combined; with distinct, elongate, transverse fossa in the middle.</paragraph>
<paragraph id="D3590C16FF84613CFF5265E2FB1AFF45" blockId="85.[151,1437,649,2006]" lastBlockId="86.[151,1168,151,176]" lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86">
Parameres with outer lobes somewhat depressed, feebly sclerotized wide apices, and deep oblique fossae on lateral sides (
<figureCitation id="4BDD1093FF87613CFEBE62CBFEF7FF45" box="[299,381,151,176]" captionStart="FIGURE 44" captionStartId="86.[151,250,1871,1893]" captionTargetBox="[185,1396,298,1847]" captionTargetId="figure@86.[184,1403,229,1850]" captionTargetPageId="86" captionText="FIGURE 44. Triodontus copridoides, holotype male (A, B, D, E), male (C). Habitus in dorsal view (A, C), labels (D), aedeagus in lateral view and parameres in dorsal and ventral view (D), abdomen (E), distributional records (F)." httpUri="https://zenodo.org/record/200487/files/figure.png" pageId="86" pageNumber="87">Fig. 44</figureCitation>
D, arrow). Internal sac of aedeagus with sclerotized and long sclerites.
</paragraph>
<caption id="87995C9EFF87613CFF026513FB12F871" httpUri="https://zenodo.org/record/200487/files/figure.png" pageId="86" pageNumber="87" targetBox="[185,1396,298,1847]" targetPageId="86">
<paragraph id="D3590C16FF87613CFF026513FB12F871" blockId="86.[151,1436,1871,1924]" pageId="86" pageNumber="87">
<emphasis id="E192D004FF87613CFF026513FE95F890" bold="true" box="[151,287,1871,1893]" pageId="86" pageNumber="87">FIGURE 44.</emphasis>
<taxonomicName id="14E67795FF87613CFEB16513FD85F890" box="[292,527,1871,1893]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="copridoides">
<emphasis id="E192D004FF87613CFEB16513FD85F890" box="[292,527,1871,1893]" italics="true" pageId="86" pageNumber="87">Triodontus copridoides</emphasis>
</taxonomicName>
, holotype male (A, B, D, E), male (C). Habitus in dorsal view (A, C), labels (D), aedeagus in lateral view and parameres in dorsal and ventral view (D), abdomen (E), distributional records (F).
</paragraph>
</caption>
<paragraph id="D3590C16FF87613CFF526596FE28F816" blockId="86.[199,418,1994,2019]" box="[199,418,1994,2019]" pageId="86" pageNumber="87">
<emphasis id="E192D004FF87613CFF526596FEAFF816" bold="true" box="[199,293,1994,2019]" pageId="86" pageNumber="87">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="D3590C16FF86613DFF5262CBFEB9FF20" blockId="87.[151,1436,151,285]" pageId="87" pageNumber="88">
<emphasis id="E192D004FF86613DFF5262CBFEB6FF45" bold="true" box="[199,316,151,176]" pageId="87" pageNumber="88">Variation.</emphasis>
Body length of examined specimens varied from 15.516.0 mm, otherwise the three specimens are rather similar.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF86613DFF526283FA8AFEE9" pageId="87" pageNumber="88" type="distribution">
<paragraph id="D3590C16FF86613DFF526283FA8AFEE9" blockId="87.[151,1436,151,285]" pageId="87" pageNumber="88">
<emphasis id="E192D004FF86613DFF526283FEEAFF0D" bold="true" box="[199,352,223,248]" pageId="87" pageNumber="88">Distribution.</emphasis>
This species is known from two rather distant localities in central
<collectingCountry id="ABF14C86FF86613DFB0C6283FAA8FF0D" box="[1177,1314,223,248]" name="Madagascar" pageId="87" pageNumber="88">Madagascar</collectingCountry>
, Ankafina Tsarafidy (as suggested by
<bibRefCitation id="B77771E7FF86613DFE526358FDFCFEE9" author="Paulian" box="[455,630,260,285]" pageId="87" pageNumber="88" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian (1977))</bibRefCitation>
and northern
<collectingCountry id="ABF14C86FF86613DFC876358FC13FEE8" box="[786,921,260,285]" name="Madagascar" pageId="87" pageNumber="88">Madagascar</collectingCountry>
(Amber Mountains) (
<figureCitation id="4BDD1093FF86613DFB1A6358FB69FEE9" box="[1167,1251,260,285]" captionStart="FIGURE 44" captionStartId="86.[151,250,1871,1893]" captionTargetBox="[185,1396,298,1847]" captionTargetId="figure@86.[184,1403,229,1850]" captionTargetPageId="86" captionText="FIGURE 44. Triodontus copridoides, holotype male (A, B, D, E), male (C). Habitus in dorsal view (A, C), labels (D), aedeagus in lateral view and parameres in dorsal and ventral view (D), abdomen (E), distributional records (F)." httpUri="https://zenodo.org/record/200487/files/figure.png" pageId="87" pageNumber="88">Fig. 44</figureCitation>
F).
</paragraph>
</subSubSection>
</treatment>
</document>