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<document id="299FB63D9402521B7AD715FBD2D2A834" ID-CLB-Dataset="39139" ID-DOI="10.11646/zootaxa.4107.4.2" ID-GBIF-Dataset="96189ebc-493c-44ef-9e46-1bf2af20c3e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="271360" ID-ZooBank="5CBAACB4-2A65-4766-8339-33D65C93FF9B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1462342958656" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2016" docId="597587E28F18FFE7FF6CFD65FCA32910" docLanguage="en" docName="zootaxa.4107.4.2.pdf.imf" docOrigin="Zootaxa 4107 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procloeon Bengtsson 1915" docType="treatment" docVersion="13" lastPageNumber="494" masterDocId="A54CFF9A8F1BFFE4FFFBFFB7FFEF2C2E" masterDocTitle="A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae)" masterLastPageNumber="516" masterPageNumber="491" pageNumber="494" updateTime="1698662244020" updateUser="ExternalLinkService">
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<mods:title id="C7ABAA8279A64871A9522CACFD8A1906">A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae)</mods:title>
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<mods:namePart id="65923B082D060F8F9B3F551C4D5F07EA">Kluge, Nikita J.</mods:namePart>
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<mods:date id="00C469DF54F58DA742DD471D397C01AA">2016</mods:date>
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<treatment id="597587E28F18FFE7FF6CFD65FCA32910" ID-DOI="http://doi.org/10.5281/zenodo.5670322" ID-GBIF-Taxon="120620936" ID-Zenodo-Dep="5670322" LSID="urn:lsid:plazi:treatment:597587E28F18FFE7FF6CFD65FCA32910" httpUri="http://treatment.plazi.org/id/597587E28F18FFE7FF6CFD65FCA32910" lastPageNumber="494" pageId="3" pageNumber="494">
<subSubSection id="99C6657F8F18FFE7FF6CFD65FDDD2EC2" box="[151,562,722,748]" pageId="3" pageNumber="494" type="nomenclature">
<paragraph id="D16336F48F18FFE7FF6CFD65FDDD2EC2" blockId="3.[151,562,722,748]" box="[151,562,722,748]" pageId="3" pageNumber="494">
<heading id="8A2B81988F18FFE7FF6CFD65FDDD2EC2" bold="true" box="[151,562,722,748]" fontSize="11" level="1" pageId="3" pageNumber="494" reason="1">
<emphasis id="E3A8EAE68F18FFE7FF6CFD65FDDD2EC2" bold="true" box="[151,562,722,748]" pageId="3" pageNumber="494">
Genus
<taxonomicName id="16DC4D778F18FFE7FF15FD65FDDD2EC2" ID-CoL="63PRK" ID-ENA="248186" authority="Bengtsson 1915" authorityName="Bengtsson" authorityYear="1915" box="[238,562,722,748]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F18FFE7FF15FD65FE852EC2" bold="true" box="[238,362,722,748]" italics="true" pageId="3" pageNumber="494">Procloeon</emphasis>
<bibRefCitation id="B54D4B058F18FFE7FE8AFD65FDDD2EC2" author="Bengtsson" box="[369,562,722,748]" pageId="3" pageNumber="494" refString="Bengtsson, S. (1915) Eine Namensanderung. Entomologisk Tidskrift, 36 (1), 34." type="journal article" year="1915">Bengtsson 1915</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="99C6657F8F18FFE7FF6CFCADFD732F42" pageId="3" pageNumber="494" type="reference_group">
<paragraph id="D16336F48F18FFE7FF6CFCADFD9E2F60" blockId="3.[151,846,793,876]" pageId="3" pageNumber="494">
<treatmentCitationGroup id="F1CC11DA8F18FFE7FF6CFCADFE0D2F01" box="[151,482,793,815]" pageId="3" pageNumber="494">
=
<taxonomicName id="16DC4D778F18FFE7FF50FCAEFE0D2F01" ID-CoL="6ZNG" authority="Bengtsson 1914" authorityName="Bengtsson" authorityYear="1914" box="[171,482,793,815]" class="Insecta" family="Baetidae" genus="Pseudocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F18FFE7FF50FCAEFED62F01" box="[171,313,793,815]" italics="true" pageId="3" pageNumber="494">Pseudocloeon</emphasis>
