1181 lines
137 KiB
XML
1181 lines
137 KiB
XML
<document id="37D97FBC929134C16329DD1ABD48A8FB" ID-CLB-Dataset="4278" ID-DOI="10.18590/euscorpius.2015.vol2015.iss194.1" ID-GBIF-Dataset="4230f888-1e02-4af9-af99-7ac1a909a647" ID-ISSN="1536-9307" ID-Zenodo-Dep="5508064" ID-ZooBank="B5D24928-5D29-40D8-8448-4953099A75EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617134406945" checkinUser="carolina" docAuthor="Ayrey, Richard F. & Soleglad, Michael E." docDate="2015" docId="595B87DEFFAFC26DFF28FE36FEE4BCAF" docLanguage="en" docName="Euscorpius.194.1-12.pdf" docOrigin="Euscorpius 194" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Vaejovis troupi Ayrey et Soleglad 2015, sp. n." docType="treatment" docUuid="8B5C2249-188E-4AAC-A26F-956F272DAEED" docUuidSource="ZooBank" docVersion="7" lastPageNumber="12" masterDocId="A562FFA6FFA8C260FFD3FFA4FFC4BF54" masterDocTitle="New species of Vaejovis from the Whetstone Mountains, southern Arizona (Scorpiones: Vaejovidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="6" updateTime="1698935388094" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods id="F9AE25AED0BB704BDE4467BF847B2DC2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="08AED081FE48E0D195E04FDF92CDA59C">
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<mods:title id="1AF3F354390DF4F3F1307DF65C348C6C">New species of Vaejovis from the Whetstone Mountains, southern Arizona (Scorpiones: Vaejovidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="CCE7B132DCCFD7C2B8A967BB95DBD09B" type="personal">
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<mods:roleTerm id="F4FADB614D79FD2CEE4ABD625745B28A">Author</mods:roleTerm>
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<mods:namePart id="F076592F01369DF5368FCE20DF2D5A05">Ayrey, Richard F.</mods:namePart>
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<mods:name id="D28AB5E6417261C95E95BBDCFEA636E2" type="personal">
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<mods:role id="44058D59884D9630A2B9EF8A467C46B8">
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<mods:roleTerm id="05E0C8FB1537E6BEE10DC232109829D3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D153DFDE9DDBDF9D8D1B5C7F1B4B3B41">Soleglad, Michael E.</mods:namePart>
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<mods:typeOfResource id="7A14ACF9B9E39639ED8F45F86DFEFB90">text</mods:typeOfResource>
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<mods:title id="65E2A27BE3B2A483055FEF48783286D3">Euscorpius</mods:title>
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</mods:titleInfo>
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<mods:part id="FF88549930157ADC9B412A878F6A32A8">
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<mods:date id="1584E9B94D57F9351D456AC46EFE2E0B">2015</mods:date>
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<mods:number id="117473355D03548C91A623D0ECFC9B6D">2015-12-31</mods:number>
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<mods:detail id="80E93085AFE33732FC4412A5F09B26E2" type="volume">
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<mods:number id="35558EB31043953CD02B3193690034EC">194</mods:number>
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<mods:classification id="C7BCA014817970BA333A4E3CACFAFEE2">journal article</mods:classification>
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<mods:identifier id="BCD55BBDE2CB7986EB1D58632942DEA8" type="DOI">10.18590/euscorpius.2015.vol2015.iss194.1</mods:identifier>
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<treatment id="595B87DEFFAFC26DFF28FE36FEE4BCAF" ID-DOI="http://doi.org/10.5281/zenodo.5508066" ID-GBIF-Taxon="187975310" ID-Zenodo-Dep="5508066" LSID="urn:lsid:zoobank.org:act:8B5C2249-188E-4AAC-A26F-956F272DAEED" httpUri="http://treatment.plazi.org/id/595B87DEFFAFC26DFF28FE36FEE4BCAF" lastPageId="13" lastPageNumber="12" pageId="7" pageNumber="6">
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<subSubSection id="99E86543FFAFC267FF28FE36FD23BEF8" box="[251,743,402,428]" pageId="7" pageNumber="6" type="nomenclature">
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<paragraph id="D14D36C8FFAFC267FF28FE36FD23BEF8" blockId="7.[195,798,402,520]" box="[251,743,402,428]" pageId="7" pageNumber="6">
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<heading id="8A0581A4FFAFC267FF28FE36FD23BEF8" box="[251,743,402,428]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="6" reason="2">
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<emphasis id="E386EADAFFAFC267FF28FE36FD23BEF8" bold="true" box="[251,743,402,428]" pageId="7" pageNumber="6">
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<taxonomicName id="16F24D4BFFAFC267FF28FE36FD52BEF8" ID-CoL="7F92D" authority="Ayrey et Soleglad" authorityName="Ayrey et Soleglad" authorityYear="2015" box="[251,662,402,428]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="troupi" status="sp. nov.">
|
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<emphasis id="E386EADAFFAFC267FF28FE36FE76BEF8" bold="true" box="[251,434,402,428]" italics="true" pageId="7" pageNumber="6">Vaejovis troupi</emphasis>
|
||
Ayrey et Soleglad
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F8B557A1FFAFC267FD76FE36FD23BEF8" box="[677,743,402,428]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99E86543FFAFC267FEBAFE16FD09BD5C" pageId="7" pageNumber="6" type="description">
|
||
<paragraph id="D14D36C8FFAFC267FEBAFE16FDBCBE9E" blockId="7.[195,798,402,520]" box="[361,632,434,458]" pageId="7" pageNumber="6">
|
||
<figureCitation id="49C92A4DFFAFC267FEBAFE16FE0ABE9E" box="[361,462,434,458]" captionStart-0="Figure 1" captionStart-1="Figures 2-6" captionStart-2="Figure 7" captionStartId-0="3.[192,267,1888,1912]" captionStartId-1="4.[192,278,1496,1520]" captionStartId-2="6.[192,267,1876,1900]" captionTargetBox-0="[191,1441,191,1857]" captionTargetBox-1="[195,1394,180,1459]" captionTargetBox-2="[202,1381,199,1841]" captionTargetId-0="figure-26@3.[192,1440,192,1856]" captionTargetId-1="figure-204@4.[192,1437,168,1467]" captionTargetId-2="figure-24@6.[192,1438,192,1847]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="Figure 1: Vaejovis troupi sp. n., female holotype. In natural habitat (top) and on a leaf (bottom)." captionText-1="Figures 2-6: Vaejovis troupi sp. n., female holotype. 2. Metasoma and telson, ventral and lateral views. 3. Carapace and closeup of lateral eyes. 4. Pedipalp chela, external view, patella and femur, dorsal views. 5. Right leg III basitarsus and tarsus, ventral view. 6. Chelal fixed and movable fingers, each exhibiting six inner denticles (ID)." captionText-2="Figure 7: Vaejovis troupi sp. n., female holotype. Trichobothrial pattern." figureDoi-0="http://doi.org/10.5281/zenodo.5508068" figureDoi-1="http://doi.org/10.5281/zenodo.5508070" figureDoi-2="http://doi.org/10.5281/zenodo.5508072" httpUri-0="https://zenodo.org/record/5508068/files/figure.png" httpUri-1="https://zenodo.org/record/5508070/files/figure.png" httpUri-2="https://zenodo.org/record/5508072/files/figure.png" pageId="7" pageNumber="6">Figs. 1–7</figureCitation>
|
||
,
|
||
<figureCitation id="49C92A4DFFAFC267FE09FE16FE33BE9E" box="[474,503,434,458]" captionStart="Figure 10" captionStartId="10.[192,267,1648,1672]" captionTargetBox="[191,1441,220,1609]" captionTargetId="figure-155@10.[192,1440,221,1608]" captionTargetPageId="10" captionText="Figure 10: Proposed geographic and molecular relationships of “sky island” species Vaejovis troupi sp. n., V. grahami, and V. vorhiesi. Top. Close-up of southeastern Arizona showing the type localities of the three species and the distinct, isolated “sky island” mountain ranges in which they occur, Whetstone, Santa Rita, and Huachuca Mountains, respectively. Bottom. A partial chronogram based on molecular data showing proposed phylogenetic relationships based on evolutionary time (red numbers). Red numbers above branch indicate estimates for a multilocus species tree, bottom numbers represent a mtDNA gene tree. Means and 95% highest posterior densities are shown for each node. Red numbers indicate proposed minimum, maximum, and mean Mya. Information is from Bryson et al. (2013: fig. S2). Gray rectangle delineates the three species discussed in this paper. Altitude data is that of the type localities." figureDoi="http://doi.org/10.5281/zenodo.5508078" httpUri="https://zenodo.org/record/5508078/files/figure.png" pageId="7" pageNumber="6">10</figureCitation>
|
||
;
|
||
<tableCitation id="9C700373FFAFC267FDD0FE16FDBCBE9E" box="[515,632,434,458]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="5.[192,257,1337,1361]" captionStartId-1="12.[192,257,524,548]" captionTargetId-0="graphics-8@5.[255,1858,192,1307]" captionTargetId-1="graphics-13@12.[187,1445,192,495]" captionTargetPageId-0="5" captionTargetPageId-1="12" captionText-0="Table 1: Morphometrics (mm) of Vaejovis troupi sp. n., Whetstone Mountains, Cochise County, Arizona, USA, V. grahami, Madera County, Arizona, USA (after Ayrey & Soleglad, 2014: tab.1, in part), and V. vorhiesi lectotype, Miller’s Canyon, Huachuca Mountains, Graham, 2007: 3–4, in part)." captionText-1="Table 2: Comparison of morphometric ratios of the metasoma based on the measurements of female specimens presented in Table 1. In these ratios, Vaejovis troupi sp. n. exhibits the more slender metasoma when compared to the other two species (as indicated by a larger ratio value), ranging from 8 to over 16 percent mean value difference (MVD). Statistical data group: minimum–maximum (mean) (±standard deviation) [number of samples]. Seg = metasomal segment, L = length, W = width." pageId="7" pageNumber="6">Tables 1–2</tableCitation>
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FF10FE76FD09BD5C" blockId="7.[195,798,402,520]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FF10FE76FD09BD5C" bold="true" pageId="7" pageNumber="6">http://zoobank.org/urn:lsid:zoobank.org:act:8B5C22 49-188E-4AAC-A26F-956F272DAEED</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99E86543FFAFC267FF13FD90FDEDBDF3" pageId="7" pageNumber="6" type="reference_group">
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||
<paragraph id="D14D36C8FFAFC267FF13FD90FE81BD18" blockId="7.[192,802,564,679]" box="[192,325,564,588]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FF13FD90FE81BD18" bold="true" box="[192,325,564,588]" pageId="7" pageNumber="6">References:</emphasis>
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FF23FDF6FDEDBDF3" blockId="7.[192,802,564,679]" pageId="7" pageNumber="6">
|
||
<taxonomicName id="16F24D4BFFAFC267FF23FDF6FD03BD3D" ID-CoL="85MN" authority=": Ayrey, 2013 a: 3" authorityName="Thorell" authorityPageNumber="3" authorityYear="1876" box="[240,711,593,617]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E386EADAFFAFC267FF23FDF6FE88BD3D" box="[240,332,594,617]" italics="true" pageId="7" pageNumber="6">Vaejovis</emphasis>
|
||
sp. cf.
|
||
<emphasis id="E386EADAFFAFC267FE74FDF5FE39BD3D" box="[423,509,593,617]" italics="true" pageId="7" pageNumber="6">vorhiesi</emphasis>
|
||
: Ayrey, 2013a: 3
|
||
</taxonomicName>
|
||
; Ayrey, 2013b: 2; Ayrey, 2014: 3;
|
||
<taxonomicName id="16F24D4BFFAFC267FDFCFDD5FEF6BDF3" ID-CoL="85MN" authorityName="Thorell" authorityYear="1876" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E386EADAFFAFC267FDFCFDD5FD4FBDDC" box="[559,651,625,648]" italics="true" pageId="7" pageNumber="6">Vaejovis</emphasis>
|
||
sp.,
|
||
<emphasis id="E386EADAFFAFC267FD1FFDD4FCE6BDDC" box="[716,802,624,648]" italics="true" pageId="7" pageNumber="6">vorhiesi</emphasis>
|
||
group
|
||
</taxonomicName>
|
||
: Bryson et al., 2013: 5.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99E86543FFAFC267FF13FD77FEBEBC95" pageId="7" pageNumber="6" type="diagnosis">
|
||
<paragraph id="D14D36C8FFAFC267FF13FD77FEBEBC95" blockId="7.[192,802,722,961]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FF13FD77FEF3BDBF" bold="true" box="[192,311,723,747]" pageId="7" pageNumber="6">Diagnosis:</emphasis>
|
||
Medium sized species,
|
||
<quantity id="160A9B2DFFAFC267FD8BFD76FD6FBDBE" box="[600,683,722,746]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="6" unit="mm" value="26.0">26 mm</quantity>
|
||
in length (only female known); light brown in color with variegated patterns on the carapace, lighter mottlings on the mesosoma, pedipalps, and legs. Pectinal tooth counts 11–12. Chelal fingers with six inner denticles (
|
||
<emphasis id="E386EADAFFAFC267FD05FCEAFD36BC31" box="[726,754,846,869]" italics="true" pageId="7" pageNumber="6">ID</emphasis>
|
||
) on both the movable and fixed fingers. Patellar external trichobothria
|
||
<emphasis id="E386EADAFFAFC267FE89FC2FFE04BCF6" box="[346,448,907,930]" italics="true" pageId="7" pageNumber="6">
|
||
est–et
|
||
<subScript id="4D76348DFFAFC267FE44FC30FE64BCF6" attach="left" box="[407,416,916,930]" fontSize="6" pageId="7" pageNumber="6">3</subScript>
|
||
–v
|
||
<subScript id="4D76348DFFAFC267FE64FC30FE04BCF6" attach="left" box="[439,448,916,930]" fontSize="6" pageId="7" pageNumber="6">3</subScript>
|
||
</emphasis>
|
||
form an angle,
|
||
<emphasis id="E386EADAFFAFC267FD5CFC2FFD6FBCF6" box="[655,683,907,930]" italics="true" pageId="7" pageNumber="6">
|
||
et
|
||
<subScript id="4D76348DFFAFC267FD71FC30FD6FBCF6" attach="left" box="[674,683,916,930]" fontSize="6" pageId="7" pageNumber="6">3</subScript>
|
||
</emphasis>
|
||
occurring more proximally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99E86543FFAFC267FF13FC42FD8EBB49" pageId="7" pageNumber="6" type="etymology">
|
||
<paragraph id="D14D36C8FFAFC267FF13FC42FD8EBB49" blockId="7.[192,802,998,1053]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FF13FC42FE86BCAA" bold="true" box="[192,322,998,1022]" pageId="7" pageNumber="6">Etymology.</emphasis>
|
||
This species is named in honor of Robert Troup who found the only specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99E86543FFAFC267FF13FBE7FD12BB2D" pageId="7" pageNumber="6" type="distribution">
|
||
<paragraph id="D14D36C8FFAFC267FF13FBE7FD12BB2D" blockId="7.[192,801,1090,1145]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FF13FBE7FE96BB0F" bold="true" box="[192,338,1091,1115]" pageId="7" pageNumber="6">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="0E49886AFFAFC267FD99FBE7FDBCBB0F" box="[586,632,1091,1115]" pageId="7" pageNumber="6">type</typeStatus>
|
||
locality, Whetstone Mountains, Cochise County,
|
||
<collectingRegion id="1336F82AFFAFC267FDE5FBC5FD4ABB2D" box="[566,654,1121,1145]" country="United States of America" name="Arizona" pageId="7" pageNumber="6">Arizona</collectingRegion>
|
||
,
|
||
<collectingCountry id="A9E57658FFAFC267FD49FBC5FD17BB2D" box="[666,723,1121,1145]" name="United States of America" pageId="7" pageNumber="6">USA</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99E86543FFAFC26DFF13FB3BFEE4BCAF" lastPageId="13" lastPageNumber="12" pageId="7" pageNumber="6" type="materials_examined">
|
||
<paragraph id="D14D36C8FFAFC267FF13FB3BFDC0BA46" blockId="7.[192,802,1182,1298]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FF13FB3BFEACBBE3" bold="true" box="[192,360,1183,1207]" pageId="7" pageNumber="6">Type material.</emphasis>