<treatmentCitation id="507D10E58F18FFE7FEBBFCADFE0D2F01" author="Bengtsson" box="[320,482,793,815]" pageId="3" pageNumber="494" year="1914">
<bibRefCitation id="B54D4B058F18FFE7FEBBFCADFE0D2F01" author="Bengtsson" box="[320,482,793,815]" pageId="3" pageNumber="494" refString="Bengtsson, S. (1914) Bemerkungen uber die nordischen Arten der Gattung Cloeon Leach. Entomologisk Tidskrift, 35 (3 - 4), 210 - 220." type="journal article" year="1914">Bengtsson 1914</bibRefCitation>
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
(non
<taxonomicName id="16DC4D778F18FFE7FDE6FCAEFCA82F01" ID-CoL="6ZNG" authority="Klapalek 1905" authorityName="Klapalek" authorityYear="1905" box="[541,839,793,815]" class="Insecta" family="Baetidae" genus="Pseudocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F18FFE7FDE6FCAEFD442F01" box="[541,683,793,815]" italics="true" pageId="3" pageNumber="494">Pseudocloeon</emphasis>
<bibRefCitation id="B54D4B058F18FFE7FD49FCAEFCA82F01" author="Klapalek" box="[690,839,793,815]" pageId="3" pageNumber="494" refString="Klapalek, F. (1905) Plecopteren und Ephemeriden aus Java, gesammelt von Prof. K. Kraepelin 1904. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 22, 103 - 107." type="journal article" year="1905">Klapalek 1905</bibRefCitation>
</taxonomicName>
)
<treatmentCitationGroup id="F1CC11DA8F18FFE7FF6CFC8FFD9E2F60" box="[151,625,824,846]" pageId="3" pageNumber="494">
Branch “11”:
<treatmentCitation id="507D10E58F18FFE7FED8FC8FFDCE2F60" author="Kluge &amp; Novikova" box="[291,545,824,846]" pageId="3" pageNumber="494" year="1992">
<bibRefCitation id="B54D4B058F18FFE7FED8FC8FFDCE2F60" author="Kluge" box="[291,545,824,846]" pageId="3" pageNumber="494" refString="Kluge, N. J. &amp; Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83. [in Russian, English translation: Entomological Review, 71 (9), 29 - 54] Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="1992">Kluge &amp; Novikova 1992</bibRefCitation>
</treatmentCitation>
:
<figureCitation id="49E72A718F18FFE7FDD0FC8FFD9E2F60" box="[555,625,824,846]" captionStart="FIGURES 11 20" captionStartId="7.[151,264,1612,1635]" captionTargetBox="[160,1415,207,1574]" captionTargetId="figure@7.[151,1436,193,1591]" captionTargetPageId="7" captionText="FIGURES 11 20. Procloeon (Oculogaster) cylindroculum, larva. 11 17, tergalii of I VII pairs (specimen from Zambia); 18, frontal suture; 19, claw; 20, fore leg, anterior view (fine setae not shown; dots show bases of fine setae forming regular rows on apical part of femur, proximal part of tibia and proximal part of tarsus)." httpUri="https://zenodo.org/record/271362/files/figure.png" pageId="3" pageNumber="494">Fig. 11</figureCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="D16336F48F18FFE7FF6CFCE1FD732F42" blockId="3.[151,846,793,876]" box="[151,668,854,876]" pageId="3" pageNumber="494">
Hierarchical name:
<taxonomicName id="16DC4D778F18FFE7FEA5FCE1FE222F42" ID-CoL="63PRK" box="[350,461,854,876]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
/g1:
<bibRefCitation id="B54D4B058F18FFE7FE08FCE1FD832F42" author="Kluge" box="[499,620,854,876]" pageId="3" pageNumber="494" refString="Kluge, N. J. (2011) Non-African representatives of the plesiomorphion Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376." type="journal article" year="2011">Kluge 2011</bibRefCitation>
: 362
</paragraph>
</subSubSection>
<subSubSection id="99C6657F8F18FFE7FF6CFC2EFD452F9C" box="[151,682,921,946]" pageId="3" pageNumber="494" type="materials_examined">
<paragraph id="D16336F48F18FFE7FF6CFC2EFD452F9C" blockId="3.[151,1436,921,1342]" box="[151,682,921,946]" pageId="3" pageNumber="494">
<emphasis id="E3A8EAE68F18FFE7FF6CFC2EFEDB2F9C" bold="true" box="[151,308,921,946]" pageId="3" pageNumber="494">