|
||
<materialsCitation id="619A3C95FFAFC267FEBCFB3AFDC0BA46" ID-GBIF-Occurrence="3352142301" collectingDate="2009-10-14" collectionCode="R, USNM" country="United States of America" county="Cochise County" elevation="1607" latitude="31.81093" location="Whetstone Mountains" longLatPrecision="1" longitude="-110.3937" pageId="7" pageNumber="6" specimenCount="1" stateProvince="Arizona" typeStatus="holotype">
|
||
<typeStatus id="0E49886AFFAFC267FEBCFB3AFE16BBE2" box="[367,466,1182,1206]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
|
||
female,
|
||
<location id="D42D6013FFAFC267FDFCFB3AFCD9BBE2" LSID="urn:lsid:plazi:treatment:595B87DEFFAFC26DFF28FE36FEE4BCAF:D42D6013FFAFC267FDFCFB3AFCD9BBE2" box="[559,797,1182,1206]" country="United States of America" county="Cochise County" latitude="31.81093" longLatPrecision="1" longitude="-110.3937" name="Whetstone Mountains" pageId="7" pageNumber="6" stateProvince="Arizona">Whetstone Mountains</location>
|
||
,
|
||
<collectingCounty id="382C4E44FFAFC267FF13FB19FEB4BB81" box="[192,368,1213,1237]" pageId="7" pageNumber="6">Cochise County</collectingCounty>
|
||
,
|
||
<collectingRegion id="1336F82AFFAFC267FEADFB19FE12BB81" box="[382,470,1213,1237]" country="United States of America" name="Arizona" pageId="7" pageNumber="6">Arizona</collectingRegion>
|
||
,
|
||
<collectingCountry id="A9E57658FFAFC267FE37FB19FDDDBB81" box="[484,537,1213,1237]" name="United States of America" pageId="7" pageNumber="6">USA</collectingCountry>
|
||
(
|
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<geoCoordinate id="B4C6500FFFAFC267FDF9FB19FD57BB81" box="[554,659,1213,1237]" degrees="31.81093" direction="north" orientation="latitude" pageId="7" pageNumber="6" precision="1" value="31.81093">31.81093</geoCoordinate>
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,
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<geoCoordinate id="B4C6500FFFAFC267FD73FB1AFCDABB81" box="[672,798,1213,1237]" degrees="110.39370" direction="west" orientation="longitude" pageId="7" pageNumber="6" precision="1" value="-110.3937">-110.39370</geoCoordinate>
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;
|
||
<elevation id="5ADFD1FBFFAFC267FF13FB78FEFFBBA0" box="[192,315,1244,1268]" metricMagnitude="3" metricUnit="m" metricValue="1.607" pageId="7" pageNumber="6" unit="m" value="1607.0">
|
||
<quantity id="160A9B2DFFAFC267FF13FB78FED7BBA7" box="[192,275,1244,1268]" metricMagnitude="3" metricUnit="m" metricValue="1.607" pageId="7" pageNumber="6" unit="m" value="1607.0">1607 m</quantity>
|
||
asl
|
||
</elevation>
|
||
),
|
||
<date id="A54C1008FFAFC267FE81FB78FDC9BBA0" box="[338,525,1244,1268]" pageId="7" pageNumber="6" value="2009-10-14">
|
||
<collectingDate id="B508E9E0FFAFC267FE81FB78FDC9BBA0" box="[338,525,1244,1268]" pageId="7" pageNumber="6" value="2009-10-14">14 October 2009</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectionCode id="B7E3AE0DFFAFC267FD99FB78FD9BBBA7" box="[586,607,1244,1267]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="6">R</collectionCode>
|
||
. Troup, specimen #RA223, deposited in
|
||
<collectionCode id="B7E3AE0DFFAFC267FE63FB5EFDC4BA46" box="[432,512,1274,1298]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="6" type="Museum">USNM</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FF13FA9CFE19BA3A" blockId="7.[192,801,1335,1390]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FF13FA9CFEFDBA04" bold="true" box="[192,313,1336,1360]" pageId="7" pageNumber="6">FEMALE.</emphasis>
|
||
Description based on
|
||
<typeStatus id="0E49886AFFAFC267FDEBFA93FD51BA1B" box="[568,661,1335,1359]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
female. See
|
||
<tableCitation id="9C700373FFAFC267FF13FAF2FED5BA3A" box="[192,273,1366,1390]" captionStart="Table 1" captionStartId="5.[192,257,1337,1361]" captionTargetId="graphics-8@5.[255,1858,192,1307]" captionTargetPageId="5" captionText="Table 1: Morphometrics (mm) of Vaejovis troupi sp. n., Whetstone Mountains, Cochise County, Arizona, USA, V. grahami, Madera County, Arizona, USA (after Ayrey & Soleglad, 2014: tab.1, in part), and V. vorhiesi lectotype, Miller’s Canyon, Huachuca Mountains, Graham, 2007: 3–4, in part)." pageId="7" pageNumber="6">Table 1</tableCitation>
|
||
for measurements.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FF13FA30FF36B953" blockId="7.[192,802,1428,1543]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FF13FA30FE44BAF8" bold="true" box="[192,384,1428,1452]" pageId="7" pageNumber="6">COLORATION.</emphasis>
|
||
Color is medium brown, slightly lighter on the legs, telson orange. Weak underlying mottling is visible on the carapace, mesosoma, pedipalps, and legs.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FF13F989FE4BB9A9" blockId="7.[192,802,1581,1790]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FF13F989FE72B911" bold="true" box="[192,438,1581,1605]" pageId="7" pageNumber="6">
|
||
CARAPACE (
|
||
<figureCitation id="49C92A4DFFAFC267FEB7F989FE63B911" box="[356,423,1581,1605]" captionStart="Figures 2-6" captionStartId="4.[192,278,1496,1520]" captionTargetBox="[195,1394,180,1459]" captionTargetId="figure-204@4.[192,1437,168,1467]" captionTargetPageId="4" captionText="Figures 2-6: Vaejovis troupi sp. n., female holotype. 2. Metasoma and telson, ventral and lateral views. 3. Carapace and closeup of lateral eyes. 4. Pedipalp chela, external view, patella and femur, dorsal views. 5. Right leg III basitarsus and tarsus, ventral view. 6. Chelal fixed and movable fingers, each exhibiting six inner denticles (ID)." figureDoi="http://doi.org/10.5281/zenodo.5508070" httpUri="https://zenodo.org/record/5508070/files/figure.png" pageId="7" pageNumber="6">Fig. 3</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Anterior margin of carapace with a conspicuous wide emargination. Carapace moderately granular occurring primarily in the subtle mottled areas. Three lateral eyes on each side, the most proximal the smallest. Ratio of median eyes position from anterior edge/carapace length 0.340; carapace length/width at median eyes 1.442.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FF13F886FC55BE6D" blockId="7.[192,802,1826,1911]" lastBlockId="7.[831,1440,198,313]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FF13F886FEA2B86E" bold="true" box="[192,358,1826,1850]" pageId="7" pageNumber="6">MESOSOMA.</emphasis>
|
||
Tergites moderately granular on proximal half on segments I–
|
||
<collectionCode id="B7E3AE0DFFAFC267FE37F8E5FDC7B80C" box="[484,515,1857,1880]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="6">VI</collectionCode>
|
||
. Tergite VII with strong crenulated median and lateral carina. Sternites III–
|
||
<collectionCode id="B7E3AE0DFFAFC267FCD5F8C4FCE6B823" box="[774,802,1888,1911]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="6">VI</collectionCode>
|
||
smooth. Sternite VII with rough surface with weak ventral lateral carinae. Stigma small and ovoid with median side rotated 35 degrees from posterior sternite margin.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FCECFEFBFAADBEC2" blockId="7.[831,1439,351,406]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FCECFEFBFC0FBE23" bold="true" box="[831,971,351,375]" pageId="7" pageNumber="6">STERNUM.</emphasis>
|
||
Sternum conforms to
|
||
<typeStatus id="0E49886AFFAFC267FB16FEC4FB37BE2C" box="[1221,1267,352,376]" pageId="7" pageNumber="6">type</typeStatus>
|
||
2, lateral lobes and apex subtly defined. Sclerite is wider than long.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FCECFE18FC2CBEA5" blockId="7.[831,1441,443,497]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FCECFE18FBB3BE80" bold="true" box="[831,1143,444,468]" pageId="7" pageNumber="6">GENITAL OPERCULUM.</emphasis>
|
||
Sclerites separated on posterior one-third.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FCECFDB3FBA3BD38" blockId="7.[831,1440,535,620]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FCECFDB3FC0FBD7B" bold="true" box="[831,971,535,559]" pageId="7" pageNumber="6">PECTINES.</emphasis>
|
||
With three anterior lamellae, 8/8 middle lamellae, and 11/12 teeth. Sensorial areas present on all teeth and fulcra are present.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FCECFD36FA58BB4E" blockId="7.[831,1441,658,1050]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FCECFD36FB97BDFE" bold="true" box="[831,1107,658,682]" pageId="7" pageNumber="6">
|
||
METASOMA (
|
||
<figureCitation id="49C92A4DFFAFC267FC24FD36FB80BDFE" box="[1015,1092,658,682]" captionStart="Figures 2-6" captionStartId="4.[192,278,1496,1520]" captionTargetBox="[195,1394,180,1459]" captionTargetId="figure-204@4.[192,1437,168,1467]" captionTargetPageId="4" captionText="Figures 2-6: Vaejovis troupi sp. n., female holotype. 2. Metasoma and telson, ventral and lateral views. 3. Carapace and closeup of lateral eyes. 4. Pedipalp chela, external view, patella and femur, dorsal views. 5. Right leg III basitarsus and tarsus, ventral view. 6. Chelal fixed and movable fingers, each exhibiting six inner denticles (ID)." figureDoi="http://doi.org/10.5281/zenodo.5508070" httpUri="https://zenodo.org/record/5508070/files/figure.png" pageId="7" pageNumber="6">Fig. 2</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Segments I–IV: dorsal and dorsolateral carinae strong and serrate with distal denticle of I–III enlarged and spinoid, and spinoid on the dorsal carinae and expanded and flared on the dorsolateral carinae for segment IV. Lateral carinae strong and serrate on I, present on posterior 2/3 of II, posterior 1/3 of III, and obsolete on IV. Ventrolateral carinae strong and serrate, ventromedian moderately granular on I and crenulate to serrate on II–IV. Segment
|
||
<collectionCode id="B7E3AE0DFFAFC267FB24FC2CFACABCCB" box="[1271,1294,904,927]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="6" type="Museum">V</collectionCode>
|
||
: Dorsolateral carinae strong and irregularly granulate. Lateral carinae granular on basal 3/5. Ventrolateral and ventromedian carinae serrate. Anal arch with approximately 16 small denticles. Dorsal intercarinal spaces irregularly granular.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FCECFB98FB66BBE4" blockId="7.[831,1440,1084,1200]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FCECFB98FBC8BB00" bold="true" box="[831,1036,1084,1108]" pageId="7" pageNumber="6">
|
||
TELSON (
|
||
<figureCitation id="49C92A4DFFAFC267FC69FB98FC39BB00" box="[954,1021,1084,1108]" captionStart="Figures 2-6" captionStartId="4.[192,278,1496,1520]" captionTargetBox="[195,1394,180,1459]" captionTargetId="figure-204@4.[192,1437,168,1467]" captionTargetPageId="4" captionText="Figures 2-6: Vaejovis troupi sp. n., female holotype. 2. Metasoma and telson, ventral and lateral views. 3. Carapace and closeup of lateral eyes. 4. Pedipalp chela, external view, patella and femur, dorsal views. 5. Right leg III basitarsus and tarsus, ventral view. 6. Chelal fixed and movable fingers, each exhibiting six inner denticles (ID)." figureDoi="http://doi.org/10.5281/zenodo.5508070" httpUri="https://zenodo.org/record/5508070/files/figure.png" pageId="7" pageNumber="6">Fig. 2</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Smooth with several setae on ventral surface. Vesicular tabs with one to two small pointed granules. Subaculear tubercle present but small. LAS present exhibiting 5–6 serrations.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC267FCECFB72FA99BA7F" blockId="7.[831,1440,1237,1323]" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FCECFB72FC3DBBBA" bold="true" box="[831,1017,1238,1262]" pageId="7" pageNumber="6">CHELICERAE.</emphasis>
|
||
Dorsal edge of movable cheliceral finger with two subdistal (
|
||
<emphasis id="E386EADAFFAFC267FBB1FB50FBBEBA58" box="[1122,1146,1268,1292]" italics="true" pageId="7" pageNumber="6">sd</emphasis>
|
||
) denticles. Ventral edge is smooth, with well-developed serrula on distal half.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFAFC268FCECFAF4FE99B96B" blockId="7.[831,1441,1360,1910]" lastBlockId="8.[192,803,1152,1599]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
|
||
<emphasis id="E386EADAFFAFC267FCECFAF4FB14BA3C" bold="true" box="[831,1232,1360,1384]" pageId="7" pageNumber="6">
|
||
PEDIPALP (
|
||
<figureCitation id="49C92A4DFFAFC267FC04FAF4FBA7BA3C" box="[983,1123,1360,1384]" captionStart-0="Figures 2-6" captionStart-1="Figure 7" captionStartId-0="4.[192,278,1496,1520]" captionStartId-1="6.[192,267,1876,1900]" captionTargetBox-0="[195,1394,180,1459]" captionTargetBox-1="[202,1381,199,1841]" captionTargetId-0="figure-204@4.[192,1437,168,1467]" captionTargetId-1="figure-24@6.[192,1438,192,1847]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="Figures 2-6: Vaejovis troupi sp. n., female holotype. 2. Metasoma and telson, ventral and lateral views. 3. Carapace and closeup of lateral eyes. 4. Pedipalp chela, external view, patella and femur, dorsal views. 5. Right leg III basitarsus and tarsus, ventral view. 6. Chelal fixed and movable fingers, each exhibiting six inner denticles (ID)." captionText-1="Figure 7: Vaejovis troupi sp. n., female holotype. Trichobothrial pattern." figureDoi-0="http://doi.org/10.5281/zenodo.5508070" figureDoi-1="http://doi.org/10.5281/zenodo.5508072" httpUri-0="https://zenodo.org/record/5508070/files/figure.png" httpUri-1="https://zenodo.org/record/5508072/files/figure.png" pageId="7" pageNumber="6">Figs. 4, 6–7</figureCitation>
|
||
). Femur.
|
||
</emphasis>
|
||
Dorsointernal and dorsoexternal carinae serrated, and ventrointernal crenulated, ventroexternal rounded. Dorsal and ventral surfaces very rough, internal surface with scattered granules, and external with line of serrated granules.
|
||
<emphasis id="E386EADAFFAFC267FCECFA4EFC50B956" bold="true" box="[831,916,1514,1538]" pageId="7" pageNumber="6">Patella.</emphasis>
|
||
Dorsointernal and ventrointernal carinae crenulated, dorsoexternal and ventroexternal carinae granulated. Dorsal patellar (
|
||
<emphasis id="E386EADAFFAFC267FBE9F983FBAFB96B" box="[1082,1131,1575,1599]" italics="true" pageId="7" pageNumber="6">
|
||
<collectionCode id="B7E3AE0DFFAFC267FBE9F983FBAFB96B" box="[1082,1131,1575,1599]" pageId="7" pageNumber="6">DPS</collectionCode>
|
||
</emphasis>
|
||
) and ventral patellar (
|
||
<emphasis id="E386EADAFFAFC267FABAF983FA53B96B" box="[1385,1431,1575,1599]" italics="true" pageId="7" pageNumber="6">
|
||
<collectionCode id="B7E3AE0DFFAFC267FABAF983FA53B96B" box="[1385,1431,1575,1599]" pageId="7" pageNumber="6">VPS</collectionCode>
|
||
</emphasis>
|
||
) spurs formed with a pointed granule,
|
||
<emphasis id="E386EADAFFAFC267FB36F9E1FADAB937" box="[1253,1310,1605,1635]" italics="true" pageId="7" pageNumber="6">
|
||
<collectionCode id="B7E3AE0DFFAFC267FB36F9E1FAD2B909" box="[1253,1302,1605,1629]" pageId="7" pageNumber="6">DPS</collectionCode>
|
||
<subScript id="4D76348DFFAFC267FAC5F9F1FADAB937" attach="left" box="[1302,1310,1621,1635]" fontSize="6" pageId="7" pageNumber="6">c</subScript>
|
||
</emphasis>
|
||
carina well developed with approximately 14 serrated granules.