<typeStatus id="0E6788568F18FFE7FF6CFC2EFF3D2F9C" box="[151,210,921,946]" pageId="3" pageNumber="494">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="16DC4D778F18FFE7FEC0FC2EFD482F9C" authority="Bengtsson 1912" authorityName="Bengtsson" authorityYear="1912" box="[315,679,921,946]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="species" species="bifidum">
<emphasis id="E3A8EAE68F18FFE7FEC0FC2EFE062F9F" box="[315,489,921,945]" italics="true" pageId="3" pageNumber="494">Cloeon bifidum</emphasis>
<bibRefCitation id="B54D4B058F18FFE7FE14FC2DFD482F9C" author="Bengtsson" box="[495,679,922,946]" pageId="3" pageNumber="494" refString="Bengtsson, S. (1912) Neue Ephemeriden aus Schweden. Entomologisk Tidskrift, 33 (1 - 2), 107 - 117." type="journal article" year="1912">Bengtsson 1912</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="99C6657F8F18FFE7FF3CFC0AFB9A2880" pageId="3" pageNumber="494" type="diagnosis">
<paragraph id="D16336F48F18FFE7FF3CFC0AFF112848" blockId="3.[151,1436,921,1342]" pageId="3" pageNumber="494">
<emphasis id="E3A8EAE68F18FFE7FF3CFC0AFE7B2FFB" bold="true" box="[199,404,957,982]" pageId="3" pageNumber="494">
Diagnosis.
<emphasis id="E3A8EAE68F18FFE7FEB2FC09FE7B2FFB" bold="true" box="[329,404,958,981]" italics="true" pageId="3" pageNumber="494">Larva.</emphasis>
</emphasis>
On lateral side of distal part of cercus each segment bears one greatly enlarged spine (
<figureCitation id="49E72A718F18FFE7FA91FC0AFF592FD4" captionStart="FIGURES 21 29" captionStartId="8.[151,264,1932,1955]" captionTargetBox="[212,1367,193,1904]" captionTargetId="figure@8.[212,1367,193,1905]" captionTargetPageId="8" captionText="FIGURES 21 29. Procloeon (Oculogaster) cylindroculum, larvae from Zambia. 21, cercus and paracercus; 22, distal part of cercus; 23 25, abdominal terga and sterna; 26, right half of pronotum and mesonotum; 27, subimaginal exuviae of right half of mesonotum; 28, maxilla; 29, labrum." httpUri="https://zenodo.org/record/271363/files/figure.png" pageId="3" pageNumber="494">Figs 22</figureCitation>
,
<figureCitation id="49E72A718F18FFE7FF3AFC56FF312FD4" box="[193,222,993,1018]" captionStart="FIGURES 59 64" captionStartId="18.[151,264,1933,1956]" captionTargetBox="[187,1393,193,1915]" captionTargetId="figure@18.[187,1394,193,1916]" captionTargetPageId="18" captionText="FIGURES 59 64. Procloeon (Oculogaster) album sp. n., larva. 59, paracercus and cercus (with subimaginal cercus inside); 60, distal part of cercus; 61, posterior margin of abdominal tergum X; 62, fragment of abdominal tergum III; 63, fragment of abdominal sternum VI; 64, tergalius VI (arrow shows additional middle rib) (59, 60, holotype)." httpUri="https://zenodo.org/record/271369/files/figure.png" pageId="3" pageNumber="494">60</figureCitation>
) (unique autapomorphy, not found in any other taxa). Cerci and paracercus without long bare last segments (the same in
<taxonomicName id="16DC4D778F18FFE7FEC8FBB2FDFC2833" box="[307,531,1029,1053]" class="Insecta" family="Baetidae" genus="Pseudocentroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F18FFE7FEC8FBB2FDFC2833" box="[307,531,1029,1053]" italics="true" pageId="3" pageNumber="494">Pseudocentroptilum</emphasis>
</taxonomicName>
). In contrast to
<taxonomicName id="16DC4D778F18FFE7FD2EFBB2FCC92833" box="[725,806,1029,1053]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F18FFE7FD2EFBB2FCC92833" box="[725,806,1029,1053]" italics="true" pageId="3" pageNumber="494">Cloeon</emphasis>
</taxonomicName>
, additional dorsal lamella of tergalii without anteriorproximal projection; in some species dorsal lamella is lost. Labial palp with 3rd segment (truncate) always widened.