|
||
<emphasis id="E386EADAFFAFC267FCECF92DFC4CB9F5" bold="true" box="[831,904,1673,1697]" pageId="7" pageNumber="6">Chela.</emphasis>
|
||
Digital (
|
||
<emphasis id="E386EADAFFAFC267FC3DF92CFBCAB9F4" box="[1006,1038,1672,1696]" italics="true" pageId="7" pageNumber="6">D1</emphasis>
|
||
) carina weak, with small irregularly placed granules, subdigital (
|
||
<emphasis id="E386EADAFFAFC267FB5FF903FB68B9EB" box="[1164,1196,1703,1727]" italics="true" pageId="7" pageNumber="6">D2</emphasis>
|
||
) represented with a single rounded granule, dorsosecondary (
|
||
<emphasis id="E386EADAFFAFC267FAC1F961FAF7B989" box="[1298,1331,1733,1757]" italics="true" pageId="7" pageNumber="6">D3</emphasis>
|
||
) rounded with slight median granules, dorsomarginal (
|
||
<emphasis id="E386EADAFFAFC267FAE1F940FA96B9A8" box="[1330,1362,1764,1788]" italics="true" pageId="7" pageNumber="6">D4</emphasis>
|
||
) rounded with scattered granules, dorsointernal (
|
||
<emphasis id="E386EADAFFAFC267FAC5F8A7FAF3B84E" box="[1302,1335,1795,1818]" italics="true" pageId="7" pageNumber="6">D5</emphasis>
|
||
) rounded and irregularly granulated with pointed granules, ventroexternal (
|
||
<emphasis id="E386EADAFFAFC267FC26F8E4FBD7B80C" box="[1013,1043,1856,1880]" italics="true" pageId="7" pageNumber="6">
|
||
<collectionCode id="B7E3AE0DFFAFC267FC26F8E4FBCCB80C" box="[1013,1032,1856,1880]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="6" type="Museum">V</collectionCode>
|
||
1
|
||
</emphasis>
|
||
) and ventromedian (
|
||
<emphasis id="E386EADAFFAFC267FACAF8E4FAF2B80C" box="[1305,1334,1856,1880]" italics="true" pageId="7" pageNumber="6">
|
||
<collectionCode id="B7E3AE0DFFAFC267FACAF8E4FAEEB80C" box="[1305,1322,1856,1880]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="6" type="Museum">V</collectionCode>
|
||
2
|
||
</emphasis>
|
||
) carinae rounded and smooth, ventrointernal (
|
||
<emphasis id="E386EADAFFAFC267FB0CF8FAFB39B822" box="[1247,1277,1886,1910]" italics="true" pageId="7" pageNumber="6">
|
||
<collectionCode id="B7E3AE0DFFAFC267FB0CF8FAFB34B822" box="[1247,1264,1886,1910]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="6" type="Museum">V</collectionCode>
|
||
3
|
||
</emphasis>
|
||
) rounded with minor granulation, and external (
|
||
<emphasis id="E386EADAFFA0C268FD9BFB25FD9CBBCC" box="[584,600,1153,1176]" italics="true" pageId="8" pageNumber="7">E</emphasis>
|
||
) carina weak to obsolete. Chelal finger median denticle (
|
||
<emphasis id="E386EADAFFA0C268FD5DFB04FD7CBBE3" box="[654,696,1184,1207]" italics="true" pageId="8" pageNumber="7">
|
||
<collectionCode id="B7E3AE0DFFA0C268FD5DFB04FD7CBBE3" box="[654,696,1184,1207]" country="Germany" name="Museum Donaueschingen" pageId="8" pageNumber="7">MD</collectionCode>
|
||
</emphasis>
|
||
) rows in straight line. Fixed finger median denticles (
|
||
<emphasis id="E386EADAFFA0C268FD61FB1BFD18BB82" box="[690,732,1215,1238]" italics="true" pageId="8" pageNumber="7">
|
||
<collectionCode id="B7E3AE0DFFA0C268FD61FB1BFD18BB82" box="[690,732,1215,1238]" country="Germany" name="Museum Donaueschingen" pageId="8" pageNumber="7">MD</collectionCode>
|
||
</emphasis>
|
||
) divided into 6 groups by 5 outer (
|
||
<emphasis id="E386EADAFFA0C268FDC6FB7AFDFFBBA1" box="[533,571,1246,1269]" italics="true" pageId="8" pageNumber="7">
|
||
<collectionCode id="B7E3AE0DFFA0C268FDC6FB7AFDFFBBA1" box="[533,571,1246,1269]" pageId="8" pageNumber="7">OD</collectionCode>
|
||
</emphasis>
|
||
) denticles, and 6
|
||
<emphasis id="E386EADAFFA0C268FCD5FB7BFCE6BBA2" box="[774,802,1247,1270]" italics="true" pageId="8" pageNumber="7">ID</emphasis>
|
||
denticles are found on the inner edge. Movable finger with 6
|
||
<emphasis id="E386EADAFFA0C268FEC2FAB9FEFEBA60" box="[273,314,1309,1332]" italics="true" pageId="8" pageNumber="7">
|
||
<collectionCode id="B7E3AE0DFFA0C268FEC2FAB9FEFEBA60" box="[273,314,1309,1332]" country="Germany" name="Museum Donaueschingen" pageId="8" pageNumber="7">MD</collectionCode>
|
||
</emphasis>
|
||
groups, 5
|
||
<emphasis id="E386EADAFFA0C268FE66FAB9FE18BA60" box="[437,476,1309,1332]" italics="true" pageId="8" pageNumber="7">OD</emphasis>
|
||
denticles and 6
|
||
<emphasis id="E386EADAFFA0C268FD45FAB9FD76BA60" box="[662,690,1309,1332]" italics="true" pageId="8" pageNumber="7">ID</emphasis>
|
||
denticles. Trichobothrial pattern
|
||
<typeStatus id="0E49886AFFA0C268FE60FA98FE25BA00" box="[435,481,1340,1364]" pageId="8" pageNumber="7">type</typeStatus>
|
||
C orthobothriotaxic (see
|
||
<figureCitation id="49C92A4DFFA0C268FD26FA98FF3BBA27" captionStart="Figure 7" captionStartId="6.[192,267,1876,1900]" captionTargetBox="[202,1381,199,1841]" captionTargetId="figure-24@6.[192,1438,192,1847]" captionTargetPageId="6" captionText="Figure 7: Vaejovis troupi sp. n., female holotype. Trichobothrial pattern." figureDoi="http://doi.org/10.5281/zenodo.5508072" httpUri="https://zenodo.org/record/5508072/files/figure.png" pageId="8" pageNumber="7">Figure 7</figureCitation>
|
||
). Chelal
|
||
<emphasis id="E386EADAFFA0C268FEB8FAFFFE44BA27" box="[363,384,1371,1395]" italics="true" pageId="8" pageNumber="7">ib</emphasis>
|
||
and
|
||
<emphasis id="E386EADAFFA0C268FE6CFAF8FE0ABA27" box="[447,462,1372,1395]" italics="true" pageId="8" pageNumber="7">it</emphasis>
|
||
trichobothria located at fixed finger’s base, considerably proximal of sixth
|
||
<emphasis id="E386EADAFFA0C268FCD5FADFFCE6BAC6" box="[774,802,1403,1426]" italics="true" pageId="8" pageNumber="7">ID</emphasis>
|
||
denticle;
|
||
<emphasis id="E386EADAFFA0C268FEF5FA3FFE85BAE6" box="[294,321,1435,1458]" italics="true" pageId="8" pageNumber="7">Dt</emphasis>
|
||
on chela is proximal of palm midpoint;
|
||
<emphasis id="E386EADAFFA0C268FCDEFA3EFCE6BAE6" box="[781,802,1434,1458]" italics="true" pageId="8" pageNumber="7">dt</emphasis>
|
||
and
|
||
<emphasis id="E386EADAFFA0C268FF3CFA1DFECABA85" box="[239,270,1465,1489]" italics="true" pageId="8" pageNumber="7">dst</emphasis>
|
||
are proximal to
|
||
<emphasis id="E386EADAFFA0C268FE16FA1EFE1CBA85" box="[453,472,1466,1489]" italics="true" pageId="8" pageNumber="7">et</emphasis>
|
||
and distal of
|
||
<emphasis id="E386EADAFFA0C268FDBCFA1EFD49BA85" box="[623,653,1466,1489]" italics="true" pageId="8" pageNumber="7">est</emphasis>
|
||
; patellar
|
||
<emphasis id="E386EADAFFA0C268FD26FA1EFCCEBA82" box="[757,778,1466,1494]" italics="true" pageId="8" pageNumber="7">
|
||
v
|
||
<subScript id="4D76348DFFA0C268FCD2FA6CFCCEBA82" attach="left" box="[769,778,1480,1494]" fontSize="6" pageId="8" pageNumber="7">3</subScript>
|
||
</emphasis>
|
||
is located on external surface and positioned distally of
|
||
<emphasis id="E386EADAFFA0C268FD2CFA7AFCDFBAAF" box="[767,795,1502,1531]" italics="true" pageId="8" pageNumber="7">
|
||
et
|
||
<subScript id="4D76348DFFA0C268FCC1FA49FCDFBAAF" attach="left" box="[786,795,1517,1531]" fontSize="6" pageId="8" pageNumber="7">3</subScript>
|
||
</emphasis>
|
||
. Trichobothrium
|
||
<emphasis id="E386EADAFFA0C268FEA3F9A7FE49B974" box="[368,397,1539,1568]" italics="true" pageId="8" pageNumber="7">
|
||
et
|
||
<subScript id="4D76348DFFA0C268FE57F9B6FE49B974" attach="left" box="[388,397,1554,1568]" fontSize="6" pageId="8" pageNumber="7">3</subScript>
|
||
</emphasis>
|
||
is located proximal of a line connecting
|
||
<emphasis id="E386EADAFFA0C268FF23F98CFECAB96B" box="[240,270,1576,1599]" italics="true" pageId="8" pageNumber="7">est</emphasis>
|
||
and
|
||
<emphasis id="E386EADAFFA0C268FE91F98CFE92B96B" box="[322,342,1576,1599]" italics="true" pageId="8" pageNumber="7">
|
||
v
|
||
<subScript id="4D76348DFFA0C268FE9EF995FE92B96B" attach="left" box="[333,342,1585,1599]" fontSize="6" pageId="8" pageNumber="7">3</subScript>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<caption id="858D6640FFA0C268FBD8FC99FA58BB18" ID-DOI="http://doi.org/10.5281/zenodo.5508074" ID-Zenodo-Dep="5508074" httpUri="https://zenodo.org/record/5508074/files/figure.png" pageId="8" pageNumber="7" startId="8.[1035,1110,829,853]" targetBox="[195,961,205,1104]" targetPageId="8">
|
||
<paragraph id="D14D36C8FFA0C268FBD8FC99FA58BB18" blockId="8.[1035,1441,829,1100]" pageId="8" pageNumber="7">
|
||
<emphasis id="E386EADAFFA0C268FBD8FC99FBBCBC01" bold="true" box="[1035,1144,829,853]" pageId="8" pageNumber="7">Figure 8:</emphasis>
|
||
Chelal fixed finger dentition showing the number of inner denticles (
|
||
<emphasis id="E386EADAFFA0C268FBC0FCD4FBE8BCD1" box="[1043,1068,880,901]" italics="true" pageId="8" pageNumber="7">ID</emphasis>
|
||
).
|
||
<emphasis id="E386EADAFFA0C268FB96FCD4FBB2BCD1" bold="true" box="[1093,1142,880,901]" pageId="8" pageNumber="7">Left.</emphasis>
|
||
<taxonomicName id="16F24D4BFFA0C268FB52FCD4FADDBCD1" authority="Ayrey & Soleglad, 2015" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[1153,1305,880,901]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="troupi" status="sp. nov.">
|
||
<emphasis id="E386EADAFFA0C268FB52FCD4FADDBCD1" box="[1153,1305,880,901]" italics="true" pageId="8" pageNumber="7">Vaejovis troupi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E386EADAFFA0C268FAF0FCD4FA99BCD1" bold="true" box="[1315,1373,880,901]" pageId="8" pageNumber="7">
|
||
<taxonomicNameLabel id="F8B557A1FFA0C268FAF0FCD4FA99BCD1" box="[1315,1373,880,901]" pageId="8" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype female (left fixed finger, reversed).
|
||
<emphasis id="E386EADAFFA0C268FBD8FC05FB9CBCE2" bold="true" box="[1035,1112,929,950]" pageId="8" pageNumber="7">Center.</emphasis>
|
||
<taxonomicName id="16F24D4BFFA0C268FBBAFC06FAEEBC9B" authority=", Madera Canyon, Santa Rita Mountains. Right." class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA0C268FBBAFC06FB1BBCE3" box="[1129,1247,929,951]" italics="true" pageId="8" pageNumber="7">V. grahami</emphasis>
|
||
, Madera Canyon, Santa Rita Mountains.
|
||
<emphasis id="E386EADAFFA0C268FB38FC1EFAEEBC9B" bold="true" box="[1259,1322,954,975]" pageId="8" pageNumber="7">Right.</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="16F24D4BFFA0C268FAE2FC1FFA5EBC84" authorityName=": Ayrey" authorityYear="2013" box="[1329,1434,954,976]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA0C268FAE2FC1FFA5EBC84" box="[1329,1434,954,976]" italics="true" pageId="8" pageNumber="7">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
, Miller’s Canyon, Huachuca Mountains. Of particular interest, note that
|
||
<taxonomicName id="16F24D4BFFA0C268FA9BFC49FA65BB56" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[1352,1441,1005,1026]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA0C268FA9BFC49FA65BB56" box="[1352,1441,1005,1026]" italics="true" pageId="8" pageNumber="7">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
has six
|
||
<emphasis id="E386EADAFFA0C268FBB2FBA2FBBEBB4F" box="[1121,1146,1030,1051]" italics="true" pageId="8" pageNumber="7">ID</emphasis>
|
||
denticles whereas the other two species exhibit only five
|
||
<emphasis id="E386EADAFFA0C268FAF6FBBAFAFABB67" box="[1317,1342,1054,1075]" italics="true" pageId="8" pageNumber="7">ID</emphasis>
|
||
denticles. Internal trichobothria
|
||
<emphasis id="E386EADAFFA0C268FB0DFB92FB34BB18" box="[1246,1264,1078,1100]" italics="true" pageId="8" pageNumber="7">ib</emphasis>
|
||
and
|
||
<emphasis id="E386EADAFFA0C268FAF3FB93FAE9BB18" box="[1312,1325,1079,1100]" italics="true" pageId="8" pageNumber="7">it</emphasis>
|
||
are shown.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D14D36C8FFA0C268FF13F9CFFEB6B994" blockId="8.[192,801,1643,1728]" pageId="8" pageNumber="7">
|
||
<emphasis id="E386EADAFFA0C268FF13F9CFFEB0B9D7" bold="true" box="[192,372,1643,1667]" pageId="8" pageNumber="7">
|
||
<collectionCode id="B7E3AE0DFFA0C268FF13F9CFFEC3B9D7" box="[192,263,1643,1667]" pageId="8" pageNumber="7">LEGS</collectionCode>
|
||
(
|
||
<figureCitation id="49C92A4DFFA0C268FECFF9CFFEA1B9D7" box="[284,357,1643,1667]" captionStart="Figures 2-6" captionStartId="4.[192,278,1496,1520]" captionTargetBox="[195,1394,180,1459]" captionTargetId="figure-204@4.[192,1437,168,1467]" captionTargetPageId="4" captionText="Figures 2-6: Vaejovis troupi sp. n., female holotype. 2. Metasoma and telson, ventral and lateral views. 3. Carapace and closeup of lateral eyes. 4. Pedipalp chela, external view, patella and femur, dorsal views. 5. Right leg III basitarsus and tarsus, ventral view. 6. Chelal fixed and movable fingers, each exhibiting six inner denticles (ID)." figureDoi="http://doi.org/10.5281/zenodo.5508070" httpUri="https://zenodo.org/record/5508070/files/figure.png" pageId="8" pageNumber="7">Fig. 5</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Ventral surface of tarsomere II with single median row of spinules terminating distally with one spinule pair.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFA0C268FF13F948FB83BBA0" blockId="8.[192,802,1772,1918]" lastBlockId="8.[831,1440,1152,1268]" pageId="8" pageNumber="7">
|
||
<emphasis id="E386EADAFFA0C268FF13F948FEA0B850" bold="true" box="[192,356,1772,1796]" pageId="8" pageNumber="7">Reproduction.</emphasis>
|
||
This species is not represented in the
|
||
<taxonomicName id="16F24D4BFFA0C268FF13F8AFFED8B876" authorityName="Thorell" authorityYear="1876" box="[192,284,1803,1826]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E386EADAFFA0C268FF13F8AFFED8B876" box="[192,284,1803,1826]" italics="true" pageId="8" pageNumber="7">Vaejovis</emphasis>
|
||
</taxonomicName>
|
||
“vorhiesi” group study (Ayrey, 2013a) but is presumed to have behavior similar to the rest of the group. The “vorhiesi” group average clutch size is 22.87 first instars (n=100 births, 2,287 first instars). Being one of the larger species it would have a relatively large clutch size, estimated to be approximately 25 first instars. Birth and postpartum behavior would be as described in Ayrey (2013a).
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFA0C268FCECFABFFBB1BA6C" blockId="8.[831,1141,1307,1336]" box="[831,1141,1307,1336]" pageId="8" pageNumber="7">
|
||
<heading id="8A0581A4FFA0C268FCECFABFFBB1BA6C" box="[831,1141,1307,1336]" pageId="8" pageNumber="7">
|
||
<emphasis id="E386EADAFFA0C268FCECFABFFBB1BA6C" bold="true" box="[831,1141,1307,1336]" pageId="8" pageNumber="7">Comparison of Species</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFA0C268FCBCFAF9FBB2B9DC" blockId="8.[831,1441,1373,1918]" pageId="8" pageNumber="7">
|
||
With the description of
|
||
<taxonomicName id="16F24D4BFFA0C268FB50FAF9FAE9BA20" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[1155,1325,1373,1396]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA0C268FB50FAF9FAE9BA20" box="[1155,1325,1373,1396]" italics="true" pageId="8" pageNumber="7">Vaejovis troupi</emphasis>
|
||
</taxonomicName>
|
||
presented herein, 18 species are now currently placed in the “
|
||
<taxonomicName id="16F24D4BFFA0C268FC99FA3EFC64BAE6" authorityName=": Ayrey" authorityYear="2013" box="[842,928,1434,1458]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA0C268FC99FA3EFC64BAE6" box="[842,928,1434,1458]" italics="true" pageId="8" pageNumber="7">vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
” group of
|
||
<taxonomicName id="16F24D4BFFA0C268FBFDFA3FFB4EBAE6" authorityName="Thorell" authorityYear="1876" box="[1070,1162,1435,1458]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E386EADAFFA0C268FBFDFA3FFB4EBAE6" box="[1070,1162,1435,1458]" italics="true" pageId="8" pageNumber="7">Vaejovis</emphasis>
|
||
</taxonomicName>
|
||
. Morphologically, these species are roughly partitioned into two presumably phylogenetic groups, those with seven inner denticles (
|
||
<emphasis id="E386EADAFFA0C268FAA8FA7CFA53BABB" box="[1403,1431,1496,1519]" italics="true" pageId="8" pageNumber="7">ID</emphasis>
|
||
) on the chelal movable finger (seven species), found primarily in central to northern Arizona, and those with six
|
||
<emphasis id="E386EADAFFA0C268FCB6F990FC45B91F" box="[869,897,1588,1611]" italics="true" pageId="8" pageNumber="7">ID</emphasis>
|
||
(eleven species) located primarily in southeastern Arizona, southwestern New
|
||
<collectingRegion id="1336F82AFFA0C268FB48F9F6FB2BB93E" box="[1179,1263,1618,1642]" country="Mexico" name="Mexico" pageId="8" pageNumber="7">Mexico</collectingRegion>
|
||
, and northern
|
||
<collectingCountry id="A9E57658FFA0C268FCECF9D4FC54B9DC" box="[831,912,1648,1672]" name="Mexico" pageId="8" pageNumber="7">Mexico</collectingCountry>
|
||
(see Map in
|
||
<figureCitation id="49C92A4DFFA0C268FBC8F9D4FBADB9DC" box="[1051,1129,1648,1672]" captionStart="Figure 11" captionStartId="11.[192,267,1642,1666]" captionTargetBox="[191,1441,191,1603]" captionTargetId="figure-171@11.[192,1440,192,1603]" captionTargetPageId="11" captionText="Figure 11: Map of Arizona, extreme western New Mexico and northern Sonora (Mexico) showing the type locality of all 17 Vaejovis “vorhiesi” group species, including new species Vaejovis troupi. Localities are divided into those species exhibiting seven inner denticles (ID) on the chelal movable finger (white rectangles with black lettering) and those with primarily six, or five ID denticles (black rectangles with white lettering). Seven IDs: 1 = V. jonesi, 2 = V. lapidicola, 3 = V. paysonensis, 4 = V. crumpi, 5 = V. bigelowi, 14 = V. trinityae, 17 = V. grayae. Six IDs: 6 = V. vorhiesi, 7 = V. cashi, 8 = V. feti, 9 = V. deboerae, 10 = V. electrum, 11 = V. tenuipalpus 12 = V. halli, 13 = V. brysoni, 15 = V. bandido, 16 = V. grahami, and 18 = V. troupi sp. n." figureDoi="http://doi.org/10.5281/zenodo.5508080" httpUri="https://zenodo.org/record/5508080/files/figure.png" pageId="8" pageNumber="7">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFA0C268FCBCF934FB61B82A" blockId="8.[831,1441,1373,1918]" pageId="8" pageNumber="7">
|
||
In this contribution we compare
|
||
<taxonomicName id="16F24D4BFFA0C268FB34F934FA8FB9F3" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[1255,1355,1680,1703]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA0C268FB34F934FA8FB9F3" box="[1255,1355,1680,1703]" italics="true" pageId="8" pageNumber="7">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
to two other species,
|
||
<taxonomicName id="16F24D4BFFA0C268FC33F90BFB98B992" box="[992,1116,1710,1734]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA0C268FC33F90BFB98B992" box="[992,1116,1710,1734]" italics="true" pageId="8" pageNumber="7">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA0C268FB49F90BFAD6B992" authorityName=": Ayrey" authorityYear="2013" box="[1178,1298,1710,1734]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA0C268FB49F90BFAD6B992" box="[1178,1298,1710,1734]" italics="true" pageId="8" pageNumber="7">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
, both which are found in adjacent sky island mountain ranges in southeastern
|
||
<collectingRegion id="1336F82AFFA0C268FC1FF94FFBE1B857" box="[972,1061,1771,1795]" country="United States of America" name="Arizona" pageId="8" pageNumber="7">Arizona</collectingRegion>
|
||
. The choice of these two species is based, in part, on the recent phylogenetic analysis based on molecular considerations (see Bryson et al., 2013), which suggests that these two species are the closest relatives of
|
||
<taxonomicName id="16F24D4BFFA0C268FC6FF8C3FBDFB82A" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[956,1051,1895,1918]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA0C268FC6FF8C3FBDFB82A" box="[956,1051,1895,1918]" italics="true" pageId="8" pageNumber="7">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
(see below).
|
||
</paragraph>
|
||
<caption id="858D6640FFA1C269FF13FB94FBD1BA5C" ID-DOI="http://doi.org/10.5281/zenodo.5508076" ID-Zenodo-Dep="5508076" httpUri="https://zenodo.org/record/5508076/files/figure.png" pageId="9" pageNumber="8" startId="9.[192,267,1072,1096]" targetBox="[191,1441,191,1038]" targetPageId="9">
|
||
<paragraph id="D14D36C8FFA1C269FF13FB94FBD1BA5C" blockId="9.[192,1440,1072,1288]" pageId="9" pageNumber="8">
|
||
<emphasis id="E386EADAFFA1C269FF13FB94FEEDBB1C" bold="true" box="[192,297,1072,1096]" pageId="9" pageNumber="8">Figure 9:</emphasis>
|
||
Comparison of the relative positions of five patellar trichobothria
|
||
<emphasis id="E386EADAFFA1C269FC7DFB96FC28BB13" box="[942,1004,1074,1095]" italics="true" pageId="9" pageNumber="8">
|
||
et
|
||
<subScript id="4D76348DFFA1C269FC6CFB9CFC03BB13" attach="left" box="[959,967,1080,1095]" fontSize="6" pageId="9" pageNumber="8">1</subScript>
|
||
–et
|
||
<subScript id="4D76348DFFA1C269FC37FB9CFC28BB13" attach="left" box="[996,1004,1080,1095]" fontSize="6" pageId="9" pageNumber="8">3</subScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="E386EADAFFA1C269FC2AFB96FBD0BB13" box="[1017,1044,1074,1095]" italics="true" pageId="9" pageNumber="8">est</emphasis>
|
||
, and
|
||
<emphasis id="E386EADAFFA1C269FB99FB96FB98BB13" box="[1098,1116,1074,1095]" italics="true" pageId="9" pageNumber="8">
|
||
v
|
||
<subScript id="4D76348DFFA1C269FB87FB9CFB98BB13" attach="left" box="[1108,1116,1080,1095]" fontSize="6" pageId="9" pageNumber="8">3</subScript>
|
||
</emphasis>
|
||
of species
|
||
<taxonomicName id="16F24D4BFFA1C269FB1AFB96FA98BB13" authority="Ayrey & Soleglad, 2015" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[1225,1372,1074,1095]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="troupi" status="sp. nov.">
|
||
<emphasis id="E386EADAFFA1C269FB1AFB96FA98BB13" box="[1225,1372,1074,1095]" italics="true" pageId="9" pageNumber="8">Vaejovis troupi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E386EADAFFA1C269FAB7FB95FA5EBB12" bold="true" box="[1380,1434,1073,1094]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="F8B557A1FFA1C269FAB7FB95FA5EBB12" box="[1380,1434,1073,1094]" pageId="9" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="16F24D4BFFA1C269FF13FBEAFEE9BB37" box="[192,301,1101,1123]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA1C269FF13FBEAFEE9BB37" box="[192,301,1101,1123]" italics="true" pageId="9" pageNumber="8">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="16F24D4BFFA1C269FEB2FBEAFE0DBB37" authorityName=": Ayrey" authorityYear="2013" box="[353,457,1101,1123]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA1C269FEB2FBEAFE0DBB37" box="[353,457,1101,1123]" italics="true" pageId="9" pageNumber="8">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="E386EADAFFA1C269FE05FBE9FDC0BB36" bold="true" box="[470,516,1101,1122]" pageId="9" pageNumber="8">Top.</emphasis>
|
||
Trichobothria positions of the three female type specimens.
|
||
<emphasis id="E386EADAFFA1C269FB9FFBE9FB61BB37" box="[1100,1189,1101,1123]" italics="true" pageId="9" pageNumber="8">Red lines</emphasis>
|
||
indicate the angle formed by trichobothria
|
||
<emphasis id="E386EADAFFA1C269FEB5FBCDFE09BB2A" box="[358,461,1128,1150]" italics="true" pageId="9" pageNumber="8">est–et3–v3</emphasis>
|
||
, contrasting
|
||
<taxonomicName id="16F24D4BFFA1C269FD9CFBCDFD6CBB2A" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[591,680,1129,1150]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA1C269FD9CFBCDFD6CBB2A" box="[591,680,1129,1150]" italics="true" pageId="9" pageNumber="8">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
from the other two species.
|
||
<emphasis id="E386EADAFFA1C269FC19FBCDFBD8BB2A" bold="true" box="[970,1052,1129,1150]" pageId="9" pageNumber="8">Bottom.</emphasis>
|
||
Each individual pattern for a pedipalp patella is anchored at trichobothrium
|
||
<emphasis id="E386EADAFFA1C269FDFBFB21FD86BBCE" box="[552,578,1157,1178]" italics="true" pageId="9" pageNumber="8">
|
||
et
|
||
<subScript id="4D76348DFFA1C269FDE9FB2FFD86BBCE" attach="left" box="[570,578,1163,1178]" fontSize="6" pageId="9" pageNumber="8">1</subScript>
|
||
</emphasis>
|
||
while other trichobothria are plotted at their relative locations forming clusters.
|
||
<emphasis id="E386EADAFFA1C269FA94FB20FA64BBCE" box="[1351,1440,1156,1178]" italics="true" pageId="9" pageNumber="8">Red lines</emphasis>
|
||
indicate the
|
||
<emphasis id="E386EADAFFA1C269FEEAFB05FE42BBE2" box="[313,390,1185,1206]" italics="true" pageId="9" pageNumber="8">average</emphasis>
|
||
angle formed by trichobothria
|
||
<emphasis id="E386EADAFFA1C269FD6FFB05FCE7BBE2" box="[700,803,1184,1206]" italics="true" pageId="9" pageNumber="8">est–et3–v3</emphasis>
|
||
. Patterns of left patellae were reversed for these plots which are shown from a right-patella perspective. Only adult female specimens were used in these plots. Number below species name indicates the number of pedipalp patellae considered. Information for
|
||
<taxonomicName id="16F24D4BFFA1C269FC5DFB7CFBFDBBB9" authorityName=": Ayrey" authorityYear="2013" box="[910,1081,1239,1261]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA1C269FC5DFB7CFBFDBBB9" box="[910,1081,1239,1261]" italics="true" pageId="9" pageNumber="8">Vaejovis vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA1C269FBA1FB7CFB27BBB9" box="[1138,1251,1239,1261]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA1C269FBA1FB7CFB27BBB9" box="[1138,1251,1239,1261]" italics="true" pageId="9" pageNumber="8">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
lectotype/holotype females was extracted from Graham (2007: fig. 9) and Ayrey & Soleglad (2014: fig. 11).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D14D36C8FFA1C269FF23FA99FBAFBA27" blockId="9.[192,802,1341,1917]" lastBlockId="9.[831,1441,1341,1917]" pageId="9" pageNumber="8">
|
||
<emphasis id="E386EADAFFA1C269FF23FA99FDE4BA01" bold="true" box="[240,544,1341,1365]" pageId="9" pageNumber="8">Geological considerations.</emphasis>
|
||
Weldon Heald (1951) coined the term “Sky Islands” to describe the distinct isolated groups of Madrean pine-oak woodlands found at higher elevations in a complex of small mountain ranges in southern and southeastern Arizona, southwestern New
|
||
<collectingRegion id="1336F82AFFA1C269FF13FA72FED0BABA" box="[192,276,1494,1518]" country="Mexico" name="Mexico" pageId="9" pageNumber="8">Mexico</collectingRegion>
|
||
, and northwestern
|
||
<collectingCountry id="A9E57658FFA1C269FE30FA72FDFCBABA" box="[483,568,1494,1518]" name="Mexico" pageId="9" pageNumber="8">Mexico</collectingCountry>
|
||
. These “Sky Islands” are surrounded at lower elevations by the Sonoran and Chihuahuan Deserts thus essentially being isolated by different extreme xeric climates and different microhabitats. Here we use an expanded definition that includes all isolated, high elevation habitats in Arizona, New
|
||
<collectingRegion id="1336F82AFFA1C269FED3F92AFE96B9F2" box="[256,338,1678,1702]" country="Mexico" name="Mexico" pageId="9" pageNumber="8">Mexico</collectingRegion>
|
||
and
|
||
<collectingCountry id="A9E57658FFA1C269FE40F92AFE2CB9F2" box="[403,488,1678,1702]" name="Mexico" pageId="9" pageNumber="8">Mexico</collectingCountry>
|
||
. Many habitats in northern Arizona and northern New
|
||
<collectingRegion id="1336F82AFFA1C269FE29F909FD8FB991" box="[506,587,1709,1733]" country="Mexico" name="Mexico" pageId="9" pageNumber="8">Mexico</collectingRegion>
|
||
are high elevation habitats isolated by the intervening Colorado Plateau. Graham (2007: 1) states: “...Faunas adapted to these isolated mountain environments are therefore disconnected from similar habitat on adjacent mountain ranges by extreme climates of xeric lowlands. ...”. This paper applies to the isolated, elevation habitats on the Colorado Plateau as well as it does the Madrean pineoak woodlands of the south.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFA1C26CFCBCFADEFDC9BD8D" blockId="9.[831,1441,1341,1917]" lastBlockId="12.[192,802,674,1894]" lastPageId="12" lastPageNumber="11" pageId="9" pageNumber="8">
|
||
These three species are found in three separate geographically close, but distinctly isolated, mountain ranges,
|
||
<taxonomicName id="16F24D4BFFA1C269FC47FA1CFC31BA9B" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[916,1013,1464,1487]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA1C269FC47FA1CFC31BA9B" box="[916,1013,1464,1487]" italics="true" pageId="9" pageNumber="8">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
found in the Whetstone Mountains,
|
||
<taxonomicName id="16F24D4BFFA1C269FA5AFA1CFC5DBABA" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA1C269FA5AFA1CFC5DBABA" italics="true" pageId="9" pageNumber="8">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
from the Santa Rita Mountains, and
|
||
<taxonomicName id="16F24D4BFFA1C269FAF8FA73FA64BABA" authorityName=": Ayrey" authorityYear="2013" box="[1323,1440,1494,1518]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA1C269FAF8FA73FA64BABA" box="[1323,1440,1494,1518]" italics="true" pageId="9" pageNumber="8">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
from the Huachuca Mountains (see
|
||
<figureCitation id="49C92A4DFFA1C269FB13FA51FAD4B959" box="[1216,1296,1525,1549]" captionStart="Figure 10" captionStartId="10.[192,267,1648,1672]" captionTargetBox="[191,1441,220,1609]" captionTargetId="figure-155@10.[192,1440,221,1608]" captionTargetPageId="10" captionText="Figure 10: Proposed geographic and molecular relationships of “sky island” species Vaejovis troupi sp. n., V. grahami, and V. vorhiesi. Top. Close-up of southeastern Arizona showing the type localities of the three species and the distinct, isolated “sky island” mountain ranges in which they occur, Whetstone, Santa Rita, and Huachuca Mountains, respectively. Bottom. A partial chronogram based on molecular data showing proposed phylogenetic relationships based on evolutionary time (red numbers). Red numbers above branch indicate estimates for a multilocus species tree, bottom numbers represent a mtDNA gene tree. Means and 95% highest posterior densities are shown for each node. Red numbers indicate proposed minimum, maximum, and mean Mya. Information is from Bryson et al. (2013: fig. S2). Gray rectangle delineates the three species discussed in this paper. Altitude data is that of the type localities." figureDoi="http://doi.org/10.5281/zenodo.5508078" httpUri="https://zenodo.org/record/5508078/files/figure.png" pageId="9" pageNumber="8">Fig. 10</figureCitation>
|
||
). These three isolated mountain ranges are situated quite close to each other, only
|
||
<quantity id="160A9B2DFFA1C269FC11F996FB8EB91E" box="[962,1098,1586,1610]" metricMagnitude="4" metricUnit="m" metricValue="5.55" metricValueMax="6.6" metricValueMin="4.5" pageId="9" pageNumber="8" unit="km" value="55.5" valueMax="66.0" valueMin="45.0">45 – 66 km</quantity>
|
||
separate the
|
||
<typeStatus id="0E49886AFFA1C269FB37F996FAD6B91E" box="[1252,1298,1586,1610]" pageId="9" pageNumber="8">type</typeStatus>
|
||
localities of these three species. The Huachuca Mountains are the most southern and highest, their Miller’s Peak reaching
|
||
<quantity id="160A9B2DFFA1C269FCECF92AFC56B9F2" box="[831,914,1678,1702]" metricMagnitude="3" metricUnit="m" metricValue="2.878" pageId="9" pageNumber="8" unit="m" value="2878.0">2878 m</quantity>
|
||
. It extends roughly
|
||
<quantity id="160A9B2DFFA1C269FBB4F92AFB6FB9F2" box="[1127,1195,1678,1702]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="8" unit="km" value="32.0">32 km</quantity>
|
||
in a north to southeast direction reaching
|
||
<collectingCountry id="A9E57658FFA1C269FBC1F909FBA3B991" box="[1042,1127,1709,1733]" name="Mexico" pageId="9" pageNumber="8">Mexico</collectingCountry>
|
||
. The Santa Rita Mountains are the largest mountain range, extending roughly
|
||
<quantity id="160A9B2DFFA1C269FA56F96FFCA0B856" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="9" pageNumber="8" unit="km" value="42.0">42 km</quantity>
|
||
, with its highest point Mount Wrightson at
|
||
<quantity id="160A9B2DFFA1C269FAE9F94EFA58B856" box="[1338,1436,1770,1794]" metricMagnitude="6" metricUnit="m" metricValue="2.863" pageId="9" pageNumber="8" unit="km" value="2863.0">2863 km</quantity>
|
||
. The Whetstone Mountains, which are located directly north of the Huachuca Mountains, is the smallest range, only
|
||
<quantity id="160A9B2DFFA1C269FCA6F8E2FC7EB80A" box="[885,954,1862,1886]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="8" unit="km" value="18.0">18 km</quantity>
|
||
long in a north to south direction, its highest point is the Apache Peak at
|
||
<quantity id="160A9B2DFFA1C269FBBCF8C1FB06B828" box="[1135,1218,1893,1917]" metricMagnitude="3" metricUnit="m" metricValue="2.346" pageId="9" pageNumber="8" unit="m" value="2346.0">2346 m</quantity>
|
||
.