</paragraph>
<paragraph id="D16336F48F18FFE7FF3CFBC6FB9A2880" blockId="3.[151,1436,921,1342]" pageId="3" pageNumber="494">
<emphasis id="E3A8EAE68F18FFE7FF3CFBC6FE9E28A7" bold="true" box="[199,369,1137,1162]" italics="true" pageId="3" pageNumber="494">Winged stages.</emphasis>
In contrast to
<taxonomicName id="16DC4D778F18FFE7FDE8FBC6FD8B28A7" box="[531,612,1137,1161]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F18FFE7FDE8FBC6FD8B28A7" box="[531,612,1137,1161]" italics="true" pageId="3" pageNumber="494">Cloeon</emphasis>
</taxonomicName>
, unpaired sclerotized projection of penial bridge semicircular or truncate; fore wing of female not colored.
<collectingCountry id="A9CB76648F18FFE7FDF3FB22FDAF2880" box="[520,576,1173,1198]" name="India" pageId="3" pageNumber="494">Hind</collectingCountry>
wings present or absent. Oviparous or viviparous.
</paragraph>
</subSubSection>
<subSubSection id="99C6657F8F18FFE7FF3CFB0EFCA32910" pageId="3" pageNumber="494" type="distribution">
<paragraph id="D16336F48F18FFE7FF3CFB0EFC8B28FC" blockId="3.[151,1436,921,1342]" box="[199,868,1209,1234]" pageId="3" pageNumber="494">
<emphasis id="E3A8EAE68F18FFE7FF3CFB0EFE8F28FC" bold="true" box="[199,352,1209,1234]" pageId="3" pageNumber="494">Distribution.</emphasis>
Holarctic, Oriental and Afrotropical Regions.
</paragraph>
<paragraph id="D16336F48F18FFE7FF3CFB6AFCA32910" blockId="3.[151,1436,921,1342]" pageId="3" pageNumber="494">
<emphasis id="E3A8EAE68F18FFE7FF3CFB6AFE8828D8" bold="true" box="[199,359,1245,1270]" pageId="3" pageNumber="494">Composition.</emphasis>
The genus
<taxonomicName id="16DC4D778F18FFE7FDF8FB6AFD9828DB" box="[515,631,1245,1269]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F18FFE7FDF8FB6AFD9828DB" box="[515,631,1245,1269]" italics="true" pageId="3" pageNumber="494">Procloeon</emphasis>
</taxonomicName>
сan be divided into several subgenera, among which the largest nominative subgenus
<taxonomicName id="16DC4D778F18FFE7FE74FAB6FDEC2937" box="[399,515,1281,1305]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="E3A8EAE68F18FFE7FE74FAB6FDEC2937" box="[399,515,1281,1305]" italics="true" pageId="3" pageNumber="494">Procloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F89B579D8F18FFE7FDF7FAB5FDAF2937" box="[524,576,1282,1305]" pageId="3" pageNumber="494" sensu="stricto">s.str.</taxonomicNameLabel>
is a plesiomorphon ancestral to subgenera
<taxonomicName id="16DC4D778F18FFE7FBCAFAB6FE2D2910" authority="Jacob (in Jacob &amp; Glazaczow)" authorityName="Jacob (in Jacob &amp; Glazaczow" authorityYear="1987" class="Insecta" family="Baetidae" genus="Pseudocentroptiloides" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="494" phylum="Arthropoda" rank="genus">
<emphasis id="E3A8EAE68F18FFE7FBCAFAB6FAC52937" box="[1073,1322,1281,1305]" italics="true" pageId="3" pageNumber="494">Pseudocentroptiloides</emphasis>
Jacob (in
<bibRefCitation id="B54D4B058F18FFE7FF6CFA92FE2D2910" author="Jacob" box="[151,450,1317,1342]" pageId="3" pageNumber="494" refString="Jacob, U. &amp; Glazaczow, A. (1987) Pseudocentroptiloides, a new baetid genus of Palaearctic and Oriental distribution (Ephemeroptera). Aquatic Insects, 8 (4), 197 - 206." type="journal article" year="1987">Jacob &amp; Glazaczow) 1987</bibRefCitation>
</taxonomicName>
,
<emphasis id="E3A8EAE68F18FFE7FE36FA92FDB82913" box="[461,599,1317,1341]" italics="true" pageId="3" pageNumber="494">Oculogaster</emphasis>
sugen. n., and others.
</paragraph>
</subSubSection>
</treatment>
</document>