|
||
<figureCitation id="49C92A4DFFA1C269FB1EF8C1FAF0B829" box="[1229,1332,1893,1917]" captionStart="Figure 10" captionStartId="10.[192,267,1648,1672]" captionTargetBox="[191,1441,220,1609]" captionTargetId="figure-155@10.[192,1440,221,1608]" captionTargetPageId="10" captionText="Figure 10: Proposed geographic and molecular relationships of “sky island” species Vaejovis troupi sp. n., V. grahami, and V. vorhiesi. Top. Close-up of southeastern Arizona showing the type localities of the three species and the distinct, isolated “sky island” mountain ranges in which they occur, Whetstone, Santa Rita, and Huachuca Mountains, respectively. Bottom. A partial chronogram based on molecular data showing proposed phylogenetic relationships based on evolutionary time (red numbers). Red numbers above branch indicate estimates for a multilocus species tree, bottom numbers represent a mtDNA gene tree. Means and 95% highest posterior densities are shown for each node. Red numbers indicate proposed minimum, maximum, and mean Mya. Information is from Bryson et al. (2013: fig. S2). Gray rectangle delineates the three species discussed in this paper. Altitude data is that of the type localities." figureDoi="http://doi.org/10.5281/zenodo.5508078" httpUri="https://zenodo.org/record/5508078/files/figure.png" pageId="9" pageNumber="8">Figure 10</figureCitation>
|
||
states the altitudes of the
|
||
<typeStatus id="0E49886AFFA4C26CFEB9FD07FE5CBDEF" box="[362,408,675,699]" pageId="12" pageNumber="11">type</typeStatus>
|
||
localities of the three species found in these three mountain ranges.
|
||
</paragraph>
|
||
<caption id="858D6640FFA2C26AFF13F9D4FD89B81C" ID-DOI="http://doi.org/10.5281/zenodo.5508078" ID-Zenodo-Dep="5508078" httpUri="https://zenodo.org/record/5508078/files/figure.png" pageId="10" pageNumber="9" startId="10.[192,267,1648,1672]" targetBox="[191,1441,220,1609]" targetPageId="10">
|
||
<paragraph id="D14D36C8FFA2C26AFF13F9D4FD89B81C" blockId="10.[192,1441,1648,1864]" pageId="10" pageNumber="9">
|
||
<emphasis id="E386EADAFFA2C26AFF13F9D4FEF3B9DC" bold="true" box="[192,311,1648,1672]" pageId="10" pageNumber="9">Figure 10:</emphasis>
|
||
Proposed geographic and molecular relationships of “sky island” species
|
||
<taxonomicName id="16F24D4BFFA2C26AFBDBF9D6FB59B9D3" authority="Ayrey & Soleglad, 2015" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[1032,1181,1650,1671]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="troupi" status="sp. nov.">
|
||
<emphasis id="E386EADAFFA2C26AFBDBF9D6FB59B9D3" box="[1032,1181,1650,1671]" italics="true" pageId="10" pageNumber="9">Vaejovis troupi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E386EADAFFA2C26AFB77F9D5FB1FB9D2" bold="true" box="[1188,1243,1649,1670]" pageId="10" pageNumber="9">
|
||
<taxonomicNameLabel id="F8B557A1FFA2C26AFB77F9D5FB1FB9D2" box="[1188,1243,1649,1670]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="16F24D4BFFA2C26AFB3BF9D6FA92B9D3" box="[1256,1366,1649,1671]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA2C26AFB3BF9D6FA92B9D3" box="[1256,1366,1649,1671]" italics="true" pageId="10" pageNumber="9">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="16F24D4BFFA2C26AFA5FF9D6FEC9B9F7" authorityName=": Ayrey" authorityYear="2013" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA2C26AFA5FF9D6FEC9B9F7" italics="true" pageId="10" pageNumber="9">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="E386EADAFFA2C26AFECFF929FE8FB9F6" bold="true" box="[284,331,1677,1698]" pageId="10" pageNumber="9">Top.</emphasis>
|
||
Close-up of southeastern Arizona showing the type localities of the three species and the distinct, isolated “sky island” mountain ranges in which they occur, Whetstone, Santa Rita, and Huachuca Mountains, respectively.
|
||
<emphasis id="E386EADAFFA2C26AFB23F90DFA86B9EA" bold="true" box="[1264,1346,1705,1726]" pageId="10" pageNumber="9">Bottom.</emphasis>
|
||
A partial chronogram based on molecular data showing proposed phylogenetic relationships based on evolutionary time (red numbers). Red numbers above branch indicate estimates for a multilocus species tree, bottom numbers represent a mtDNA gene tree. Means and 95% highest posterior densities are shown for each node. Red numbers indicate proposed minimum, maximum, and mean Mya. Information is from Bryson et al. (2013: fig. S2). Gray rectangle delineates the three species discussed in this paper. Altitude data is that of the type localities.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="858D6640FFA3C26BFF13F9CEFAB5B9A9" ID-DOI="http://doi.org/10.5281/zenodo.5508080" ID-Zenodo-Dep="5508080" httpUri="https://zenodo.org/record/5508080/files/figure.png" pageId="11" pageNumber="10" startId="11.[192,267,1642,1666]" targetBox="[191,1441,191,1603]" targetPageId="11">
|
||
<paragraph id="D14D36C8FFA3C26BFF13F9CEFAB5B9A9" blockId="11.[192,1441,1642,1789]" pageId="11" pageNumber="10">
|
||
<emphasis id="E386EADAFFA3C26BFF13F9CEFEFCB9D6" bold="true" box="[192,312,1642,1666]" pageId="11" pageNumber="10">Figure 11:</emphasis>
|
||
Map of Arizona, extreme western New Mexico and northern Sonora (Mexico) showing the type locality of all 17
|
||
<taxonomicName id="16F24D4BFFA3C26BFF13F921FED7B9CE" authorityName="Thorell" authorityYear="1876" box="[192,275,1669,1690]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E386EADAFFA3C26BFF13F921FED7B9CE" box="[192,275,1669,1690]" italics="true" pageId="11" pageNumber="10">Vaejovis</emphasis>
|
||
</taxonomicName>
|
||
“vorhiesi” group species, including new species
|
||
<taxonomicName id="16F24D4BFFA3C26BFCD2F921FC5CB9CE" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[769,920,1669,1690]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA3C26BFCD2F921FC5CB9CE" box="[769,920,1669,1690]" italics="true" pageId="11" pageNumber="10">Vaejovis troupi</emphasis>
|
||
</taxonomicName>
|
||
. Localities are divided into those species exhibiting seven inner denticles (
|
||
<emphasis id="E386EADAFFA3C26BFE4EF93AFE72B9E7" box="[413,438,1694,1715]" italics="true" pageId="11" pageNumber="10">ID</emphasis>
|
||
) on the chelal movable finger (white rectangles with black lettering) and those with primarily six, or five
|
||
<emphasis id="E386EADAFFA3C26BFF3EF912FEC2B99F" box="[237,262,1718,1739]" italics="true" pageId="11" pageNumber="10">ID</emphasis>
|
||
denticles (black rectangles with white lettering).
|
||
<emphasis id="E386EADAFFA3C26BFD39F912FCA9B99F" bold="true" box="[746,877,1718,1739]" pageId="11" pageNumber="10">
|
||
Seven
|
||
<emphasis id="E386EADAFFA3C26BFCFEF912FC8CB99F" bold="true" box="[813,840,1718,1739]" italics="true" pageId="11" pageNumber="10">ID</emphasis>
|
||
s: 1
|
||
</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFC5AF912FC1AB99F" authorityName="Stahnke" authorityYear="1940" box="[905,990,1718,1739]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="jonesi">
|
||
<emphasis id="E386EADAFFA3C26BFC5AF912FC1AB99F" box="[905,990,1718,1739]" italics="true" pageId="11" pageNumber="10">V. jonesi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E386EADAFFA3C26BFC38F912FC33B99F" bold="true" box="[1003,1015,1718,1739]" pageId="11" pageNumber="10">2</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFBC7F912FB54B99F" authorityName="Stahnke" authorityYear="1940" box="[1044,1168,1717,1739]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="lapidicola">
|
||
<emphasis id="E386EADAFFA3C26BFBC7F912FB54B99F" box="[1044,1168,1717,1739]" italics="true" pageId="11" pageNumber="10">V. lapidicola</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E386EADAFFA3C26BFB4DF912FB6EB99F" bold="true" box="[1182,1194,1718,1739]" pageId="11" pageNumber="10">3</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFB15F912FA92B99F" authorityName="Soleglad" authorityYear="1973" box="[1222,1366,1718,1739]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="paysonensis">
|
||
<emphasis id="E386EADAFFA3C26BFB15F912FA92B99F" box="[1222,1366,1718,1739]" italics="true" pageId="11" pageNumber="10">V. paysonensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E386EADAFFA3C26BFAB7F912FAB4B99F" bold="true" box="[1380,1392,1718,1739]" pageId="11" pageNumber="10">4</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFA5FF912FEC0B9B0" authorityName="Ayrey & Soleglad" authorityYear="2011" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="crumpi">
|
||
<emphasis id="E386EADAFFA3C26BFA5FF912FEC0B9B0" italics="true" pageId="11" pageNumber="10">V. crumpi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E386EADAFFA3C26BFEC2F96BFED9B9B0" bold="true" box="[273,285,1743,1764]" pageId="11" pageNumber="10">5</emphasis>
|
||
=
|
||
<emphasis id="E386EADAFFA3C26BFEE4F96BFE6FB9B0" box="[311,427,1742,1764]" italics="true" pageId="11" pageNumber="10">
|
||
<taxonomicName id="16F24D4BFFA3C26BFEE4F96BFE63B9B0" authorityName="Sissom" authorityYear="2011" box="[311,423,1742,1764]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="bigelowi">V. bigelowi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="E386EADAFFA3C26BFE62F96BFE0DB9B0" bold="true" box="[433,457,1743,1764]" pageId="11" pageNumber="10">14</emphasis>
|
||
<emphasis id="E386EADAFFA3C26BFE1CF96BFD96B9B0" box="[463,594,1743,1764]" italics="true" pageId="11" pageNumber="10">
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFE35F96BFD96B9B0" authorityName="Ayrey" authorityYear="2013" box="[486,594,1743,1764]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="trinityae">V. trinityae</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="E386EADAFFA3C26BFD8DF96BFD4FB9B0" bold="true" box="[606,651,1743,1764]" pageId="11" pageNumber="10">17 =</emphasis>
|
||
<taxonomicName id="16F24D4BFFA3C26BFD42F96BFD31B9B0" authorityName="Ayrey" authorityYear="2014" box="[657,757,1743,1764]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="grayae">
|
||
<emphasis id="E386EADAFFA3C26BFD42F96BFD31B9B0" box="[657,757,1743,1764]" italics="true" pageId="11" pageNumber="10">V. grayae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="E386EADAFFA3C26BFCD2F96BFCA2B9B0" bold="true" box="[769,870,1743,1764]" pageId="11" pageNumber="10">
|
||
Six
|
||
<emphasis id="E386EADAFFA3C26BFCFBF96BFC87B9B0" bold="true" box="[808,835,1743,1764]" italics="true" pageId="11" pageNumber="10">ID</emphasis>
|
||
s: 6
|
||
</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFC53F96BFC2CB9B0" authorityName=": Ayrey" authorityYear="2013" box="[896,1000,1742,1764]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA3C26BFC53F96BFC2CB9B0" box="[896,1000,1742,1764]" italics="true" pageId="11" pageNumber="10">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E386EADAFFA3C26BFC26F96BFBC5B9B0" bold="true" box="[1013,1025,1743,1764]" pageId="11" pageNumber="10">7</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFBC8F96BFBACB9B0" authorityName="Graham" authorityYear="2007" box="[1051,1128,1742,1764]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="cashi">
|
||
<emphasis id="E386EADAFFA3C26BFBC8F96BFBACB9B0" box="[1051,1128,1742,1764]" italics="true" pageId="11" pageNumber="10">V. cashi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E386EADAFFA3C26BFBA6F96BFB45B9B0" bold="true" box="[1141,1153,1743,1764]" pageId="11" pageNumber="10">8</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFB48F96BFB11B9B0" authorityName="Graham" authorityYear="2007" box="[1179,1237,1742,1764]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="feti">
|
||
<emphasis id="E386EADAFFA3C26BFB48F96BFB11B9B0" box="[1179,1237,1742,1764]" italics="true" pageId="11" pageNumber="10">V. feti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E386EADAFFA3C26BFB32F96BFAC6B9B0" bold="true" box="[1249,1282,1743,1764]" pageId="11" pageNumber="10">9 =</emphasis>
|
||
<emphasis id="E386EADAFFA3C26BFADBF96BFA46B9B0" box="[1288,1410,1742,1764]" italics="true" pageId="11" pageNumber="10">
|
||
<taxonomicName id="16F24D4BFFA3C26BFADBF96BFABBB9B0" authorityName="Ayrey" authorityYear="2009" box="[1288,1407,1742,1764]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="deboerae">V. deboerae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="E386EADAFFA3C26BFA5BF96BFA64B9B0" bold="true" box="[1416,1440,1743,1764]" pageId="11" pageNumber="10">10</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFF00F94CFE86B9A9" authorityName="Hughes" authorityYear="2011" box="[211,322,1767,1789]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="electrum">
|
||
<emphasis id="E386EADAFFA3C26BFF00F94CFE86B9A9" box="[211,322,1767,1789]" italics="true" pageId="11" pageNumber="10">V. electrum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E386EADAFFA3C26BFE9CF94CFEA3B9A9" bold="true" box="[335,359,1768,1789]" pageId="11" pageNumber="10">11</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFE53F94CFDCEB9A9" box="[384,522,1767,1789]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="tenuipalpus">
|
||
<emphasis id="E386EADAFFA3C26BFE53F94CFDCEB9A9" box="[384,522,1767,1789]" italics="true" pageId="11" pageNumber="10">V. tenuipalpus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E386EADAFFA3C26BFDC3F94CFDF9B9A9" bold="true" box="[528,573,1768,1789]" pageId="11" pageNumber="10">12 =</emphasis>
|
||
<emphasis id="E386EADAFFA3C26BFD91F94CFD4BB9A9" box="[578,655,1767,1789]" italics="true" pageId="11" pageNumber="10">
|
||
<taxonomicName id="16F24D4BFFA3C26BFD91F94CFD4FB9A9" authorityName="Ayrey" authorityYear="2012" box="[578,651,1767,1789]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="halli">V. halli</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="E386EADAFFA3C26BFD46F94CFD69B9A9" bold="true" box="[661,685,1768,1789]" pageId="11" pageNumber="10">13</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFD15F94CFCEDB9A9" authorityName="Ayrey & Webber" authorityYear="2013" box="[710,809,1767,1789]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="brysoni">
|
||
<emphasis id="E386EADAFFA3C26BFD15F94CFCEDB9A9" box="[710,809,1767,1789]" italics="true" pageId="11" pageNumber="10">V. brysoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="E386EADAFFA3C26BFCE6F94CFCA5B9A9" bold="true" box="[821,865,1768,1789]" pageId="11" pageNumber="10">15 =</emphasis>
|
||
<emphasis id="E386EADAFFA3C26BFCB4F94CFC13B9A9" box="[871,983,1767,1789]" italics="true" pageId="11" pageNumber="10">
|
||
<taxonomicName id="16F24D4BFFA3C26BFCB4F94CFC17B9A9" authorityName="Graham, Ayrey & Bryson" authorityYear="2012" box="[871,979,1767,1789]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="bandido">V. bandido</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="E386EADAFFA3C26BFC0FF94CFC30B9A9" bold="true" box="[988,1012,1768,1789]" pageId="11" pageNumber="10">16</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFBDDF94CFBBFB9A9" box="[1038,1147,1767,1789]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA3C26BFBDDF94CFBBFB9A9" box="[1038,1147,1767,1789]" italics="true" pageId="11" pageNumber="10">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="E386EADAFFA3C26BFB7CF94CFB03B9A9" bold="true" box="[1199,1223,1768,1789]" pageId="11" pageNumber="10">18</emphasis>
|
||
=
|
||
<taxonomicName id="16F24D4BFFA3C26BFB33F94CFAF2B9A9" authority="Ayrey & Soleglad, 2015" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[1248,1334,1768,1789]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="troupi" status="sp. nov.">
|
||
<emphasis id="E386EADAFFA3C26BFB33F94CFAF2B9A9" box="[1248,1334,1768,1789]" italics="true" pageId="11" pageNumber="10">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F8B557A1FFA3C26BFAE8F94CFAB5B9A9" box="[1339,1393,1768,1789]" pageId="11" pageNumber="10" rank="species">
|
||
<emphasis id="E386EADAFFA3C26BFAE8F94CFAAFB9A9" bold="true" box="[1339,1387,1768,1789]" pageId="11" pageNumber="10">sp. n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D14D36C8FFA4C26CFE18FF6DFA4EBE8D" pageId="12" pageNumber="11">
|
||
<table id="A3F2C468FFA43D9FFF19FF6CFA4EBE8E" box="[202,1418,200,474]" gridcols="4" gridrows="6" pageId="12" pageNumber="11">
|
||
<tr id="6FC2348AFFA43D9FFF19FF6CFA4EBFB7" box="[202,1418,200,227]" gridrow="0" pageId="12" pageNumber="11" rowspan-0="1">
|
||
<th id="2C135DF6FFA43D9FFE51FF6CFDB9BFB7" box="[386,637,200,227]" gridcol="1" gridrow="0" pageId="12" pageNumber="11">
|
||
<taxonomicName id="16F24D4BFFA4C26CFE18FF6DFDF3BFB7" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[459,567,201,227]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFE18FF6DFDF3BFB7" bold="true" box="[459,567,201,227]" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="2C135DF6FFA43D9FFD48FF6CFC39BFB7" box="[667,1021,200,227]" gridcol="2" gridrow="0" pageId="12" pageNumber="11">
|
||
<taxonomicName id="16F24D4BFFA4C26CFCDCFF6CFC5CBFB6" box="[783,920,200,226]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA4C26CFCDCFF6CFC5CBFB6" bold="true" box="[783,920,200,226]" italics="true" pageId="12" pageNumber="11">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="2C135DF6FFA43D9FFBFBFF6CFA4EBFB7" box="[1064,1418,200,227]" gridcol="3" gridrow="0" pageId="12" pageNumber="11">
|
||
<taxonomicName id="16F24D4BFFA4C26CFB4DFF6DFAE4BFB7" authorityName=": Ayrey" authorityYear="2013" box="[1182,1312,201,227]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFB4DFF6DFAE4BFB7" bold="true" box="[1182,1312,201,227]" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="6FC2348AFFA43D9FFF19FF4BFA4EBE5E" box="[202,1418,239,266]" gridrow="1" pageId="12" pageNumber="11">
|
||
<th id="2C135DF6FFA43D9FFF19FF4BFE99BE5E" box="[202,349,239,266]" gridcol="0" gridrow="1" pageId="12" pageNumber="11">
|
||
<emphasis id="E386EADAFFA4C26CFF19FF56FE8DBE5E" bold="true" box="[202,329,242,266]" pageId="12" pageNumber="11">Seg-I: L/W</emphasis>
|
||
</th>
|
||
<td id="2C135DF6FFA43D9FFE51FF4BFDB9BE5E" box="[386,637,239,266]" gridcol="1" gridrow="1" pageId="12" pageNumber="11">
|
||
0.917>
|
||
<emphasis id="E386EADAFFA4C26CFE08FF4BFE31BE57" bold="true" box="[475,501,239,259]" italics="true" pageId="12" pageNumber="11"></emphasis>
|
||
16.6, 15.6 %
|
||
</td>
|
||
<td id="2C135DF6FFA43D9FFD48FF4BFC39BE5E" box="[667,1021,239,266]" gridcol="2" gridrow="1" pageId="12" pageNumber="11">0.744–0.854 (0.787) (±0.059) [3]</td>
|
||
<td id="2C135DF6FFA43D9FFBFBFF4BFA4EBE5E" box="[1064,1418,239,266]" gridcol="3" gridrow="1" pageId="12" pageNumber="11">0.778–0.811 (0.794) (±0.015) [4]</td>
|
||
</tr>
|
||
<tr id="6FC2348AFFA43D9FFF19FE87FA4EBE6A" box="[202,1418,291,318]" gridrow="2" pageId="12" pageNumber="11">
|
||
<th id="2C135DF6FFA43D9FFF19FE87FE99BE6A" box="[202,349,291,318]" gridcol="0" gridrow="2" pageId="12" pageNumber="11">
|
||
<emphasis id="E386EADAFFA4C26CFF19FE82FE97BE6A" bold="true" box="[202,339,294,318]" pageId="12" pageNumber="11">Seg-II: L/W</emphasis>
|
||
</th>
|
||
<td id="2C135DF6FFA43D9FFE51FE87FDB9BE6A" box="[386,637,291,318]" gridcol="1" gridrow="2" pageId="12" pageNumber="11">
|
||
1.086:>
|
||
<emphasis id="E386EADAFFA4C26CFE0FFE87FE32BE63" bold="true" box="[476,502,291,311]" italics="true" pageId="12" pageNumber="11"></emphasis>
|
||
14.6, 15.0 %
|
||
</td>
|
||
<td id="2C135DF6FFA43D9FFD48FE87FC39BE6A" box="[667,1021,291,318]" gridcol="2" gridrow="2" pageId="12" pageNumber="11">0.946–0.949 (0.948) (±0.002) [3]</td>
|
||
<td id="2C135DF6FFA43D9FFBFBFE87FA4EBE6A" box="[1064,1418,291,318]" gridcol="3" gridrow="2" pageId="12" pageNumber="11">0.923–0.971 (0.945) (±0.020) [4]</td>
|
||
</tr>
|
||
<tr id="6FC2348AFFA43D9FFF19FEF3FA4EBE26" box="[202,1418,343,370]" gridrow="3" pageId="12" pageNumber="11">
|
||
<th id="2C135DF6FFA43D9FFF19FEF3FE99BE26" box="[202,349,343,370]" gridcol="0" gridrow="3" pageId="12" pageNumber="11">
|
||
<emphasis id="E386EADAFFA4C26CFF19FEFEFE99BE26" bold="true" box="[202,349,346,370]" pageId="12" pageNumber="11">Seg-III: L/W</emphasis>
|
||
</th>
|
||
<td id="2C135DF6FFA43D9FFE51FEF3FDB9BE26" box="[386,637,343,370]" gridcol="1" gridrow="3" pageId="12" pageNumber="11">
|
||
1.176:>
|
||
<emphasis id="E386EADAFFA4C26CFE0FFEF3FE32BE3F" bold="true" box="[476,502,343,363]" italics="true" pageId="12" pageNumber="11"></emphasis>
|
||
13.5, 10.6 %
|
||
</td>
|
||
<td id="2C135DF6FFA43D9FFD48FEF3FC39BE26" box="[667,1021,343,370]" gridcol="2" gridrow="3" pageId="12" pageNumber="11">1.026–1.054 (1.036) (±0.016) [3]</td>
|
||
<td id="2C135DF6FFA43D9FFBFBFEF3FA4EBE26" box="[1064,1418,343,370]" gridcol="3" gridrow="3" pageId="12" pageNumber="11">1.017–1.091 (1.064) (±0.034) [4]</td>
|
||
</tr>
|
||
<tr id="6FC2348AFFA43D9FFF19FE2FFA4EBEF2" box="[202,1418,395,422]" gridrow="4" pageId="12" pageNumber="11">
|
||
<th id="2C135DF6FFA43D9FFF19FE2FFE99BEF2" box="[202,349,395,422]" gridcol="0" gridrow="4" pageId="12" pageNumber="11">
|
||
<emphasis id="E386EADAFFA4C26CFF19FE2AFE98BEF2" bold="true" box="[202,348,398,422]" pageId="12" pageNumber="11">Seg-IV: L/W</emphasis>
|
||
</th>
|
||
<td id="2C135DF6FFA43D9FFE51FE2FFDB9BEF2" box="[386,637,395,422]" gridcol="1" gridrow="4" pageId="12" pageNumber="11">
|
||
1.688:>
|
||
<emphasis id="E386EADAFFA4C26CFE0FFE2FFE32BECB" bold="true" box="[476,502,395,415]" italics="true" pageId="12" pageNumber="11"></emphasis>
|
||
13.1, 7.7 %
|
||
</td>
|
||
<td id="2C135DF6FFA43D9FFD48FE2FFC39BEF2" box="[667,1021,395,422]" gridcol="2" gridrow="4" pageId="12" pageNumber="11">1.432–1.545 (1.492) (±0.057) [3]</td>
|
||
<td id="2C135DF6FFA43D9FFBFBFE2FFA4EBEF2" box="[1064,1418,395,422]" gridcol="3" gridrow="4" pageId="12" pageNumber="11">1.527–1.613 (1.567) (±0.038) [4]</td>
|
||
</tr>
|
||
<tr id="6FC2348AFFA43D9FFF19FE1BFA4EBE8E" box="[202,1418,447,474]" gridrow="5" pageId="12" pageNumber="11">
|
||
<th id="2C135DF6FFA43D9FFF19FE1BFE99BE8E" box="[202,349,447,474]" gridcol="0" gridrow="5" pageId="12" pageNumber="11">
|
||
<emphasis id="E386EADAFFA4C26CFF19FE66FE96BE8E" bold="true" box="[202,338,450,474]" pageId="12" pageNumber="11">Seg-V: L/W</emphasis>
|
||
</th>
|
||
<td id="2C135DF6FFA43D9FFE51FE1BFDB9BE8E" box="[386,637,447,474]" gridcol="1" gridrow="5" pageId="12" pageNumber="11">
|
||
2.433:>
|
||
<emphasis id="E386EADAFFA4C26CFE0FFE1BFE32BE87" bold="true" box="[476,502,447,467]" italics="true" pageId="12" pageNumber="11"></emphasis>
|
||
7.7, 7.6 %
|
||
</td>
|
||
<td id="2C135DF6FFA43D9FFD48FE1BFC39BE8E" box="[667,1021,447,474]" gridcol="2" gridrow="5" pageId="12" pageNumber="11">2.182–2.364 (2.258) (±0.094) [3]</td>
|
||
<td id="2C135DF6FFA43D9FFBFBFE1BFA4EBE8E" box="[1064,1418,447,474]" gridcol="3" gridrow="5" pageId="12" pageNumber="11">2.194–2.333 (2.262) (±0.062) [4]</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="858D6640FFA4C26CFF13FDA8FAA2BD3A" pageId="12" pageNumber="11" startId="12.[192,257,524,548]" targetBox="[202,1418,200,474]" targetIsTable="true" targetPageId="12">
|
||
<paragraph id="D14D36C8FFA4C26CFF13FDA8FAA2BD3A" blockId="12.[192,1441,524,622]" pageId="12" pageNumber="11">
|
||
<emphasis id="E386EADAFFA4C26CFF13FDA8FEE5BD70" bold="true" box="[192,289,524,548]" pageId="12" pageNumber="11">Table 2:</emphasis>
|
||
Comparison of morphometric ratios of the metasoma based on the measurements of female specimens presented in Table 1. In these ratios,
|
||
<taxonomicName id="16F24D4BFFA4C26CFE7CFD83FD81BD68" authority="Ayrey & Soleglad, 2015" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[431,581,551,572]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi" status="sp. nov.">
|
||
<emphasis id="E386EADAFFA4C26CFE7CFD83FD81BD68" box="[431,581,551,572]" italics="true" pageId="12" pageNumber="11">Vaejovis troupi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E386EADAFFA4C26CFD9FFD83FD40BD68" bold="true" box="[588,644,551,572]" pageId="12" pageNumber="11">
|
||
<taxonomicNameLabel id="F8B557A1FFA4C26CFD9FFD83FD40BD68" box="[588,644,551,572]" pageId="12" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
exhibits the more slender metasoma when compared to the other two species (as indicated by a larger ratio value), ranging from 8 to over 16 percent mean value difference (MVD). Statistical data group: minimum–maximum (mean) (±standard deviation) [number of samples]. Seg = metasomal segment, L = length, W = width.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D14D36C8FFA4C26CFF23FD7BFECCB906" blockId="12.[192,802,674,1894]" pageId="12" pageNumber="11">
|
||
<emphasis id="E386EADAFFA4C26CFF23FD7BFDD9BDA3" bold="true" box="[240,541,735,759]" pageId="12" pageNumber="11">Molecular considerations.</emphasis>
|
||
In an important study conducted by Bryson et al. (2013) using multilocus data (mtDNA and two nuclear genes), they explored the phylogeographic structure of the “
|
||
<taxonomicName id="16F24D4BFFA4C26CFDEEFC9FFD57BC07" authorityName=": Ayrey" authorityYear="2013" box="[573,659,827,851]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFDEEFC9FFD57BC07" box="[573,659,827,851]" italics="true" pageId="12" pageNumber="11">vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
” group. The dataset was generated from 63 samples of “
|
||
<taxonomicName id="16F24D4BFFA4C26CFD6CFCFEFCD1BC26" authorityName=": Ayrey" authorityYear="2013" box="[703,789,858,882]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFD6CFCFEFCD1BC26" box="[703,789,858,882]" italics="true" pageId="12" pageNumber="11">vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
” group scorpions including 11 of the then described species as well as many other populations. Important to this paper, the dataset included species
|
||
<taxonomicName id="16F24D4BFFA4C26CFDEDFC13FD77BC9A" authorityName=": Ayrey" authorityYear="2013" box="[574,691,950,974]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFDEDFC13FD77BC9A" box="[574,691,950,974]" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
and specimens from two populations, the Santa Rita Mountains, now named
|
||
<taxonomicName id="16F24D4BFFA4C26CFE9BFC50FE07BB5F" box="[328,451,1011,1035]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA4C26CFE9BFC50FE07BB5F" box="[328,451,1011,1035]" italics="true" pageId="12" pageNumber="11">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
, and the Whetstone Mountains, the location of
|
||
<taxonomicName id="16F24D4BFFA4C26CFEBEFBB7FE14BB7E" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[365,464,1043,1066]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFEBEFBB7FE14BB7E" box="[365,464,1043,1066]" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
, the new species described in this paper. In their study divergence times were estimated across the mtDNA dataset and the multilocus species tree, and the temporal distribution of divergence events was modeled across southwestern North America. The result of their study placed
|
||
<taxonomicName id="16F24D4BFFA4C26CFDF8FB08FD63BB97" box="[555,679,1195,1219]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA4C26CFDF8FB08FD63BB97" box="[555,679,1195,1219]" italics="true" pageId="12" pageNumber="11">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="16F24D4BFFA4C26CFD6AFB08FCDFBB97" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[697,795,1196,1219]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFD6AFB08FCDFBB97" box="[697,795,1196,1219]" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="16F24D4BFFA4C26CFF22FB6FFEADBBB6" authorityName=": Ayrey" authorityYear="2013" box="[241,361,1226,1250]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFF22FB6FFEADBBB6" box="[241,361,1226,1250]" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
in a terminal clade, implying that they were the most closely related. Both the multilocus species tree and mtDNA gene tree analyses supported this result. Their chronograms (Bryson et al., 2013: fig. S2) showed the estimated divergence times (in millions of years, Mya) as follows: for the multilocus species tree, 3.25–8.48 (5.75) Mya for
|
||
<taxonomicName id="16F24D4BFFA4C26CFD93FA27FD06BACE" box="[576,706,1410,1434]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA4C26CFD93FA27FD06BACE" box="[576,706,1410,1434]" italics="true" pageId="12" pageNumber="11">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA4C26CFCD8FA27FED2BAEC" authorityName=": Ayrey" authorityYear="2013" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFCD8FA27FED2BAEC" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="16F24D4BFFA4C26CFEE5FA05FE52BAEC" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[310,406,1441,1464]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFEE5FA05FE52BAEC" box="[310,406,1441,1464]" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
, and for the mtDNA gene tree, 3.3– 8.07 (5.45) Mya for
|
||
<taxonomicName id="16F24D4BFFA4C26CFE7CFA64FDE9BA83" box="[431,557,1471,1495]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA4C26CFE7CFA64FDE9BA83" box="[431,557,1471,1495]" italics="true" pageId="12" pageNumber="11">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA4C26CFDB8FA64FD27BA83" authorityName=": Ayrey" authorityYear="2013" box="[619,739,1471,1495]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFDB8FA64FD27BA83" box="[619,739,1471,1495]" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="16F24D4BFFA4C26CFCD8FA64FEC5BAA2" authorityName="Ayrey & Soleglad" authorityYear="2015" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFCD8FA64FEC5BAA2" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
. Similarly, multilocus species tree, 1.05–5.12 (2.98) Mya for
|
||
<taxonomicName id="16F24D4BFFA4C26CFEA2FA59FE2DB940" authorityName=": Ayrey" authorityYear="2013" box="[369,489,1532,1556]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFEA2FA59FE2DB940" box="[369,489,1532,1556]" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA4C26CFDF5FA59FD4CB940" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[550,648,1533,1556]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFDF5FA59FD4CB940" box="[550,648,1533,1556]" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
, and mtDNA gene tree, 1.53–5.80 (3.53) Mya for
|
||
<taxonomicName id="16F24D4BFFA4C26CFD8AF9B8FD14B967" authorityName=": Ayrey" authorityYear="2013" box="[601,720,1563,1587]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFD8AF9B8FD14B967" box="[601,720,1563,1587]" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA4C26CFCD8F9B8FEC5B906" authorityName="Ayrey & Soleglad" authorityYear="2015" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFCD8F9B8FEC5B906" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFA4C26CFF23F9FDFD7AB9BF" blockId="12.[192,802,674,1894]" pageId="12" pageNumber="11">
|
||
It should also be noted that the Bryson et al. (2013) study placed species
|
||
<taxonomicName id="16F24D4BFFA4C26CFE7AF9DCFDE8B9DB" authorityName="Ayrey" authorityYear="2009" box="[425,556,1655,1679]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="deboerae">
|
||
<emphasis id="E386EADAFFA4C26CFE7AF9DCFDE8B9DB" box="[425,556,1655,1679]" italics="true" pageId="12" pageNumber="11">V. deboerae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA4C26CFDB4F9DCFCC7B9DB" box="[615,771,1655,1679]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="tenuipalpus">
|
||
<emphasis id="E386EADAFFA4C26CFDB4F9DCFCC7B9DB" box="[615,771,1655,1679]" italics="true" pageId="12" pageNumber="11">V. tenuipalpus</emphasis>
|
||
</taxonomicName>
|
||
in the same basic clade as our three subject species, but earlier chronologically, exhibiting 4.45–11.08 (7.71) Mya and 8.07–18.19 (12.78) Mya, respectively.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFA4C26CFF23F956FB8ABC26" blockId="12.[192,802,674,1894]" lastBlockId="12.[831,1441,674,1894]" pageId="12" pageNumber="11">
|
||
<emphasis id="E386EADAFFA4C26CFF23F956FD94B85E" bold="true" box="[240,592,1778,1802]" pageId="12" pageNumber="11">Morphological considerations.</emphasis>
|
||
We have isolated three morphology-based characters that distinguish
|
||
<taxonomicName id="16F24D4BFFA4C26CFCD9F8B5FEC5B813" authorityName="Ayrey & Soleglad" authorityYear="2015" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFCD9F8B5FEC5B813" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
from species
|
||
<taxonomicName id="16F24D4BFFA4C26CFE62F894FDF6B813" box="[433,562,1839,1863]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA4C26CFE62F894FDF6B813" box="[433,562,1839,1863]" italics="true" pageId="12" pageNumber="11">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA4C26CFDABF894FD31B813" authorityName=": Ayrey" authorityYear="2013" box="[632,757,1839,1863]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFDABF894FD31B813" box="[632,757,1839,1863]" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
: 1) number of inner denticles (
|
||
<emphasis id="E386EADAFFA4C26CFE38F8EBFDC3B832" box="[491,519,1871,1894]" italics="true" pageId="12" pageNumber="11">ID</emphasis>
|
||
) on the chelal fingers; 2) comparative positions of certain patellar trichobothria; and 3) morphometric ratios of the metasomal segments. Of course, it must be emphasized here that only one specimen of
|
||
<taxonomicName id="16F24D4BFFA4C26CFC03FD5BFBF0BC42" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[976,1076,767,790]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFC03FD5BFBF0BC42" box="[976,1076,767,790]" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
, a female, is currently available for examination, and the male is unknown. Therefore, some variability should be expected as more specimens are collected and studied.
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFA4C26CFCBCFCDDFA52BAA2" blockId="12.[831,1441,674,1894]" pageId="12" pageNumber="11">
|
||
All three species exhibit six
|
||
<emphasis id="E386EADAFFA4C26CFB07FCDEFB34BCC5" box="[1236,1264,890,913]" italics="true" pageId="12" pageNumber="11">ID</emphasis>
|
||
on the chelal movable finger, also common to eight other species in the “
|
||
<taxonomicName id="16F24D4BFFA4C26CFCA2FC12FC03BC9A" authorityName=": Ayrey" authorityYear="2013" box="[881,967,950,974]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFCA2FC12FC03BC9A" box="[881,967,950,974]" italics="true" pageId="12" pageNumber="11">vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
” group (see map in
|
||
<figureCitation id="49C92A4DFFA4C26CFB72FC12FB2BBC9A" box="[1185,1263,950,974]" captionStart="Figure 11" captionStartId="11.[192,267,1642,1666]" captionTargetBox="[191,1441,191,1603]" captionTargetId="figure-171@11.[192,1440,192,1603]" captionTargetPageId="11" captionText="Figure 11: Map of Arizona, extreme western New Mexico and northern Sonora (Mexico) showing the type locality of all 17 Vaejovis “vorhiesi” group species, including new species Vaejovis troupi. Localities are divided into those species exhibiting seven inner denticles (ID) on the chelal movable finger (white rectangles with black lettering) and those with primarily six, or five ID denticles (black rectangles with white lettering). Seven IDs: 1 = V. jonesi, 2 = V. lapidicola, 3 = V. paysonensis, 4 = V. crumpi, 5 = V. bigelowi, 14 = V. trinityae, 17 = V. grayae. Six IDs: 6 = V. vorhiesi, 7 = V. cashi, 8 = V. feti, 9 = V. deboerae, 10 = V. electrum, 11 = V. tenuipalpus 12 = V. halli, 13 = V. brysoni, 15 = V. bandido, 16 = V. grahami, and 18 = V. troupi sp. n." figureDoi="http://doi.org/10.5281/zenodo.5508080" httpUri="https://zenodo.org/record/5508080/files/figure.png" pageId="12" pageNumber="11">Fig. 11</figureCitation>
|
||
). Unusual, however, are the six
|
||
<emphasis id="E386EADAFFA4C26CFC24FC71FBD7BCB8" box="[1015,1043,981,1004]" italics="true" pageId="12" pageNumber="11">ID</emphasis>
|
||
also found on the fixed finger in
|
||
<taxonomicName id="16F24D4BFFA4C26CFA5AFC71FC44BB5F" authorityName="Ayrey & Soleglad" authorityYear="2015" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFA5AFC71FC44BB5F" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
(found on both chelae), somewhat unusual in this species subgroup that only have six
|
||
<emphasis id="E386EADAFFA4C26CFB02FBB7FB29BB7E" box="[1233,1261,1043,1066]" italics="true" pageId="12" pageNumber="11">ID</emphasis>
|
||
on the movable finger. In contrast,
|
||
<taxonomicName id="16F24D4BFFA4C26CFBCAFB96FB52BB1D" box="[1049,1174,1073,1097]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA4C26CFBCAFB96FB52BB1D" box="[1049,1174,1073,1097]" italics="true" pageId="12" pageNumber="11">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA4C26CFB06FB96FA8ABB1D" authorityName=": Ayrey" authorityYear="2013" box="[1237,1358,1073,1097]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFB06FB96FA8ABB1D" box="[1237,1358,1073,1097]" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
, on an average, only have five
|
||
<emphasis id="E386EADAFFA4C26CFB94FBF4FBA7BB33" box="[1095,1123,1104,1127]" italics="true" pageId="12" pageNumber="11">ID</emphasis>
|
||
on the fixed finger (see
|
||
<figureCitation id="49C92A4DFFA4C26CFAA5FBEBFC8ABBD2" captionStart="Figure 8" captionStartId="8.[1035,1110,829,853]" captionTargetBox="[195,961,205,1104]" captionTargetId="figure-13@8.[180,1011,194,1104]" captionTargetPageId="8" captionText="Figure 8: Chelal fixed finger dentition showing the number of inner denticles (ID). Left. Vaejovis troupi sp. n., holotype female (left fixed finger, reversed). Center. V. grahami, Madera Canyon, Santa Rita Mountains. Right. V. vorhiesi, Miller’s Canyon, Huachuca Mountains. Of particular interest, note that V. troupi has six ID denticles whereas the other two species exhibit only five ID denticles. Internal trichobothria ib and it are shown." figureDoi="http://doi.org/10.5281/zenodo.5508074" httpUri="https://zenodo.org/record/5508074/files/figure.png" pageId="12" pageNumber="11">Fig. 8</figureCitation>
|
||
). Other species contained in the species set with six
|
||
<emphasis id="E386EADAFFA4C26CFA57FBCBFA64BBD2" box="[1412,1440,1135,1158]" italics="true" pageId="12" pageNumber="11">ID</emphasis>
|
||
on the movable finger, only
|
||
<taxonomicName id="16F24D4BFFA4C26CFBBCFB29FA64BBF1" authority=", Sissom et al." authorityName="Sissom" box="[1135,1440,1164,1189]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="tenuipalpus">
|
||
<emphasis id="E386EADAFFA4C26CFBBCFB29FACEBBF0" box="[1135,1290,1164,1188]" italics="true" pageId="12" pageNumber="11">V. tenuipalpus</emphasis>
|
||
, Sissom et al.
|
||
</taxonomicName>
|
||
(2012: fig. 4D) and
|
||
<taxonomicName id="16F24D4BFFA4C26CFBCDFB08FB32BB97" authority="Hughes" authorityName="Hughes" authorityYear="2011" box="[1054,1270,1195,1219]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="electrum">
|
||
<emphasis id="E386EADAFFA4C26CFBCDFB08FB58BB97" box="[1054,1180,1195,1219]" italics="true" pageId="12" pageNumber="11">V. electrum</emphasis>
|
||
Hughes
|
||
</taxonomicName>
|
||
(2011: fig. 8d) match
|
||
<taxonomicName id="16F24D4BFFA4C26CFC5BFB6FFC23BBB6" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[904,999,1227,1250]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFC5BFB6FFC23BBB6" box="[904,999,1227,1250]" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
with six
|
||
<emphasis id="E386EADAFFA4C26CFB9DFB6FFBAEBBB6" box="[1102,1130,1227,1250]" italics="true" pageId="12" pageNumber="11">ID</emphasis>
|
||
on the fixed finger, as these authors illustrated for the type specimens. It should be stated here, however, that variability does occur on the number of
|
||
<emphasis id="E386EADAFFA4C26CFC6FFA83FC1CBA6A" box="[956,984,1319,1342]" italics="true" pageId="12" pageNumber="11">ID</emphasis>
|
||
found on the fixed finger for “vorhiesi” group species. For example, Sissom (2012: 12) reported 4–6 (5.732) [56] for
|
||
<taxonomicName id="16F24D4BFFA4C26CFBF0FAC0FB7BBA2F" box="[1059,1215,1379,1403]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="tenuipalpus">
|
||
<emphasis id="E386EADAFFA4C26CFBF0FAC0FB7BBA2F" box="[1059,1215,1379,1403]" italics="true" pageId="12" pageNumber="11">V. tenuipalpus</emphasis>
|
||
</taxonomicName>
|
||
; for
|
||
<taxonomicName id="16F24D4BFFA4C26CFB2BFAC0FAB6BA2F" box="[1272,1394,1379,1403]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA4C26CFB2BFAC0FAB6BA2F" box="[1272,1394,1379,1403]" italics="true" pageId="12" pageNumber="11">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
, six specimens, we encountered significant variability, 5–6 (5.417) [12]; however, in stark contrast,
|
||
<taxonomicName id="16F24D4BFFA4C26CFAF7FA05FA64BAEC" authorityName=": Ayrey" authorityYear="2013" box="[1316,1440,1440,1464]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFAF7FA05FA64BAEC" box="[1316,1440,1440,1464]" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
exhibited 5
|
||
<emphasis id="E386EADAFFA4C26CFC15FA64FC26BA83" box="[966,994,1472,1495]" italics="true" pageId="12" pageNumber="11">ID</emphasis>
|
||
on the fixed finger for four specimens (including
|
||
<typeStatus id="0E49886AFFA4C26CFC60FA7AFBD0BAA2" box="[947,1044,1502,1526]" pageId="12" pageNumber="11" type="lectotype">lectotype</typeStatus>
|
||
as described by Graham (2007: 6)).
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFA4C26DFCBCFA59FD2ABD9C" blockId="12.[831,1441,674,1894]" lastBlockId="13.[192,802,198,1019]" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11">
|
||
We compared the relative positions of five external patellar trichobothria
|
||
<emphasis id="E386EADAFFA4C26CFBE2F9B8FBB3B967" box="[1073,1143,1564,1587]" italics="true" pageId="12" pageNumber="11">
|
||
et
|
||
<subScript id="4D76348DFFA4C26CFB97F981FB89B967" attach="left" box="[1092,1101,1573,1587]" fontSize="6" pageId="12" pageNumber="11">1</subScript>
|
||
–et
|
||
<subScript id="4D76348DFFA4C26CFBBDF981FBB3B967" attach="left" box="[1134,1143,1573,1587]" fontSize="6" pageId="12" pageNumber="11">3</subScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="E386EADAFFA4C26CFB59F9B8FB63B967" box="[1162,1191,1564,1587]" italics="true" pageId="12" pageNumber="11">est</emphasis>
|
||
, and
|
||
<emphasis id="E386EADAFFA4C26CFB3EF9B8FB3DB967" box="[1261,1273,1564,1587]" italics="true" pageId="12" pageNumber="11">v</emphasis>
|
||
<quantity id="160A9B2DFFA4C26CFB2AF981FAE7B967" box="[1273,1315,1563,1587]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="11" unit="in" value="3.0">
|
||
<emphasis id="E386EADAFFA4C26CFB2AF981FAC6B967" box="[1273,1282,1573,1587]" italics="true" pageId="12" pageNumber="11">
|
||
<subScript id="4D76348DFFA4C26CFB2AF981FAC6B967" attach="left" box="[1273,1282,1573,1587]" fontSize="6" pageId="12" pageNumber="11">3</subScript>
|
||
</emphasis>
|
||
in
|
||
</quantity>
|
||
species
|
||
<taxonomicName id="16F24D4BFFA4C26CFA5AF9B8FC44B906" authorityName="Ayrey & Soleglad" authorityYear="2015" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFA5AF9B8FC44B906" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="16F24D4BFFA4C26CFC41F99FFBCBB906" box="[914,1039,1594,1618]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA4C26CFC41F99FFBCBB906" box="[914,1039,1594,1618]" italics="true" pageId="12" pageNumber="11">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="16F24D4BFFA4C26CFB80F99FFB08B906" authorityName=": Ayrey" authorityYear="2013" box="[1107,1228,1594,1618]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFB80F99FFB08B906" box="[1107,1228,1594,1618]" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="49C92A4DFFA4C26CFADDF99EFA97B906" box="[1294,1363,1594,1618]" captionStart="Figure 9" captionStartId="9.[192,267,1072,1096]" captionTargetBox="[191,1441,191,1038]" captionTargetId="figure-509@9.[192,1440,192,1037]" captionTargetPageId="9" captionText="Figure 9: Comparison of the relative positions of five patellar trichobothria et1–et3, est, and v3 of species Vaejovis troupi sp. n., V. grahami, and V. vorhiesi. Top. Trichobothria positions of the three female type specimens. Red lines indicate the angle formed by trichobothria est–et3–v3, contrasting V. troupi from the other two species. Bottom. Each individual pattern for a pedipalp patella is anchored at trichobothrium et1 while other trichobothria are plotted at their relative locations forming clusters. Red lines indicate the average angle formed by trichobothria est–et3–v3. Patterns of left patellae were reversed for these plots which are shown from a right-patella perspective. Only adult female specimens were used in these plots. Number below species name indicates the number of pedipalp patellae considered. Information for Vaejovis vorhiesi and V. grahami lectotype/holotype females was extracted from Graham (2007: fig. 9) and Ayrey & Soleglad (2014: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.5508076" httpUri="https://zenodo.org/record/5508076/files/figure.png" pageId="12" pageNumber="11">Fig. 9</figureCitation>
|
||
). Both patellae of
|
||
<taxonomicName id="16F24D4BFFA4C26CFC6AF9FDFBDDB924" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[953,1049,1625,1648]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA4C26CFC6AF9FDFBDDB924" box="[953,1049,1625,1648]" italics="true" pageId="12" pageNumber="11">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
and thirteen patellae of
|
||
<taxonomicName id="16F24D4BFFA4C26CFAF5F9FDFA64B924" box="[1318,1440,1624,1648]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA4C26CFAF5F9FDFA64B924" box="[1318,1440,1624,1648]" italics="true" pageId="12" pageNumber="11">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA4C26CFCA3F9DCFC23B9DB" authorityName=": Ayrey" authorityYear="2013" box="[880,999,1655,1679]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA4C26CFCA3F9DCFC23B9DB" box="[880,999,1655,1679]" italics="true" pageId="12" pageNumber="11">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
were included in this analysis, demonstrating little variability in the relative positions of these five trichobothria. In
|
||
<figureCitation id="49C92A4DFFA4C26CFBE1F911FB54B999" box="[1074,1168,1717,1741]" captionStart="Figure 9" captionStartId="9.[192,267,1072,1096]" captionTargetBox="[191,1441,191,1038]" captionTargetId="figure-509@9.[192,1440,192,1037]" captionTargetPageId="9" captionText="Figure 9: Comparison of the relative positions of five patellar trichobothria et1–et3, est, and v3 of species Vaejovis troupi sp. n., V. grahami, and V. vorhiesi. Top. Trichobothria positions of the three female type specimens. Red lines indicate the angle formed by trichobothria est–et3–v3, contrasting V. troupi from the other two species. Bottom. Each individual pattern for a pedipalp patella is anchored at trichobothrium et1 while other trichobothria are plotted at their relative locations forming clusters. Red lines indicate the average angle formed by trichobothria est–et3–v3. Patterns of left patellae were reversed for these plots which are shown from a right-patella perspective. Only adult female specimens were used in these plots. Number below species name indicates the number of pedipalp patellae considered. Information for Vaejovis vorhiesi and V. grahami lectotype/holotype females was extracted from Graham (2007: fig. 9) and Ayrey & Soleglad (2014: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.5508076" httpUri="https://zenodo.org/record/5508076/files/figure.png" pageId="12" pageNumber="11">Figure 9</figureCitation>
|
||
we plotted the five trichobothria by anchoring each patellar pattern to trichobothrium
|
||
<emphasis id="E386EADAFFA4C26CFC1EF957FC2EB85E" box="[973,1002,1779,1802]" italics="true" pageId="12" pageNumber="11">
|
||
et
|
||
<subScript id="4D76348DFFA4C26CFC32F958FC2EB85E" attach="right" box="[993,1002,1788,1802]" fontSize="6" pageId="12" pageNumber="11">1</subScript>
|
||
</emphasis>
|
||
, allowing the other trichobothria to form clusters. It is apparent that these trichobothria in general form fairly tight clusters, thus demonstrating minimal variability in their relative positions. In
|
||
<figureCitation id="49C92A4DFFA4C26CFB3DF8EAFA8EB832" box="[1262,1354,1870,1894]" captionStart="Figure 9" captionStartId="9.[192,267,1072,1096]" captionTargetBox="[191,1441,191,1038]" captionTargetId="figure-509@9.[192,1440,192,1037]" captionTargetPageId="9" captionText="Figure 9: Comparison of the relative positions of five patellar trichobothria et1–et3, est, and v3 of species Vaejovis troupi sp. n., V. grahami, and V. vorhiesi. Top. Trichobothria positions of the three female type specimens. Red lines indicate the angle formed by trichobothria est–et3–v3, contrasting V. troupi from the other two species. Bottom. Each individual pattern for a pedipalp patella is anchored at trichobothrium et1 while other trichobothria are plotted at their relative locations forming clusters. Red lines indicate the average angle formed by trichobothria est–et3–v3. Patterns of left patellae were reversed for these plots which are shown from a right-patella perspective. Only adult female specimens were used in these plots. Number below species name indicates the number of pedipalp patellae considered. Information for Vaejovis vorhiesi and V. grahami lectotype/holotype females was extracted from Graham (2007: fig. 9) and Ayrey & Soleglad (2014: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.5508076" httpUri="https://zenodo.org/record/5508076/files/figure.png" pageId="12" pageNumber="11">Figure 9</figureCitation>
|
||
, we see that
|
||
<emphasis id="E386EADAFFA5C26DFF26FF63FECCBF8A" box="[245,264,199,222]" italics="true" pageId="13" pageNumber="12">et</emphasis>
|
||
<quantity id="160A9B2DFFA5C26DFEDBFF74FEF0BF8A" box="[264,308,198,222]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="13" pageNumber="12" unit="in" value="3.0">
|
||
<emphasis id="E386EADAFFA5C26DFEDBFF74FED5BF8A" box="[264,273,208,222]" italics="true" pageId="13" pageNumber="12">
|
||
<subScript id="4D76348DFFA5C26DFEDBFF74FED5BF8A" attach="left" box="[264,273,208,222]" fontSize="6" pageId="13" pageNumber="12">3</subScript>
|
||
</emphasis>
|
||
in
|
||
</quantity>
|
||
<taxonomicName id="16F24D4BFFA5C26DFE92FF63FE62BF8A" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[321,422,199,222]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA5C26DFE92FF63FE62BF8A" box="[321,422,199,222]" italics="true" pageId="13" pageNumber="12">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
is noticeably proximal to a line connecting
|
||
<emphasis id="E386EADAFFA5C26DFE81FF41FEB4BFA8" box="[338,368,229,252]" italics="true" pageId="13" pageNumber="12">est</emphasis>
|
||
and
|
||
<emphasis id="E386EADAFFA5C26DFE02FF41FE22BFA9" box="[465,486,229,253]" italics="true" pageId="13" pageNumber="12">
|
||
v
|
||
<subScript id="4D76348DFFA5C26DFE0EFF4BFE22BFA9" attach="left" box="[477,486,239,253]" fontSize="6" pageId="13" pageNumber="12">3</subScript>
|
||
</emphasis>
|
||
, forming an angle of approximately 130 degrees. In the other two species
|
||
<emphasis id="E386EADAFFA5C26DFCD5FEA0FCE6BE4F" box="[774,802,260,283]" italics="true" pageId="13" pageNumber="12">
|
||
et
|
||
<subScript id="4D76348DFFA5C26DFCCAFEA9FCE6BE4F" attach="left" box="[793,802,269,283]" fontSize="6" pageId="13" pageNumber="12">3</subScript>
|
||
</emphasis>
|
||
is essentially located in this connecting line, thus the angle is quite obtuse, over 165 degrees. This difference in the relative pattern can be attributed to a couple of observations. In general the distance between
|
||
<emphasis id="E386EADAFFA5C26DFD60FEDBFD14BEC2" box="[691,720,383,406]" italics="true" pageId="13" pageNumber="12">
|
||
et
|
||
<subScript id="4D76348DFFA5C26DFD14FE2CFD14BEC2" attach="left" box="[711,720,392,406]" fontSize="6" pageId="13" pageNumber="12">1</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="E386EADAFFA5C26DFCD5FEDBFCE6BEC2" box="[774,802,383,406]" italics="true" pageId="13" pageNumber="12">
|
||
et
|
||
<subScript id="4D76348DFFA5C26DFCCAFE2CFCE6BEC2" attach="left" box="[793,802,392,406]" fontSize="6" pageId="13" pageNumber="12">2</subScript>
|
||
</emphasis>
|
||
is approximately the same as between
|
||
<emphasis id="E386EADAFFA5C26DFDBCFE39FD4FBEE1" box="[623,651,413,437]" italics="true" pageId="13" pageNumber="12">
|
||
et
|
||
<subScript id="4D76348DFFA5C26DFD51FE03FD4FBEE1" attach="left" box="[642,651,423,437]" fontSize="6" pageId="13" pageNumber="12">2</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="E386EADAFFA5C26DFD15FE39FD1DBEE0" box="[710,729,413,436]" italics="true" pageId="13" pageNumber="12">et</emphasis>
|
||
<quantity id="160A9B2DFFA5C26DFD0AFE03FCC4BEE0" box="[729,768,412,437]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="13" pageNumber="12" unit="in" value="3.0">
|
||
<emphasis id="E386EADAFFA5C26DFD0AFE03FD26BEE1" box="[729,738,423,437]" italics="true" pageId="13" pageNumber="12">
|
||
<subScript id="4D76348DFFA5C26DFD0AFE03FD26BEE1" attach="left" box="[729,738,423,437]" fontSize="6" pageId="13" pageNumber="12">3</subScript>
|
||
</emphasis>
|
||
in
|
||
</quantity>
|
||
<taxonomicName id="16F24D4BFFA5C26DFCD8FE39FEC5BE87" authorityName="Ayrey & Soleglad" authorityYear="2015" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA5C26DFCD8FE39FEC5BE87" italics="true" pageId="13" pageNumber="12">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
. For the other two species, the distance between
|
||
<emphasis id="E386EADAFFA5C26DFF13FE7EFF18BEA6" box="[192,220,474,498]" italics="true" pageId="13" pageNumber="12">
|
||
et
|
||
<subScript id="4D76348DFFA5C26DFF00FE40FF18BEA6" attach="left" box="[211,220,484,498]" fontSize="6" pageId="13" pageNumber="12">2</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="E386EADAFFA5C26DFEC3FE7EFEE9BEA6" box="[272,301,474,498]" italics="true" pageId="13" pageNumber="12">
|
||
et
|
||
<subScript id="4D76348DFFA5C26DFEF7FE40FEE9BEA6" attach="left" box="[292,301,484,498]" fontSize="6" pageId="13" pageNumber="12">3</subScript>
|
||
</emphasis>
|
||
is shorter. Trichobothria
|
||
<emphasis id="E386EADAFFA5C26DFDEEFE7EFD47BEA6" box="[573,643,474,498]" italics="true" pageId="13" pageNumber="12">
|
||
et
|
||
<subScript id="4D76348DFFA5C26DFD83FE40FD9DBEA6" attach="left" box="[592,601,484,498]" fontSize="6" pageId="13" pageNumber="12">1</subScript>
|
||
–et
|
||
<subScript id="4D76348DFFA5C26DFDA9FE40FD47BEA6" attach="left" box="[634,643,484,498]" fontSize="6" pageId="13" pageNumber="12">3</subScript>
|
||
</emphasis>
|
||
are essentially in a straight line in
|
||
<taxonomicName id="16F24D4BFFA5C26DFE4AFE5DFE3FBD44" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[409,507,505,528]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA5C26DFE4AFE5DFE3FBD44" box="[409,507,505,528]" italics="true" pageId="13" pageNumber="12">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
, whereas
|
||
<emphasis id="E386EADAFFA5C26DFDB9FE5DFD42BD44" box="[618,646,505,528]" italics="true" pageId="13" pageNumber="12">
|
||
et
|
||
<subScript id="4D76348DFFA5C26DFDAEFDA6FD42BD44" attach="left" box="[637,646,514,528]" fontSize="6" pageId="13" pageNumber="12">3</subScript>
|
||
</emphasis>
|
||
is out of line, closer to the ventral edge in the other two species. Based on the analysis of published trichobothrial patterns of other sky island species which exhibit six
|
||
<emphasis id="E386EADAFFA5C26DFD7CFDF1FD0FBD38" box="[687,715,597,620]" italics="true" pageId="13" pageNumber="12">ID</emphasis>
|
||
on the movable finger, we see that species
|
||
<taxonomicName id="16F24D4BFFA5C26DFD9CFDD0FD16BDDF" authorityName="Ayrey" authorityYear="2009" box="[591,722,627,651]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="deboerae">
|
||
<emphasis id="E386EADAFFA5C26DFD9CFDD0FD16BDDF" box="[591,722,627,651]" italics="true" pageId="13" pageNumber="12">V. deboerae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA5C26DFCD8FDD0FF35BDFE" authorityName="Ayrey" authorityYear="2012" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="halli">
|
||
<emphasis id="E386EADAFFA5C26DFCD8FDD0FF35BDFE" italics="true" pageId="13" pageNumber="12">V. halli</emphasis>
|
||
</taxonomicName>
|
||
have patterns similar to that described for
|
||
<taxonomicName id="16F24D4BFFA5C26DFD12FD37FCE6BDFE" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[705,802,659,682]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA5C26DFD12FD37FCE6BDFE" box="[705,802,659,682]" italics="true" pageId="13" pageNumber="12">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
(see Ayrey, 2009: fig. 14; and Ayrey, 2012: fig. 10).
|
||
</paragraph>
|
||
<paragraph id="D14D36C8FFA5C26DFF23FD74FEE4BCAF" blockId="13.[192,802,198,1019]" pageId="13" pageNumber="12">
|
||
We compared several morphometric ratios between the
|
||
<taxonomicName id="16F24D4BFFA5C26DFF3BFD4BFE8CBC52" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[232,328,751,774]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA5C26DFF3BFD4BFE8CBC52" box="[232,328,751,774]" italics="true" pageId="13" pageNumber="12">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
female and
|
||
<taxonomicName id="16F24D4BFFA5C26DFE03FD4BFD8DBC52" box="[464,585,750,774]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA5C26DFE03FD4BFD8DBC52" box="[464,585,750,774]" italics="true" pageId="13" pageNumber="12">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
(three females) and
|
||
<taxonomicName id="16F24D4BFFA5C26DFF13FCA9FEFCBC70" authorityName=": Ayrey" authorityYear="2013" box="[192,312,780,804]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA5C26DFF13FCA9FEFCBC70" box="[192,312,780,804]" italics="true" pageId="13" pageNumber="12">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
(four females, including the
|
||
<typeStatus id="0E49886AFFA5C26DFD53FCA8FD26BC70" box="[640,738,780,804]" pageId="13" pageNumber="12" type="lectotype">lectotype</typeStatus>
|
||
measurements contained in Graham (2007: 3–4)). These measurements are presented in
|
||
<tableCitation id="9C700373FFA5C26DFDF2FCEEFDBCBC36" box="[545,632,842,866]" captionStart="Table 1" captionStartId="5.[192,257,1337,1361]" captionTargetId="graphics-8@5.[255,1858,192,1307]" captionTargetPageId="5" captionText="Table 1: Morphometrics (mm) of Vaejovis troupi sp. n., Whetstone Mountains, Cochise County, Arizona, USA, V. grahami, Madera County, Arizona, USA (after Ayrey & Soleglad, 2014: tab.1, in part), and V. vorhiesi lectotype, Miller’s Canyon, Huachuca Mountains, Graham, 2007: 3–4, in part)." pageId="13" pageNumber="12">Table 1</tableCitation>
|
||
. The
|
||
<taxonomicName id="16F24D4BFFA5C26DFD6CFCEFFCE6BC36" authorityName="Ayrey & Soleglad" authorityYear="2015" box="[703,802,843,866]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="troupi">
|
||
<emphasis id="E386EADAFFA5C26DFD6CFCEFFCE6BC36" box="[703,802,843,866]" italics="true" pageId="13" pageNumber="12">V. troupi</emphasis>
|
||
</taxonomicName>
|
||
female has a relatively thinner metasoma than female
|
||
<taxonomicName id="16F24D4BFFA5C26DFCD8FCCDFEDEBCCB" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="E386EADAFFA5C26DFCD8FCCDFEDEBCCB" italics="true" pageId="13" pageNumber="12">V. grahami</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="16F24D4BFFA5C26DFE9CFC2CFE07BCCB" authorityName=": Ayrey" authorityYear="2013" box="[335,451,903,927]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="vorhiesi">
|
||
<emphasis id="E386EADAFFA5C26DFE9CFC2CFE07BCCB" box="[335,451,903,927]" italics="true" pageId="13" pageNumber="12">V. vorhiesi</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<tableCitation id="9C700373FFA5C26DFE2EFC23FD94BCCB" box="[509,592,903,927]" captionStart="Table 2" captionStartId="12.[192,257,524,548]" captionTargetId="graphics-13@12.[187,1445,192,495]" captionTargetPageId="12" captionText="Table 2: Comparison of morphometric ratios of the metasoma based on the measurements of female specimens presented in Table 1. In these ratios, Vaejovis troupi sp. n. exhibits the more slender metasoma when compared to the other two species (as indicated by a larger ratio value), ranging from 8 to over 16 percent mean value difference (MVD). Statistical data group: minimum–maximum (mean) (±standard deviation) [number of samples]. Seg = metasomal segment, L = length, W = width." pageId="13" pageNumber="12">Table 2</tableCitation>
|
||
), ranging from 8 to over 16 percent mean value differences (
|
||
<collectionCode id="B7E3AE0DFFA5C26DFDACFC02FD04BCEA" box="[639,704,934,958]" pageId="13" pageNumber="12">MVD</collectionCode>
|
||
). In particular, the basal segments I–III exhibit the largest differences.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |