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<document id="9796611312ED3EF1D4F38F48FFC68414" ID-DOI="10.5852/ejt.2023.908.2345" ID-ISSN="2118-9773" ID-Zenodo-Dep="10202951" ID-ZooBank="738E2549-40F3-48E5-9123-F4F2B0D2B834" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700815481197" checkinUser="plazi" docAuthor="Jonishi, Taro &amp; Nakano, Takafumi" docDate="2023" docId="4E6387C65F729D59BE0AF91F1BE0F93C" docLanguage="en" docName="EJT.2023.908.155-182.pdf" docOrigin="European Journal of Taxonomy 908" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2345/10223" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.33:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="33" docTitle="Scolopocryptops brevisulcatus Jonishi &amp; Nakano 2023, sp. nov." docType="treatment" docUuid="3C9DB52B-2691-4980-B339-1B31E177F10C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="174" masterDocId="B25AFFBE5F7F9D4ABC39FFF21F60FFFA" masterDocTitle="Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae)" masterLastPageNumber="182" masterPageNumber="155" pageNumber="168" updateTime="1704507926729" updateUser="felipe" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7CFF31F8F119AFDAD92C7827D353EED9">Two new species of Scolopocryptops centipedes from southern Japan (Chilopoda: Scolopendromorpha: Scolopocryptopidae)</mods:title>
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<mods:namePart id="4F9774007F36C9BF09E1EB6D5D9D9E74">Jonishi, Taro</mods:namePart>
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<mods:affiliation id="76EF44E665BAECC30145DE65EA6819FA">Department of Zoology, Graduate School of Science, Kyoto University, Kyoto 606 - 8502, Japan.</mods:affiliation>
<mods:nameIdentifier id="387802E3D4EF7AA5F371E90AD594DEEF" type="email">ykn347635@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0F0261196279E90767BE9542527698E1">Nakano, Takafumi</mods:namePart>
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<mods:affiliation id="FD68066581C73BE4EECA46C5B3E74691">Department of Zoology, Graduate School of Science, Kyoto University, Kyoto 606 - 8502, Japan.</mods:affiliation>
<mods:nameIdentifier id="7C12E02DEBD7BAB8A3BACD80F390EB60" type="email">nakano@zoo.zool.kyoto-u.ac.jp</mods:nameIdentifier>
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<mods:title id="9B4DE04212AC4CEB26E2AF2DA096EB4C">European Journal of Taxonomy</mods:title>
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<mods:date id="D8933CE4FDAD1046FA7F2D1F78CACD1D">2023</mods:date>
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<treatment id="4E6387C65F729D59BE0AF91F1BE0F93C" LSID="urn:lsid:zoobank.org:act:3C9DB52B-2691-4980-B339-1B31E177F10C" httpUri="http://treatment.plazi.org/id/4E6387C65F729D59BE0AF91F1BE0F93C" lastPageId="19" lastPageNumber="174" pageId="13" pageNumber="168">
<subSubSection id="8ED0655B5F729D47BE0AF91F1B61F8F2" box="[563,1025,1773,1800]" pageId="13" pageNumber="168" type="nomenclature">
<paragraph id="C67536D05F729D47BE0AF91F1B61F8F2" blockId="13.[372,1215,1773,1869]" box="[563,1025,1773,1800]" pageId="13" pageNumber="168">
<heading id="9D3D81BC5F729D47BE0AF91F1B61F8F2" box="[563,1025,1773,1800]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="168" reason="2">
<taxonomicName id="01CA4D535F729D47BE0AF91F1CFFF8F2" authority="Jonishi &amp; Nakano, 2023" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[563,927,1773,1800]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="13" pageNumber="168" phylum="Arthropoda" rank="species" species="brevisulcatus" status="sp. nov.">
<emphasis id="F4BEEAC25F729D47BE0AF91F1CFFF8F2" bold="true" box="[563,927,1773,1800]" italics="true" pageId="13" pageNumber="168">Scolopocryptops brevisulcatus</emphasis>
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<taxonomicNameLabel id="EF8D57B95F729D47BF9FF91C1B61F8F2" box="[934,1025,1774,1800]" pageId="13" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="8ED0655B5F729D47BD4DF8E21C34F8B7" pageId="13" pageNumber="168" type="description">
<paragraph id="C67536D05F729D47BD4DF8E21BDFF8D1" blockId="13.[372,1215,1773,1869]" box="[372,1215,1808,1835]" pageId="13" pageNumber="168">
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urn:lsid:zoobank.org:act:
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</paragraph>
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<figureCitation id="5EF12A555F729D47BEE6F8C01C34F8B7" box="[735,852,1842,1869]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="14.[189,232,1963,1989]" captionStartId-1="15.[189,232,1928,1954]" captionStartId-2="16.[189,232,1636,1662]" captionStartId-3="17.[189,232,1773,1799]" captionStartId-4="18.[189,232,1822,1848]" captionStartId-5="19.[189,232,731,757]" captionTargetBox-0="[189,1383,1529,1913]" captionTargetBox-1="[210,1387,1436,1868]" captionTargetBox-2="[189,1398,265,1570]" captionTargetBox-3="[230,1369,322,1671]" captionTargetBox-4="[189,1398,1004,1759]" captionTargetBox-5="[189,1398,270,686]" captionTargetId-0="figure-370@14.[189,1398,1529,1913]" captionTargetId-1="figure-392@15.[189,1398,1421,1882]" captionTargetId-2="figure-17@16.[189,1398,265,1570]" captionTargetId-3="figure-11@17.[189,1398,280,1693]" captionTargetId-4="figure-235@18.[189,1398,1004,1759]" captionTargetId-5="figure-376@19.[189,1398,270,686]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionTargetPageId-3="17" captionTargetPageId-4="18" captionTargetPageId-5="19" captionText-0="Fig. 9. Scolopocryptops brevisulcatus sp. nov., holotype, ♂ (KUZ Z4390) and habitat at the type locality. A. Habitat (laurel tree forest) on Okinawa Island. B. Live specimen, dorsal view. Scale bar = 10 mm." captionText-1="Fig. 10. Scolopocryptops brevisulcatus sp. nov., holotype, ♂ (KUZ Z4390). A. Basal articles of left antenna, dorsal view. B. Cephalic plate, dorsal view; arrowhead indicates left cephalic marginal sulcus. Scale bars = 1 mm." captionText-2="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." captionText-3="Fig. 12. Scolopocryptops brevisulcatus sp. nov. A. Holotype, ♂ (KUZ Z4390). B. Paratype, subadult, sex undetermined (KUZ Z4386). Forcipules and forcipular coxosternite, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; md = median diastema; op = outer part of anterior margin of forcipular coxosternite; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A = 1 mm; B = 0.2 mm." captionText-4="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." captionText-5="Fig. 14. Scolopocryptops brevisulcatus sp. nov. AB. Holotype, ♂ (KUZ Z4390). C. Paratype, ♀ (KUZ Z4389). A. Right ultimate leg, lateral view. B. Male genital segments, lamina subanalis, and anal valves, ventral view. C. Female genital segment, lamina subanalis, and anal valves, ventral view. Abbreviations: av = anal valve; ls = lamina subanalis; sgs I = sternite of genital segment 1; sgs II = sternite of genital segment 2. Scale bars: A = 2 mm; BC = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10202969" figureDoi-1="http://doi.org/10.5281/zenodo.10202971" figureDoi-2="http://doi.org/10.5281/zenodo.10202975" figureDoi-3="http://doi.org/10.5281/zenodo.10202977" figureDoi-4="http://doi.org/10.5281/zenodo.10202979" figureDoi-5="http://doi.org/10.5281/zenodo.10202981" httpUri-0="https://zenodo.org/record/10202969/files/figure.png" httpUri-1="https://zenodo.org/record/10202971/files/figure.png" httpUri-2="https://zenodo.org/record/10202975/files/figure.png" httpUri-3="https://zenodo.org/record/10202977/files/figure.png" httpUri-4="https://zenodo.org/record/10202979/files/figure.png" httpUri-5="https://zenodo.org/record/10202981/files/figure.png" pageId="13" pageNumber="168">Figs 914</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8ED0655B5F729D44BC84F8811E00FE54" lastPageId="14" lastPageNumber="169" pageId="13" pageNumber="168" type="diagnosis">
<paragraph id="C67536D05F729D47BC84F8811E56F877" blockId="13.[189,310,1907,1933]" box="[189,310,1907,1933]" pageId="13" pageNumber="168">
<heading id="9D3D81BC5F729D47BC84F8811E56F877" bold="true" box="[189,310,1907,1933]" fontSize="11" level="3" pageId="13" pageNumber="168" reason="3">
<emphasis id="F4BEEAC25F729D47BC84F8811E56F877" bold="true" box="[189,310,1907,1933]" pageId="13" pageNumber="168">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="C67536D05F729D44BC84F8501E00FE54" blockId="13.[189,1401,1953,2015]" lastBlockId="14.[189,1400,265,430]" lastPageId="14" lastPageNumber="169" pageId="13" pageNumber="168">
Antenna with sparse minute hairs and short setae on dorsal surface of basal four articles, subsequent articles densely setose. Cephalic plate with short and shallow lateral marginal sulci in posterior half. Forcipular coxosternite with anterior margin slightly bilobed but median diastema almost lacking, darkly sclerotized bands not reaching outer part of anterior margin, with a pair of almost-reduced teeth. Coxopleuron approx. 1.61.7
<emphasis id="F4BEEAC25F719D44BE27FEBD1D52FE92" box="[542,562,335,360]" italics="true" pageId="14" pageNumber="169">×</emphasis>
as long as sternite 23; pleural dorsal margin slightly protruding from lateral margin of tergite 23, posterior and ventral margins forming approx. 7080° angle; coxopleural process short.
</paragraph>
</subSubSection>
<subSubSection id="8ED0655B5F719D44BC84FE271B6FFDBB" pageId="14" pageNumber="169" type="etymology">
<paragraph id="C67536D05F719D44BC84FE271E24FE15" blockId="14.[189,324,469,495]" box="[189,324,469,495]" pageId="14" pageNumber="169">
<heading id="9D3D81BC5F719D44BC84FE271E24FE15" bold="true" box="[189,324,469,495]" fontSize="11" level="3" pageId="14" pageNumber="169" reason="3">
<emphasis id="F4BEEAC25F719D44BC84FE271E24FE15" bold="true" box="[189,324,469,495]" pageId="14" pageNumber="169">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C67536D05F719D44BC84FDF61B6FFDBB" blockId="14.[189,1399,516,577]" pageId="14" pageNumber="169">
The specific name is derived from the Latin words
<emphasis id="F4BEEAC25F719D44BF09FDF61C17FDE4" box="[816,887,516,542]" italics="true" pageId="14" pageNumber="169">brevis</emphasis>
(short, shallow) and
<emphasis id="F4BEEAC25F719D44B8A5FDF61B85FDE4" box="[1180,1253,516,542]" italics="true" pageId="14" pageNumber="169">sulcus</emphasis>
(groove), referring to the unapparent cephalic marginal sulcus of this new species.
</paragraph>
</subSubSection>
<subSubSection id="8ED0655B5F719D44BC84FD9B1B38FD78" box="[189,1112,616,643]" pageId="14" pageNumber="169" type="vernacular_names">
<paragraph id="C67536D05F719D44BC84FD9B1B38FD78" blockId="14.[189,1112,616,643]" box="[189,1112,616,643]" pageId="14" pageNumber="169">We herein suggest a Japanese name for this species as Kirekomi-akamukade.</paragraph>
</subSubSection>
<subSubSection id="8ED0655B5F719D44BC84FD5B1D07FA98" pageId="14" pageNumber="169" type="materials_examined">
<paragraph id="C67536D05F719D44BC84FD5B1ECAFD39" blockId="14.[189,426,681,707]" box="[189,426,681,707]" pageId="14" pageNumber="169">
<heading id="9D3D81BC5F719D44BC84FD5B1ECAFD39" bold="true" box="[189,426,681,707]" fontSize="11" level="3" pageId="14" pageNumber="169" reason="3">
<emphasis id="F4BEEAC25F719D44BC84FD5B1ECAFD39" bold="true" box="[189,426,681,707]" pageId="14" pageNumber="169">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="C67536D05F719D44BCD3FD2A1C51FCC3" blockId="14.[187,1398,728,825]" pageId="14" pageNumber="169">
<materialsCitation id="76A23C8D5F719D44BCD3FD2A1C51FCC3" collectingDate="2022-06-14" collectionCode="KUZ" collectorName="T. Jonishi" country="Japan" county="Island" elevation="311" latitude="26.63157" location="Mt. Katsuu-dake" longLatPrecision="1" longitude="127.93771" municipality="Motobu" pageId="14" pageNumber="169" specimenCode="Z4390" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa Island" typeStatus="holotype">
<emphasis id="F4BEEAC25F719D44BCD3FD2A1E3CFD08" bold="true" box="[234,348,728,754]" pageId="14" pageNumber="169">
<typeStatus id="197188725F719D44BCD3FD2A1E3CFD08" box="[234,348,728,754]" pageId="14" pageNumber="169" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="BEDD76405F719D44BC84FD0E1E76FCEC" box="[189,278,764,790]" name="Japan" pageId="14" pageNumber="169">JAPAN</collectingCountry>
<emphasis id="F4BEEAC25F719D44BD22FD0E1E94FCEC" bold="true" box="[283,500,763,790]" pageId="14" pageNumber="169">
<collectingRegion id="040EF8325F719D44BD16FD091E94FCEC" box="[303,500,763,790]" pageId="14" pageNumber="169">Okinawa Island</collectingRegion>
</emphasis>
<specimenCount id="D0CCFD595F719D44BE30FD0E1D42FCEC" box="[521,546,764,790]" count="1" pageId="14" pageNumber="169" type="male"></specimenCount>
;
<collectingMunicipality id="2611ACAA5F719D44BE14FD091DEDFCEF" box="[557,653,763,789]" pageId="14" pageNumber="169">Motobu</collectingMunicipality>
, near
<location id="C315600B5F719D44BEF7FD0E1CF1FCEF" LSID="urn:lsid:plazi:treatment:4E6387C65F729D59BE0AF91F1BE0F93C:C315600B5F719D44BEF7FD0E1CF1FCEF" box="[718,913,763,790]" country="Japan" county="Island" latitude="26.63157" longLatPrecision="1" longitude="127.93771" municipality="Motobu" name="Mt. Katsuu-dake" pageId="14" pageNumber="169" stateProvince="Okinawa Island">Mt. Katsuu-dake</location>
(
<figureCitation id="5EF12A555F719D44BFA7FD091C87FCEC" box="[926,999,763,790]" captionStart="Fig" captionStartId="2.[189,232,1787,1813]" captionTargetBox="[189,1398,889,1744]" captionTargetId="figure-181@2.[189,1398,889,1744]" captionTargetPageId="2" captionText="Fig. 1. Map showing the collection localities of Scolopocryptops elegans (Takakuwa, 1937), S. miyosii sp. nov., S. brevisulcatus sp. nov., and S. curtus (Takakuwa, 1939) in the present study and Jonishi &amp; Nakano (2022).Orange circles:S. elegans; pink diamonds:S. miyosii sp. nov.; red squares:S. brevisulcatus sp. nov.; yellow triangles: S. curtus; black triangles: localities of the sequence data of S. curtus obtained from INSD (see Table 1). Shapes with two lines indicate the type localities of each species (S. curtus from its type locality was not examined). Locality numbers (E1E5, M1M5, B1B4, C1 and C2) are shown in Fig. 2 and Table 1." figureDoi="http://doi.org/10.5281/zenodo.10202953" httpUri="https://zenodo.org/record/10202953/files/figure.png" pageId="14" pageNumber="169">Figs 1</figureCitation>
,
<figureCitation id="5EF12A555F719D44BFCAFD0E1B7BFCEC" box="[1011,1051,764,790]" captionStart="Fig" captionStartId="14.[189,232,1963,1989]" captionTargetBox="[189,1383,1529,1913]" captionTargetId="figure-370@14.[189,1398,1529,1913]" captionTargetPageId="14" captionText="Fig. 9. Scolopocryptops brevisulcatus sp. nov., holotype, ♂ (KUZ Z4390) and habitat at the type locality. A. Habitat (laurel tree forest) on Okinawa Island. B. Live specimen, dorsal view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10202969" httpUri="https://zenodo.org/record/10202969/files/figure.png" pageId="14" pageNumber="169">9A</figureCitation>
);
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26.63157
<emphasis id="F4BEEAC25F719D44B8A5FD0E1BC8FCEC" box="[1180,1192,764,790]" italics="true" pageId="14" pageNumber="169">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="A3FE50175F719D44B8F6FD0E1A11FCEC" box="[1231,1393,764,790]" degrees="127.93771" direction="east" orientation="longitude" pageId="14" pageNumber="169" precision="1" value="127.93771">
127.93771
<emphasis id="F4BEEAC25F719D44B972FD0E1A37FCEC" box="[1355,1367,764,790]" italics="true" pageId="14" pageNumber="169">°</emphasis>
E
</geoCoordinate>
;
<quantity id="01329B355F719D44BC82FCED1E64FCC3" box="[187,260,799,825]" metricMagnitude="2" metricUnit="m" metricValue="3.11" pageId="14" pageNumber="169" unit="m" value="311.0">
<elevation id="4DE7D1E35F719D44BC82FCED1E64FCC3" box="[187,260,799,825]" metricMagnitude="2" metricUnit="m" metricValue="3.11" pageId="14" pageNumber="169" unit="m" value="311.0">311 m</elevation>
</quantity>
alt.;
<date id="B27410105F719D44BD06FCED1EBAFCC3" box="[319,474,799,825]" pageId="14" pageNumber="169" value="2022-06-14">
<collectingDate id="A230E9F85F719D44BD06FCED1EBAFCC3" box="[319,474,799,825]" pageId="14" pageNumber="169" value="2022-06-14">14 Jun. 2022</collectingDate>
</date>
;
<collectorName id="6B3F53065F719D44BDDCFCED1D37FCC3" box="[485,599,799,825]" pageId="14" pageNumber="169">T. Jonishi</collectorName>
leg.;
<collectionCode id="A0DBAE155F719D44BEA0FCED1DB5FCC3" box="[665,725,799,825]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">KUZ</collectionCode>
<specimenCode id="966C9EAB5F719D44BEE5FCED1C4DFCC3" box="[732,813,799,825]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">Z4390</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="C67536D05F719D44BCD3FC921A58FBB7" blockId="14.[189,1399,864,1102]" pageId="14" pageNumber="169">
<materialsCitation id="76A23C8D5F719D44BCD3FC921C34FC3A" collectingDate="2021-05-04" collectionCode="KUZ" collectorName="F. Okuyama" country="Japan" county="Island" elevation="298" latitude="26.80745" location="Mt. Nishime-dake" longLatPrecision="1" longitude="128.26617" municipality="Kunigami" pageId="14" pageNumber="169" specimenCode="Z4386" specimenCount="1" specimenCount-subadult="1" stateProvince="Okinawa Island" typeStatus="paratype">
<emphasis id="F4BEEAC25F719D44BCD3FC921E07FC80" bold="true" box="[234,359,864,890]" pageId="14" pageNumber="169">
<typeStatus id="197188725F719D44BCD3FC921E07FC80" box="[234,359,864,890]" pageId="14" pageNumber="169" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="BEDD76405F719D44BC84FC711E76FC67" box="[189,278,899,925]" name="Japan" pageId="14" pageNumber="169">JAPAN</collectingCountry>
<emphasis id="F4BEEAC25F719D44BD22FC711E95FC67" bold="true" box="[283,501,899,925]" pageId="14" pageNumber="169">
<collectingRegion id="040EF8325F719D44BD09FC711E95FC67" box="[304,501,899,925]" pageId="14" pageNumber="169">Okinawa Island</collectingRegion>
</emphasis>
<specimenCount id="D0CCFD595F719D44BE32FC711DE7FC67" box="[523,647,899,925]" count="1" pageId="14" pageNumber="169" type="subadult">1 subadult</specimenCount>
;
<collectingMunicipality id="2611ACAA5F719D44BEA8FC711C69FC67" box="[657,777,899,925]" pageId="14" pageNumber="169">Kunigami</collectingMunicipality>
, near
<location id="C315600B5F719D44BF72FC761B42FC67" LSID="urn:lsid:plazi:treatment:4E6387C65F729D59BE0AF91F1BE0F93C:C315600B5F719D44BF72FC761B42FC67" box="[843,1058,899,926]" country="Japan" county="Island" latitude="26.80745" longLatPrecision="1" longitude="128.26617" municipality="Kunigami" name="Mt. Nishime-dake" pageId="14" pageNumber="169" stateProvince="Okinawa Island">Mt. Nishime-dake</location>
;
<geoCoordinate id="A3FE50175F719D44B814FC711BA4FC64" box="[1069,1220,899,926]" degrees="26.80745" direction="north" orientation="latitude" pageId="14" pageNumber="169" precision="1" value="26.80745">
26.80745
<emphasis id="F4BEEAC25F719D44B8A2FC711BC7FC67" box="[1179,1191,899,925]" italics="true" pageId="14" pageNumber="169">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="A3FE50175F719D44B8F7FC711A11FC64" box="[1230,1393,899,926]" degrees="128.26618" direction="east" orientation="longitude" pageId="14" pageNumber="169" precision="1" value="128.26617">
128.26618
<emphasis id="F4BEEAC25F719D44B972FC711A37FC67" box="[1355,1367,899,925]" italics="true" pageId="14" pageNumber="169">°</emphasis>
E
</geoCoordinate>
;
<quantity id="01329B355F719D44BC84FC551E69FC3B" box="[189,265,935,961]" metricMagnitude="2" metricUnit="m" metricValue="2.98" pageId="14" pageNumber="169" unit="m" value="298.0">
<elevation id="4DE7D1E35F719D44BC84FC551E69FC3B" box="[189,265,935,961]" metricMagnitude="2" metricUnit="m" metricValue="2.98" pageId="14" pageNumber="169" unit="m" value="298.0">298 m</elevation>
</quantity>
alt.;
<date id="B27410105F719D44BD71FC551EBDFC3B" box="[328,477,935,961]" pageId="14" pageNumber="169" value="2021-05-04">
<collectingDate id="A230E9F85F719D44BD71FC551EBDFC3B" box="[328,477,935,961]" pageId="14" pageNumber="169" value="2021-05-04">4 May 2021</collectingDate>
</date>
;
<collectorName id="6B3F53065F719D44BDD2FC551D1CFC3A" box="[491,636,934,961]" pageId="14" pageNumber="169">F. Okuyama</collectorName>
leg.;
<collectionCode id="A0DBAE155F719D44BEFBFC551D9EFC3B" box="[706,766,935,961]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">KUZ</collectionCode>
<specimenCode id="966C9EAB5F719D44BF31FC541C34FC3A" box="[776,852,934,960]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">Z4386</specimenCode>
</materialsCitation>
<materialsCitation id="76A23C8D5F719D44BF48FC551A17FC1E" collectingDate="2022-04-08" collectionCode="KUZ" collectorName="F. Okuyama" country="Japan" county="Island" elevation="107" latitude="26.81195" location="Mt. Nishime-dake" longLatPrecision="1" longitude="128.28415" municipality="Kunigami" pageId="14" pageNumber="169" specimenCode="Z4387" specimenCount="1" specimenCount-adult="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="D0CCFD595F719D44BF48FC551CA7FC3A" box="[881,967,934,961]" count="1" pageId="14" pageNumber="169" type="adult">1 adult</specimenCount>
, sex undetermined; same locality as for preceding;
<geoCoordinate id="A3FE50175F719D44BD4BFC3B1D6EFC1E" box="[370,526,969,996]" degrees="26.81195" direction="north" orientation="latitude" pageId="14" pageNumber="169" precision="1" value="26.81195">
26.81195
<emphasis id="F4BEEAC25F719D44BDE7FC381E8AFC1E" box="[478,490,970,996]" italics="true" pageId="14" pageNumber="169">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="A3FE50175F719D44BE24FC3B1DA6FC1E" box="[541,710,969,996]" degrees="128.28415" direction="east" orientation="longitude" pageId="14" pageNumber="169" precision="1" value="128.28415">
128.28415
<emphasis id="F4BEEAC25F719D44BEA3FC381DC6FC1E" box="[666,678,970,996]" italics="true" pageId="14" pageNumber="169">°</emphasis>
E
</geoCoordinate>
;
<quantity id="01329B355F719D44BEEFFC381C45FC1E" box="[726,805,970,996]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" pageId="14" pageNumber="169" unit="m" value="107.0">
<elevation id="4DE7D1E35F719D44BEEFFC381C45FC1E" box="[726,805,970,996]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" pageId="14" pageNumber="169" unit="m" value="107.0">107 m</elevation>
</quantity>
alt.;
<date id="B27410105F719D44BF51FC381C9DFC1E" box="[872,1021,970,996]" pageId="14" pageNumber="169" value="2022-04-08">
<collectingDate id="A230E9F85F719D44BF51FC381C9DFC1E" box="[872,1021,970,996]" pageId="14" pageNumber="169" value="2022-04-08">8 Apr. 2022</collectingDate>
</date>
;
<collectorName id="6B3F53065F719D44B834FC381BC0FC1E" box="[1037,1184,970,996]" pageId="14" pageNumber="169">F. Okuyama</collectorName>
leg;
<collectionCode id="A0DBAE155F719D44B8DBFC381A7EFC1E" box="[1250,1310,970,996]" collectionName="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">KUZ</collectionCode>
<specimenCode id="966C9EAB5F719D44B913FC381A17FC1E" box="[1322,1399,970,996]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">Z4387</specimenCode>
</materialsCitation>
<materialsCitation id="76A23C8D5F719D44BCEBFC1F1C11FBFD" box="[210,881,1005,1031]" collectingDate="2022-06-14" collectionCode="KUZ" collectorName="Jonishi" country="Japan" county="Island" elevation="311" latitude="26.63157" location="Mt. Katsuu-dake" longLatPrecision="1" longitude="127.93771" municipality="Motobu" pageId="14" pageNumber="169" specimenCode="Z4389" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="D0CCFD595F719D44BCEBFC1F1E62FBFD" box="[210,258,1005,1031]" count="1" pageId="14" pageNumber="169" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="A0DBAE155F719D44BEE4FC1F1C79FBFD" box="[733,793,1005,1031]" collectionName="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">KUZ</collectionCode>
<specimenCode id="966C9EAB5F719D44BF1DFC1F1C11FBFD" box="[804,881,1005,1031]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">Z4389</specimenCode>
</materialsCitation>
<materialsCitation id="76A23C8D5F719D44BFA9FC1F1B01FBD0" collectingDate="2022-06-15" collectionCode="KUZ" collectorName="T. Jonishi" country="Japan" county="Island" elevation="355" latitude="26.8008" location="Mt. Nishime-dake" longLatPrecision="1" longitude="128.27728" municipality="Kunigami" pageId="14" pageNumber="169" specimenCode="Z4391" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="D0CCFD595F719D44BFA9FC1F1CA2FBFD" box="[912,962,1005,1031]" count="1" pageId="14" pageNumber="169" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="2611ACAA5F719D44BFE8FC1F1B29FBFD" box="[977,1097,1005,1031]" pageId="14" pageNumber="169">Kunigami</collectingMunicipality>
, near
<location id="C315600B5F719D44B8AFFC1F1A12FBFD" LSID="urn:lsid:plazi:treatment:4E6387C65F729D59BE0AF91F1BE0F93C:C315600B5F719D44B8AFFC1F1A12FBFD" box="[1174,1394,1005,1031]" country="Japan" county="Island" latitude="26.8008" longLatPrecision="1" longitude="128.27728" municipality="Kunigami" name="Mt. Nishime-dake" pageId="14" pageNumber="169" stateProvince="Okinawa">Mt. Nishime-dake</location>
;
<geoCoordinate id="A3FE50175F719D44BC84FBE21E34FBD1" box="[189,340,1040,1067]" degrees="26.80080" direction="north" orientation="latitude" pageId="14" pageNumber="169" precision="1" value="26.8008">
26.80080
<emphasis id="F4BEEAC25F719D44BD12FBE21E57FBD0" box="[299,311,1040,1066]" italics="true" pageId="14" pageNumber="169">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="A3FE50175F719D44BD64FBE21D60FBD1" box="[349,512,1040,1067]" degrees="128.27728" direction="east" orientation="longitude" pageId="14" pageNumber="169" precision="1" value="128.27728">128.27728° E</geoCoordinate>
;
<quantity id="01329B355F719D44BE33FBE21D32FBD1" box="[522,594,1040,1067]" metricMagnitude="2" metricUnit="m" metricValue="3.55" pageId="14" pageNumber="169" unit="m" value="355.0">
<elevation id="4DE7D1E35F719D44BE33FBE21D32FBD1" box="[522,594,1040,1067]" metricMagnitude="2" metricUnit="m" metricValue="3.55" pageId="14" pageNumber="169" unit="m" value="355.0">355 m</elevation>
</quantity>
alt.;
<date id="B27410105F719D44BEB1FBE21C7EFBD0" box="[648,798,1040,1067]" pageId="14" pageNumber="169" value="2022-06-15">
<collectingDate id="A230E9F85F719D44BEB1FBE21C7EFBD0" box="[648,798,1040,1067]" pageId="14" pageNumber="169" value="2022-06-15">15 Jun. 2022</collectingDate>
</date>
;
<collectorName id="6B3F53065F719D44BF1EFBE31CF6FBD0" box="[807,918,1040,1067]" pageId="14" pageNumber="169">T. Jonishi</collectorName>
leg.;
<collectionCode id="A0DBAE155F719D44BFEAFBE31B6FFBD1" box="[979,1039,1041,1067]" collectionName="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">KUZ</collectionCode>
<specimenCode id="966C9EAB5F719D44B82CFBE21B01FBD0" box="[1045,1121,1040,1066]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">Z4391</specimenCode>
</materialsCitation>
<materialsCitation id="76A23C8D5F719D44B84FFBE21A54FBB7" collectingDate="2022-06-16" collectionCode="KUZ" collectorName="T. Jonishi" country="Japan" county="Island" elevation="384" latitude="26.75251" location="Mt. Fuenchiji" longLatPrecision="1" longitude="128.24303" municipality="Kunigami" pageId="14" pageNumber="169" specimenCode="Z4392" specimenCount="1" specimenCount-subadult="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="D0CCFD595F719D44B84FFBE21B90FBD0" box="[1142,1264,1040,1066]" count="1" pageId="14" pageNumber="169" type="subadult">1 subadult</specimenCount>
;
<collectingMunicipality id="2611ACAA5F719D44B8C3FBE21A12FBD0" box="[1274,1394,1040,1066]" pageId="14" pageNumber="169">Kunigami</collectingMunicipality>
,
<location id="C315600B5F719D44BC84FBC61E01FBB7" LSID="urn:lsid:plazi:treatment:4E6387C65F729D59BE0AF91F1BE0F93C:C315600B5F719D44BC84FBC61E01FBB7" box="[189,353,1075,1102]" country="Japan" county="Island" latitude="26.75251" longLatPrecision="1" longitude="128.24303" municipality="Kunigami" name="Mt. Fuenchiji" pageId="14" pageNumber="169" stateProvince="Okinawa">Mt. Fuenchiji</location>
;
<geoCoordinate id="A3FE50175F719D44BD55FBC11D65FBB4" box="[364,517,1075,1102]" degrees="26.75251" direction="north" orientation="latitude" pageId="14" pageNumber="169" precision="1" value="26.75251">
26.75251
<emphasis id="F4BEEAC25F719D44BDE3FBC11E86FBB7" box="[474,486,1075,1101]" italics="true" pageId="14" pageNumber="169">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="A3FE50175F719D44BE28FBC11DD5FBB4" box="[529,693,1075,1102]" degrees="128.24303" direction="east" orientation="longitude" pageId="14" pageNumber="169" precision="1" value="128.24303">
128.24303
<emphasis id="F4BEEAC25F719D44BEB7FBC11DFAFBB7" box="[654,666,1075,1101]" italics="true" pageId="14" pageNumber="169">°</emphasis>
E
</geoCoordinate>
,
<quantity id="01329B355F719D44BEF8FBC11C6CFBB4" box="[705,780,1075,1102]" metricMagnitude="2" metricUnit="m" metricValue="3.84" pageId="14" pageNumber="169" unit="m" value="384.0">
<elevation id="4DE7D1E35F719D44BEF8FBC11C6CFBB4" box="[705,780,1075,1102]" metricMagnitude="2" metricUnit="m" metricValue="3.84" pageId="14" pageNumber="169" unit="m" value="384.0">384 m</elevation>
</quantity>
alt.;
<date id="B27410105F719D44BF7EFBC11C81FBB7" box="[839,993,1075,1102]" pageId="14" pageNumber="169" value="2022-06-16">
<collectingDate id="A230E9F85F719D44BF7EFBC11C81FBB7" box="[839,993,1075,1102]" pageId="14" pageNumber="169" value="2022-06-16">16 Jun. 2022</collectingDate>
</date>
;
<collectorName id="6B3F53065F719D44BFD4FBC61B3EFBB7" box="[1005,1118,1075,1102]" pageId="14" pageNumber="169">T. Jonishi</collectorName>
leg.;
<collectionCode id="A0DBAE155F719D44B899FBC61BBCFBB4" box="[1184,1244,1076,1102]" collectionName="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">KUZ</collectionCode>
<specimenCode id="966C9EAB5F719D44B8DDFBC11A54FBB7" box="[1252,1332,1075,1101]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">Z4392</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="C67536D05F719D44BCD3FB861E81FB74" blockId="14.[189,1399,1140,1378]" box="[234,481,1140,1166]" pageId="14" pageNumber="169">
<emphasis id="F4BEEAC25F719D44BCD3FB861E81FB74" bold="true" box="[234,481,1140,1166]" pageId="14" pageNumber="169">Additional material</emphasis>
</paragraph>
<paragraph id="C67536D05F719D44BC84FB6A1D07FA98" blockId="14.[189,1399,1140,1378]" pageId="14" pageNumber="169">
<materialsCitation id="76A23C8D5F719D44BC84FB6A1C8AFB2C" collectingDate="2019-09-20" collectionCode="KUZ" collectorName="T. Jonishi" country="Japan" county="Island" elevation="291" latitude="26.63145" location="Okinawa Island" longLatPrecision="1" longitude="127.93812" municipality="Motobu" pageId="14" pageNumber="169" specimenCode="Z4385" specimenCount="1" specimenCount-adult="1" stateProvince="Okinawa Island">
<collectingCountry id="BEDD76405F719D44BC84FB6A1E76FB48" box="[189,278,1176,1202]" name="Japan" pageId="14" pageNumber="169">JAPAN</collectingCountry>
<emphasis id="F4BEEAC25F719D44BD26FB6A1D62FB48" bold="true" box="[287,514,1176,1202]" pageId="14" pageNumber="169">
<collectingRegion id="040EF8325F719D44BD01FB6A1D62FB48" box="[312,514,1176,1202]" pageId="14" pageNumber="169">Okinawa Island</collectingRegion>
</emphasis>
<specimenCount id="D0CCFD595F719D44BE19FB6A1D16FB48" box="[544,630,1176,1202]" count="1" pageId="14" pageNumber="169" type="adult">1 adult</specimenCount>
, sex undetermined; same locality as for holotype;
<geoCoordinate id="A3FE50175F719D44B8EFFB6A1A12FB48" box="[1238,1394,1176,1203]" degrees="26.63145" direction="north" orientation="latitude" pageId="14" pageNumber="169" precision="1" value="26.63145">
26.63145
<emphasis id="F4BEEAC25F719D44B97DFB6A1A30FB48" box="[1348,1360,1176,1202]" italics="true" pageId="14" pageNumber="169">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="A3FE50175F719D44BC84FB491E03FB2C" box="[189,355,1211,1238]" degrees="127.93812" direction="east" orientation="longitude" pageId="14" pageNumber="169" precision="1" value="127.93812">
127.93812
<emphasis id="F4BEEAC25F719D44BD03FB491E26FB2F" box="[314,326,1211,1237]" italics="true" pageId="14" pageNumber="169">°</emphasis>
E
</geoCoordinate>
;
<quantity id="01329B355F719D44BD49FB491EDBFB2C" box="[368,443,1211,1238]" metricMagnitude="2" metricUnit="m" metricValue="2.91" pageId="14" pageNumber="169" unit="m" value="291.0">
<elevation id="4DE7D1E35F719D44BD49FB491EDBFB2C" box="[368,443,1211,1238]" metricMagnitude="2" metricUnit="m" metricValue="2.91" pageId="14" pageNumber="169" unit="m" value="291.0">291 m</elevation>
</quantity>
alt.;
<date id="B27410105F719D44BDC1FB491DF7FB2F" box="[504,663,1211,1237]" pageId="14" pageNumber="169" value="2019-09-20">
<collectingDate id="A230E9F85F719D44BDC1FB491DF7FB2F" box="[504,663,1211,1237]" pageId="14" pageNumber="169" value="2019-09-20">20 Sep. 2019</collectingDate>
</date>
;
<collectorName id="6B3F53065F719D44BE9AFB4E1C75FB2F" box="[675,789,1211,1238]" pageId="14" pageNumber="169">T. Jonishi</collectorName>
leg.;
<collectionCode id="A0DBAE155F719D44BF60FB4E1CF5FB2C" box="[857,917,1212,1238]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">KUZ</collectionCode>
<specimenCode id="966C9EAB5F719D44BFA4FB491C8AFB2C" box="[925,1002,1211,1238]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">Z4385</specimenCode>
</materialsCitation>
<materialsCitation id="76A23C8D5F719D44B83CFB491A70FB02" collectingDate="2022-06-14" collectionCode="KUZ" collectorName="T. Jonishi" country="Japan" county="Island" elevation="307" latitude="26.63139" location="Mt. Katsuu-dake" longLatPrecision="1" longitude="127.93792" municipality="Motobu" pageId="14" pageNumber="169" specimenCode="Z4388" specimenCount="1" specimenCount-subadult="1" stateProvince="Okinawa">
<specimenCount id="D0CCFD595F719D44B83CFB491BE2FB2F" box="[1029,1154,1211,1237]" count="1" pageId="14" pageNumber="169" type="subadult">1 subadult</specimenCount>
; same locality as for holotype;
<geoCoordinate id="A3FE50175F719D44BD0CFB2C1EAFFB03" box="[309,463,1246,1273]" degrees="26.63139" direction="north" orientation="latitude" pageId="14" pageNumber="169" precision="1" value="26.63139">
26.63139
<emphasis id="F4BEEAC25F719D44BD9AFB2C1ECFFB02" box="[419,431,1246,1272]" italics="true" pageId="14" pageNumber="169">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="A3FE50175F719D44BDE4FB2C1DE3FB03" box="[477,643,1246,1273]" degrees="127.93792" direction="east" orientation="longitude" pageId="14" pageNumber="169" precision="1" value="127.93792">127.93792° E</geoCoordinate>
;
<quantity id="01329B355F719D44BEA8FB2C1DBDFB03" box="[657,733,1246,1273]" metricMagnitude="2" metricUnit="m" metricValue="3.07" pageId="14" pageNumber="169" unit="m" value="307.0">
<elevation id="4DE7D1E35F719D44BEA8FB2C1DBDFB03" box="[657,733,1246,1273]" metricMagnitude="2" metricUnit="m" metricValue="3.07" pageId="14" pageNumber="169" unit="m" value="307.0">307 m</elevation>
</quantity>
alt.;
<date id="B27410105F719D44BF22FB2C1CD9FB02" box="[795,953,1246,1273]" pageId="14" pageNumber="169" value="2022-06-14">
<collectingDate id="A230E9F85F719D44BF22FB2C1CD9FB02" box="[795,953,1246,1273]" pageId="14" pageNumber="169" value="2022-06-14">14 Jun. 2022</collectingDate>
</date>
;
<collectorName id="6B3F53065F719D44BFFFFB2D1B59FB02" box="[966,1081,1246,1273]" pageId="14" pageNumber="169">T. Jonishi</collectorName>
leg.;
<collectionCode id="A0DBAE155F719D44B847FB2D1BDAFB03" box="[1150,1210,1247,1273]" collectionName="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">KUZ</collectionCode>
<specimenCode id="966C9EAB5F719D44B8FAFB2C1A70FB02" box="[1219,1296,1246,1272]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">Z4388</specimenCode>
</materialsCitation>
<materialsCitation id="76A23C8D5F719D44B915FB2C1E33FAC5" collectingDate="2022-06-16" collectionCode="KUZ" collectorName="T. Jonishi" country="Japan" county="Island" elevation="244" latitude="26.68491" location="Mt. Nekumachiji" longLatPrecision="1" longitude="128.1343" municipality="Ogimi" pageId="14" pageNumber="169" specimenCode="Z4393" specimenCount="1" stateProvince="Okinawa">
<specimenCount id="D0CCFD595F719D44B915FB2C1A12FB02" box="[1324,1394,1246,1272]" count="1" pageId="14" pageNumber="169" type="generic">1 juv.</specimenCount>
;
<collectingMunicipality id="2611ACAA5F719D44BC84FAF31E6BFAE1" box="[189,267,1281,1307]" pageId="14" pageNumber="169">Ogimi</collectingMunicipality>
, near
<location id="C315600B5F719D44BD6DFAF01D43FAE1" LSID="urn:lsid:plazi:treatment:4E6387C65F729D59BE0AF91F1BE0F93C:C315600B5F719D44BD6DFAF01D43FAE1" box="[340,547,1281,1308]" country="Japan" county="Island" latitude="26.68491" longLatPrecision="1" longitude="128.1343" municipality="Ogimi" name="Mt. Nekumachiji" pageId="14" pageNumber="169" stateProvince="Okinawa">Mt. Nekumachiji</location>
;
<geoCoordinate id="A3FE50175F719D44BE09FAF31DACFAE6" box="[560,716,1281,1308]" degrees="26.68491" direction="north" orientation="latitude" pageId="14" pageNumber="169" precision="1" value="26.68491">
26.68491
<emphasis id="F4BEEAC25F719D44BEA7FAF01DCAFAE6" box="[670,682,1282,1308]" italics="true" pageId="14" pageNumber="169">°</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="A3FE50175F719D44BEE0FAF01CE0FAE6" box="[729,896,1282,1308]" degrees="128.13430" direction="east" orientation="longitude" pageId="14" pageNumber="169" precision="1" value="128.1343">
128.13430
<emphasis id="F4BEEAC25F719D44BF6FFAF01C02FAE6" box="[854,866,1282,1308]" italics="true" pageId="14" pageNumber="169">°</emphasis>
E
</geoCoordinate>
;
<quantity id="01329B355F719D44BFB6FAF01CBBFAE6" box="[911,987,1282,1308]" metricMagnitude="2" metricUnit="m" metricValue="2.44" pageId="14" pageNumber="169" unit="m" value="244.0">
<elevation id="4DE7D1E35F719D44BFB6FAF01CBBFAE6" box="[911,987,1282,1308]" metricMagnitude="2" metricUnit="m" metricValue="2.44" pageId="14" pageNumber="169" unit="m" value="244.0">244 m</elevation>
</quantity>
alt.;
<date id="B27410105F719D44B823FAF31BD8FAE6" box="[1050,1208,1281,1308]" pageId="14" pageNumber="169" value="2022-06-16">
<collectingDate id="A230E9F85F719D44B823FAF31BD8FAE6" box="[1050,1208,1281,1308]" pageId="14" pageNumber="169" value="2022-06-16">16 Jun. 2022</collectingDate>
</date>
;
<collectorName id="6B3F53065F719D44B8FFFAF01A5AFAE1" box="[1222,1338,1281,1308]" pageId="14" pageNumber="169">T. Jonishi</collectorName>
leg.;
<collectionCode id="A0DBAE155F719D44BC86FAD71F9BFAC5" box="[191,251,1317,1343]" collectionName="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">KUZ</collectionCode>
<specimenCode id="966C9EAB5F719D44BD3BFAD71E33FAC5" box="[258,339,1317,1343]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">Z4393</specimenCode>
</materialsCitation>
.
<materialsCitation id="76A23C8D5F719D44BD48FAD71D03FA98" collectingDate="2022-11-08" collectionCode="KUZ" collectorName="Ryobu Fukuyama" country="Japan" county="Izena Island" elevation="48" latitude="26.94288" location="Mt. Ufu-yama" longLatPrecision="1" longitude="127.92713" pageId="14" pageNumber="169" specimenCode="Z4394" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa">
<emphasis id="F4BEEAC25F719D44BD48FAD71D6AFAC5" bold="true" box="[369,522,1317,1343]" pageId="14" pageNumber="169">
<collectingCounty id="2F144E5C5F719D44BD48FAD71D6AFAC5" box="[369,522,1317,1343]" pageId="14" pageNumber="169">Izena Island</collectingCounty>
</emphasis>
<specimenCount id="D0CCFD595F719D44BE1AFAD71D30FAC5" box="[547,592,1317,1343]" count="1" pageId="14" pageNumber="169" type="female">1 ♀</specimenCount>
;
<location id="C315600B5F719D44BE67FAD71C69FAC5" LSID="urn:lsid:plazi:treatment:4E6387C65F729D59BE0AF91F1BE0F93C:C315600B5F719D44BE67FAD71C69FAC5" box="[606,777,1317,1343]" country="Japan" county="Izena Island" latitude="26.94288" longLatPrecision="1" longitude="127.92713" name="Mt. Ufu-yama" pageId="14" pageNumber="169" stateProvince="Okinawa">Mt. Ufu-yama</location>
;
<geoCoordinate id="A3FE50175F719D44BF2DFAD71CCDFAC5" box="[788,941,1317,1343]" degrees="26.94288" direction="north" orientation="latitude" pageId="14" pageNumber="169" precision="1" value="26.94288">26.94288° N</geoCoordinate>
,
<geoCoordinate id="A3FE50175F719D44BF80FAD71B3DFAC5" box="[953,1117,1317,1343]" degrees="127.92713" direction="east" orientation="longitude" pageId="14" pageNumber="169" precision="1" value="127.92713">127.92713° E</geoCoordinate>
;
<quantity id="01329B355F719D44B850FAD71BC4FAC5" box="[1129,1188,1317,1343]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="169" unit="m" value="48.0">
<elevation id="4DE7D1E35F719D44B850FAD71BC4FAC5" box="[1129,1188,1317,1343]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="169" unit="m" value="48.0">48 m</elevation>
</quantity>
alt.;
<date id="B27410105F719D44B8E6FAD71A12FAC5" box="[1247,1394,1317,1343]" pageId="14" pageNumber="169" value="2022-11-08">
<collectingDate id="A230E9F85F719D44B8E6FAD71A12FAC5" box="[1247,1394,1317,1343]" pageId="14" pageNumber="169" value="2022-11-08">8 Nov. 2022</collectingDate>
</date>
;
<collectorName id="6B3F53065F719D44BC84FABA1EEDFA98" box="[189,397,1352,1378]" pageId="14" pageNumber="169">Ryobu Fukuyama</collectorName>
leg.;
<collectionCode id="A0DBAE155F719D44BDF6FABA1D6BFA98" box="[463,523,1352,1378]" collectionName="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">KUZ</collectionCode>
<specimenCode id="966C9EAB5F719D44BE2AFABA1D03FA98" box="[531,611,1352,1378]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="14" pageNumber="169">Z4394</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="92B566585F719D44BC84F8591A3DF813" ID-DOI="http://doi.org/10.5281/zenodo.10202969" ID-Zenodo-Dep="10202969" httpUri="https://zenodo.org/record/10202969/files/figure.png" pageId="14" pageNumber="169" startId="14.[189,232,1963,1989]" targetBox="[189,1383,1529,1913]" targetPageId="14">
<paragraph id="C67536D05F719D44BC84F8591A3DF813" blockId="14.[189,1399,1963,2025]" pageId="14" pageNumber="169">
<emphasis id="F4BEEAC25F719D44BC84F8591E6AF83F" bold="true" box="[189,266,1963,1989]" pageId="14" pageNumber="169">Fig. 9.</emphasis>
<taxonomicName id="01CA4D535F719D44BD29F8591D14F83F" authority="Jonishi &amp; Nakano, 2023" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[272,628,1963,1989]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="14" pageNumber="169" phylum="Arthropoda" rank="species" species="brevisulcatus" status="sp. nov.">
<emphasis id="F4BEEAC25F719D44BD29F8591D14F83F" box="[272,628,1963,1989]" italics="true" pageId="14" pageNumber="169">Scolopocryptops brevisulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF8D57B95F719D44BE43F85E1DB5F83C" box="[634,725,1964,1990]" pageId="14" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (KUZ Z4390) and habitat at the type locality.
<emphasis id="F4BEEAC25F719D44BC84F83C1FB2F812" bold="true" box="[189,210,1998,2024]" pageId="14" pageNumber="169">A</emphasis>
. Habitat (laurel tree forest) on Okinawa Island.
<emphasis id="F4BEEAC25F719D44BF3FF83C1C7AF812" bold="true" box="[774,794,1998,2024]" pageId="14" pageNumber="169">B</emphasis>
. Live specimen, dorsal view. Scale bar = 10 mm.
</paragraph>
</caption>
<subSubSection id="8ED0655B5F709D59BC84FEFB1A2CFBBA" lastPageId="19" lastPageNumber="174" pageId="15" pageNumber="170" type="description">
<paragraph id="C67536D05F709D45BC84FEFB1B0AFED9" blockId="15.[189,1130,265,291]" box="[189,1130,265,291]" pageId="15" pageNumber="170">
<emphasis id="F4BEEAC25F709D45BC84FEFB1E81FED9" bold="true" box="[189,481,265,291]" pageId="15" pageNumber="170">
Description of
<typeStatus id="197188725F709D45BD4CFEFB1E81FED9" box="[373,481,265,291]" pageId="15" pageNumber="170" type="holotype">holotype</typeStatus>
</emphasis>
[variation in other specimens given in square brackets]
</paragraph>
<paragraph id="C67536D05F709D45BC84FEC51BD6FE8F" blockId="15.[189,1398,311,373]" pageId="15" pageNumber="170">
Body length 58.0 mm [
<quantity id="01329B355F709D45BDE9FEC51D1FFEA8" box="[464,639,311,338]" metricMagnitude="-2" metricUnit="m" metricValue="5.135" metricValueMax="5.91" metricValueMin="4.36" pageId="15" pageNumber="170" unit="mm" value="51.35" valueMax="59.1" valueMin="43.6">43.659.1 mm</quantity>
in adults] in 80% ethanol, 64.0 mm before fixation. Body color brownish red [brownish orange]; antennae, legs, and ultimate legs yellowish (
<figureCitation id="5EF12A555F709D45B875FEA91BC9FE8F" box="[1100,1193,347,373]" captionStart="Fig" captionStartId="14.[189,232,1963,1989]" captionTargetBox="[189,1383,1529,1913]" captionTargetId="figure-370@14.[189,1398,1529,1913]" captionTargetPageId="14" captionText="Fig. 9. Scolopocryptops brevisulcatus sp. nov., holotype, ♂ (KUZ Z4390) and habitat at the type locality. A. Habitat (laurel tree forest) on Okinawa Island. B. Live specimen, dorsal view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10202969" httpUri="https://zenodo.org/record/10202969/files/figure.png" pageId="15" pageNumber="170">Fig. 9B</figureCitation>
).
</paragraph>
<paragraph id="C67536D05F709D45BC84FE6E1C97FD2A" blockId="15.[187,1399,411,720]" pageId="15" pageNumber="170">
Antennae damaged, 13
<superScript id="31BF9B985F709D45BDF5FE691EB9FE53" attach="left" box="[460,473,411,425]" fontSize="6" pageId="15" pageNumber="170">th</superScript>
and subsequent articles lost [
<quantity id="01329B355F709D45BF0FFE6E1CF3FE4D" box="[822,915,412,439]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="15" pageNumber="170" unit="mm" value="9.9">9.9 mm</quantity>
in length, approx. 0.2
<emphasis id="F4BEEAC25F709D45B8A4FE6F1BD1FE4C" box="[1181,1201,413,438]" italics="true" pageId="15" pageNumber="170">×</emphasis>
as long as body, composed of 17 articles in
<typeStatus id="197188725F709D45BDCAFE321D38FE20" box="[499,600,448,474]" pageId="15" pageNumber="170" type="paratype">paratype</typeStatus>
<collectionCode id="A0DBAE155F709D45BE67FE321DFAFE20" box="[606,666,448,474]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="15" pageNumber="170">KUZ</collectionCode>
Z4391]; density of setae increasing toward distal articles; basal 4 articles with sparse minute hairs (see
<bibRefCitation id="A25B4B215F709D45BEB4FE111C0AFE07" author="Bonato L. &amp; Edgecombe G. D. &amp; Lewis J. G. E. &amp; Minelli A. &amp; Pereira L. A. &amp; Shelley R. M. &amp; Zapparoli M." box="[653,874,482,509]" pageId="15" pageNumber="170" pagination="17 - 51" refId="ref11376" refString="Bonato L., Edgecombe G. D., Lewis J. G. E., Minelli A., Pereira L. A., Shelley R. M. &amp; Zapparoli M. 2010. A common terminology for the external anatomy of centipedes (Chilopoda). ZooKeys 69: 17 - 51. https: // doi. org / 10.3897 / zookeys. 69.737" type="journal article" year="2010">
Bonato
<emphasis id="F4BEEAC25F709D45BED5FE111C47FE06" box="[748,807,482,508]" italics="true" pageId="15" pageNumber="170">et al.</emphasis>
2010
</bibRefCitation>
) and short setae dorsally, 5
<superScript id="31BF9B985F709D45B894FE101BDAFE0A" attach="left" box="[1197,1210,482,496]" fontSize="6" pageId="15" pageNumber="170">th</superScript>
and subsequent articles setose, 6
<superScript id="31BF9B985F709D45BD47FDF71EEBFDE9" attach="right" box="[382,395,517,531]" fontSize="6" pageId="15" pageNumber="170">th</superScript>
17
<superScript id="31BF9B985F709D45BD8EFDF71EA4FDE9" attach="left" box="[439,452,517,531]" fontSize="6" pageId="15" pageNumber="170">th</superScript>
articles more densely covered with setae [basal 2 articles sparsely setose, 3
<superScript id="31BF9B985F709D45B90EFDF71A25FDE9" attach="left" box="[1335,1349,517,531]" fontSize="6" pageId="15" pageNumber="170">rd</superScript>
and subsequent articles densely covered with setae in juvenile
<collectionCode id="A0DBAE155F709D45BF45FDDB1CD8FDB9" box="[892,952,553,579]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="15" pageNumber="170">KUZ</collectionCode>
Z4393; 4
<superScript id="31BF9B985F709D45B809FDDA1B5DFDCC" attach="left" box="[1072,1085,552,566]" fontSize="6" pageId="15" pageNumber="170">th</superScript>
article moderately setose, 5
<superScript id="31BF9B985F709D45BCF5FDB91FB9FDA3" attach="right" box="[204,217,587,601]" fontSize="6" pageId="15" pageNumber="170">th</superScript>
17
<superScript id="31BF9B985F709D45BD3CFDB91E72FDA3" attach="left" box="[261,274,587,601]" fontSize="6" pageId="15" pageNumber="170">th</superScript>
articles more densely covered with setae in subadults] (
<figureCitation id="5EF12A555F709D45BF87FDBE1B4FFD9C" box="[958,1071,588,614]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[210,1387,1436,1868]" captionTargetId="figure-392@15.[189,1398,1421,1882]" captionTargetPageId="15" captionText="Fig. 10. Scolopocryptops brevisulcatus sp. nov., holotype, ♂ (KUZ Z4390). A. Basal articles of left antenna, dorsal view. B. Cephalic plate, dorsal view; arrowhead indicates left cephalic marginal sulcus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202971" httpUri="https://zenodo.org/record/10202971/files/figure.png" pageId="15" pageNumber="170">Fig. 10A</figureCitation>
). Cephalic plate as long as wide, sides converging anteriorly (
<figureCitation id="5EF12A555F709D45BE66FD9D1DB1FD73" box="[607,721,623,649]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[210,1387,1436,1868]" captionTargetId="figure-392@15.[189,1398,1421,1882]" captionTargetPageId="15" captionText="Fig. 10. Scolopocryptops brevisulcatus sp. nov., holotype, ♂ (KUZ Z4390). A. Basal articles of left antenna, dorsal view. B. Cephalic plate, dorsal view; arrowhead indicates left cephalic marginal sulcus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202971" httpUri="https://zenodo.org/record/10202971/files/figure.png" pageId="15" pageNumber="170">Figs 10B</figureCitation>
,
<figureCitation id="5EF12A555F709D45BED9FD9D1C76FD73" box="[736,790,623,649]" captionStart="Fig" captionStartId="16.[189,232,1636,1662]" captionTargetBox="[189,1398,265,1570]" captionTargetId="figure-17@16.[189,1398,265,1570]" captionTargetPageId="16" captionText="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="15" pageNumber="170">11A</figureCitation>
); its surface finely punctate [minute setae on each punctum present in juvenile and subadults]; short and shallow lateral marginal sulci present in posterior half [marginal sulci absent in juvenile and subadults] (
<figureCitation id="5EF12A555F709D45BF00FD441CC7FD2A" box="[825,935,694,720]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[210,1387,1436,1868]" captionTargetId="figure-392@15.[189,1398,1421,1882]" captionTargetPageId="15" captionText="Fig. 10. Scolopocryptops brevisulcatus sp. nov., holotype, ♂ (KUZ Z4390). A. Basal articles of left antenna, dorsal view. B. Cephalic plate, dorsal view; arrowhead indicates left cephalic marginal sulcus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202971" httpUri="https://zenodo.org/record/10202971/files/figure.png" pageId="15" pageNumber="170">Figs 10B</figureCitation>
,
<figureCitation id="5EF12A555F709D45BF8DFD441C89FD2A" box="[948,1001,694,720]" captionStart="Fig" captionStartId="16.[189,232,1636,1662]" captionTargetBox="[189,1398,265,1570]" captionTargetId="figure-17@16.[189,1398,265,1570]" captionTargetPageId="16" captionText="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="15" pageNumber="170">11A</figureCitation>
).
</paragraph>
<paragraph id="C67536D05F709D45BC84FD051A23FBF2" blockId="15.[188,1400,759,1032]" pageId="15" pageNumber="170">
Second maxillae article 2 with elongated and semi-transparent dorsal spur distally; dorsal brush without transparent margin (
<figureCitation id="5EF12A555F709D45BD81FCE81D2DFCCE" box="[440,589,794,820]" captionStart="Fig" captionStartId="16.[189,232,1636,1662]" captionTargetBox="[189,1398,265,1570]" captionTargetId="figure-17@16.[189,1398,265,1570]" captionTargetPageId="16" captionText="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="15" pageNumber="170">Fig. 11BC</figureCitation>
); pretarsus consisting of dark brown basal and semi-transparent short apical parts (
<figureCitation id="5EF12A555F709D45BDA7FCCF1D4EFCA2" box="[414,558,829,856]" captionStart="Fig" captionStartId="16.[189,232,1636,1662]" captionTargetBox="[189,1398,265,1570]" captionTargetId="figure-17@16.[189,1398,265,1570]" captionTargetPageId="16" captionText="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="15" pageNumber="170">Fig. 11BC</figureCitation>
). Forcipular coxosternite and trochanteroprefemora sparsely punctate, coxosternite without sutures (
<figureCitation id="5EF12A555F709D45BE1FFC931DF6FC81" box="[550,662,865,891]" captionStart="Fig" captionStartId="16.[189,232,1636,1662]" captionTargetBox="[189,1398,265,1570]" captionTargetId="figure-17@16.[189,1398,265,1570]" captionTargetPageId="16" captionText="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="15" pageNumber="170">Fig. 11D</figureCitation>
); forcipular trochanteroprefemur with small and blunt black process, with apparent basal suture (
<figureCitation id="5EF12A555F709D45BE47FC761D92FC64" box="[638,754,900,926]" captionStart="Fig" captionStartId="16.[189,232,1636,1662]" captionTargetBox="[189,1398,265,1570]" captionTargetId="figure-17@16.[189,1398,265,1570]" captionTargetPageId="16" captionText="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="15" pageNumber="170">Figs 11D</figureCitation>
,
<figureCitation id="5EF12A555F709D45BF3AFC761C43FC64" box="[771,803,900,926]" captionStart="Fig" captionStartId="17.[189,232,1773,1799]" captionTargetBox="[230,1369,322,1671]" captionTargetId="figure-11@17.[189,1398,280,1693]" captionTargetPageId="17" captionText="Fig. 12. Scolopocryptops brevisulcatus sp. nov. A. Holotype, ♂ (KUZ Z4390). B. Paratype, subadult, sex undetermined (KUZ Z4386). Forcipules and forcipular coxosternite, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; md = median diastema; op = outer part of anterior margin of forcipular coxosternite; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A = 1 mm; B = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202977" httpUri="https://zenodo.org/record/10202977/files/figure.png" pageId="15" pageNumber="170">12</figureCitation>
); anterior margin of coxosternite almost straight and slightly bilobed but median diastema almost lacking; darkly sclerotized bands not reaching outer part of anterior margin of coxosternite; a pair of teeth almost reduced, forming antero-lateral corner of sclerotized bands [teeth on anterior margin prominent in juvenile and subadults] (
<figureCitation id="5EF12A555F709D45B840FC1F1A68FBF2" box="[1145,1288,1005,1032]" captionStart="Fig" captionStartId="16.[189,232,1636,1662]" captionTargetBox="[189,1398,265,1570]" captionTargetId="figure-17@16.[189,1398,265,1570]" captionTargetPageId="16" captionText="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="15" pageNumber="170">Figs 11DE</figureCitation>
,
<figureCitation id="5EF12A555F709D45B92CFC1C1A56FBF2" box="[1301,1334,1006,1032]" captionStart="Fig" captionStartId="17.[189,232,1773,1799]" captionTargetBox="[230,1369,322,1671]" captionTargetId="figure-11@17.[189,1398,280,1693]" captionTargetPageId="17" captionText="Fig. 12. Scolopocryptops brevisulcatus sp. nov. A. Holotype, ♂ (KUZ Z4390). B. Paratype, subadult, sex undetermined (KUZ Z4386). Forcipules and forcipular coxosternite, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; md = median diastema; op = outer part of anterior margin of forcipular coxosternite; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A = 1 mm; B = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202977" httpUri="https://zenodo.org/record/10202977/files/figure.png" pageId="15" pageNumber="170">12</figureCitation>
).
</paragraph>
<paragraph id="C67536D05F709D45BC84FBDD1E81FB6A" blockId="15.[189,1399,1071,1168]" pageId="15" pageNumber="170">
Tergites finely punctate; tergite 1 with anterior transverse suture, anterior margin covered by cephalic plate (
<figureCitation id="5EF12A555F709D45BD3EFBA01E18FB96" box="[263,376,1106,1132]" captionStart="Fig" captionStartId="16.[189,232,1636,1662]" captionTargetBox="[189,1398,265,1570]" captionTargetId="figure-17@16.[189,1398,265,1570]" captionTargetPageId="16" captionText="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="15" pageNumber="170">Figs 11A</figureCitation>
,
<figureCitation id="5EF12A555F709D45BDBFFBA01EDDFB96" box="[390,445,1106,1132]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="15" pageNumber="170">13A</figureCitation>
). Paramedian sutures present on tergites 222, lateral marginations complete on tergites 621 (
<figureCitation id="5EF12A555F709D45BD5EFB871EB4FB6A" box="[359,468,1141,1168]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="15" pageNumber="170">Fig. 13A</figureCitation>
).
</paragraph>
<paragraph id="C67536D05F709D45BC84FB451BE3FAC1" blockId="15.[189,1399,1207,1339]" pageId="15" pageNumber="170">
Sternites lacking paramedian sutures, finely punctate (
<figureCitation id="5EF12A555F709D45BF12FB451CF4FB2B" box="[811,916,1207,1233]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="15" pageNumber="170">Fig. 13B</figureCitation>
); sternites 220 with shallow longitudinal sulcus and slight median depression (
<figureCitation id="5EF12A555F709D45BEB9FB281D8DFB0E" box="[640,749,1242,1268]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="15" pageNumber="170">Fig. 13B</figureCitation>
), both shallower and unapparent on anterior sternites. Sternite 23 damaged, right posterior margin lost [sides of sternite 23 converging posteriorly, posterior margin slightly concave in
<typeStatus id="197188725F709D45BDC4FAD31D02FAC1" box="[509,610,1313,1339]" pageId="15" pageNumber="170" type="paratype">paratype</typeStatus>
<collectionCode id="A0DBAE155F709D45BE53FAD31DC6FAC1" box="[618,678,1313,1339]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="15" pageNumber="170">KUZ</collectionCode>
Z4387 and other specimens] (
<figureCitation id="5EF12A555F709D45B833FAD21B16FAC1" box="[1034,1142,1312,1339]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="15" pageNumber="170">Fig. 13C</figureCitation>
).
</paragraph>
<caption id="92B566585F709D45BC84F87A1EFFF813" ID-DOI="http://doi.org/10.5281/zenodo.10202971" ID-Zenodo-Dep="10202971" httpUri="https://zenodo.org/record/10202971/files/figure.png" pageId="15" pageNumber="170" startId="15.[189,232,1928,1954]" targetBox="[210,1387,1436,1868]" targetPageId="15">
<paragraph id="C67536D05F709D45BC84F87A1EFFF813" blockId="15.[189,1399,1928,2025]" pageId="15" pageNumber="170">
<emphasis id="F4BEEAC25F709D45BC84F87A1E7DF858" bold="true" box="[189,285,1928,1954]" pageId="15" pageNumber="170">Fig. 10.</emphasis>
<taxonomicName id="01CA4D535F709D45BD11F87A1DF0F858" authority="Jonishi &amp; Nakano, 2023" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[296,656,1928,1954]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="15" pageNumber="170" phylum="Arthropoda" rank="species" species="brevisulcatus" status="sp. nov.">
<emphasis id="F4BEEAC25F709D45BD11F87A1DF0F858" box="[296,656,1928,1954]" italics="true" pageId="15" pageNumber="170">Scolopocryptops brevisulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF8D57B95F709D45BEA2F87A1D9BF858" box="[667,763,1928,1954]" pageId="15" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (KUZ Z4390).
<emphasis id="F4BEEAC25F709D45B862F87A1B10F858" bold="true" box="[1115,1136,1928,1954]" pageId="15" pageNumber="170">A</emphasis>
. Basal articles of left antenna, dorsal view.
<emphasis id="F4BEEAC25F709D45BD85F8591EB0F83F" bold="true" box="[444,464,1963,1989]" pageId="15" pageNumber="170">B</emphasis>
. Cephalic plate, dorsal view; arrowhead indicates left cephalic marginal sulcus. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="92B566585F6F9D5ABC84F9961D1EF88F" ID-DOI="http://doi.org/10.5281/zenodo.10202975" ID-Zenodo-Dep="10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="16" pageNumber="171" startId="16.[189,232,1636,1662]" targetBox="[189,1398,265,1570]" targetPageId="16">
<paragraph id="C67536D05F6F9D5ABC84F9961D1EF88F" blockId="16.[187,1399,1636,1909]" pageId="16" pageNumber="171">
<emphasis id="F4BEEAC25F6F9D5ABC84F9961E78F984" bold="true" box="[189,280,1636,1662]" pageId="16" pageNumber="171">Fig. 11.</emphasis>
<taxonomicName id="01CA4D535F6F9D5ABD27F9961DE3F984" authority="Jonishi &amp; Nakano, 2023" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[286,643,1636,1662]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="16" pageNumber="171" phylum="Arthropoda" rank="species" species="brevisulcatus" status="sp. nov.">
<emphasis id="F4BEEAC25F6F9D5ABD27F9961DE3F984" box="[286,643,1636,1662]" italics="true" pageId="16" pageNumber="171">Scolopocryptops brevisulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF8D57B95F6F9D5ABEB0F9971D83F985" box="[649,739,1637,1663]" pageId="16" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="F4BEEAC25F6F9D5ABEDEF9961C41F984" bold="true" box="[743,801,1636,1662]" pageId="16" pageNumber="171">AD</emphasis>
. Holotype, ♂ (KUZ Z4390).
<emphasis id="F4BEEAC25F6F9D5AB84FF9961BEAF984" bold="true" box="[1142,1162,1636,1662]" pageId="16" pageNumber="171">E</emphasis>
. Paratype, subadult, sex undetermined (KUZ Z4386).
<emphasis id="F4BEEAC25F6F9D5ABE7CF9751D3AF95B" bold="true" box="[581,602,1671,1697]" pageId="16" pageNumber="171">A</emphasis>
. Cephalic plate and tergite 1, dorsal view.
<emphasis id="F4BEEAC25F6F9D5AB875F9751B00F95B" bold="true" box="[1100,1120,1671,1697]" pageId="16" pageNumber="171">B</emphasis>
. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view.
<emphasis id="F4BEEAC25F6F9D5ABFA8F9581CC6F93E" bold="true" box="[913,934,1706,1732]" pageId="16" pageNumber="171">C</emphasis>
. Article 3 and pretarsus of left second maxilla, lateral view.
<emphasis id="F4BEEAC25F6F9D5ABDF3F93C1D61F912" bold="true" box="[458,513,1742,1768]" pageId="16" pageNumber="171">DE</emphasis>
. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm.
</paragraph>
</caption>
<caption id="92B566585F6E9D5BBC84F91F1D75F842" ID-DOI="http://doi.org/10.5281/zenodo.10202977" ID-Zenodo-Dep="10202977" httpUri="https://zenodo.org/record/10202977/files/figure.png" pageId="17" pageNumber="172" startId="17.[189,232,1773,1799]" targetBox="[230,1369,322,1671]" targetPageId="17">
<paragraph id="C67536D05F6E9D5BBC84F91F1D75F842" blockId="17.[189,1401,1773,1976]" pageId="17" pageNumber="172">
<emphasis id="F4BEEAC25F6E9D5BBC84F91F1E7CF8FD" bold="true" box="[189,284,1773,1799]" pageId="17" pageNumber="172">Fig. 12.</emphasis>
<taxonomicName id="01CA4D535F6E9D5BBD1CF91F1DECF8FD" authority="Jonishi &amp; Nakano, 2023" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[293,652,1773,1799]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="17" pageNumber="172" phylum="Arthropoda" rank="species" species="brevisulcatus" status="sp. nov.">
<emphasis id="F4BEEAC25F6E9D5BBD1CF91F1DECF8FD" box="[293,652,1773,1799]" italics="true" pageId="17" pageNumber="172">Scolopocryptops brevisulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF8D57B95F6E9D5BBEACF91C1D91F8F2" box="[661,753,1774,1800]" pageId="17" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="F4BEEAC25F6E9D5BBEC2F91F1C6FF8FD" bold="true" box="[763,783,1773,1799]" pageId="17" pageNumber="172">A</emphasis>
. Holotype, ♂ (KUZ Z4390).
<emphasis id="F4BEEAC25F6E9D5BB848F91F1BE5F8FD" bold="true" box="[1137,1157,1773,1799]" pageId="17" pageNumber="172">B</emphasis>
. Paratype, subadult, sex undetermined (KUZ Z4386). Forcipules and forcipular coxosternite, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; md = median diastema; op = outer part of anterior margin of forcipular coxosternite; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A = 1 mm; B = 0.2 mm.
</paragraph>
</caption>
<paragraph id="C67536D05F6D9D58BC84FEFB1BDAFED9" blockId="18.[189,1210,265,292]" box="[189,1210,265,292]" pageId="18" pageNumber="173">Spiracles ovoid, present on leg-bearing segments 3, 5, 8, 10, 12, 14, 16, 18, 20, and 22.</paragraph>
<paragraph id="C67536D05F6D9D58BC84FEB91EDCFE51" blockId="18.[188,1399,330,427]" pageId="18" pageNumber="173">Legs lacking setae; tarsi of legs 121 undivided; legs 119 with lateral and ventral tibial spurs and tarsal spur, legs 20 and 21, respectively, with tibial spur and tarsal spur; leg 22 without spurs. All legs with two accessory spines.</paragraph>
<paragraph id="C67536D05F6D9D58BC84FE201EC8FD3A" blockId="18.[189,1400,466,704]" pageId="18" pageNumber="173">
Coxopleuron approx. 1.7
<emphasis id="F4BEEAC25F6D9D58BDD4FE211D61FE16" box="[493,513,467,492]" italics="true" pageId="18" pageNumber="173">×</emphasis>
[1.61.7
<emphasis id="F4BEEAC25F6D9D58BE4BFE211DE6FE16" box="[626,646,467,492]" italics="true" pageId="18" pageNumber="173">×</emphasis>
] as long as sternite 23 (
<figureCitation id="5EF12A555F6D9D58BF8AFE201B43FE16" box="[947,1059,466,492]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="18" pageNumber="173">Fig. 13D</figureCitation>
). Dorsal margin of ultimate pleuron slightly protruding from lateral side of tergite 23 (
<figureCitation id="5EF12A555F6D9D58BF4CFE071B65FDEA" box="[885,1029,501,528]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="18" pageNumber="173">Fig. 13DE</figureCitation>
), posterior margin with minute dark spine (
<figureCitation id="5EF12A555F6D9D58BD74FDEB1EDCFDC9" box="[333,444,537,563]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="18" pageNumber="173">Fig. 13D</figureCitation>
). Posterior and ventral margins of coxopleuron converging posteriorly, forming approx. 70° [7080°] angle (
<figureCitation id="5EF12A555F6D9D58BE36FDCE1D1CFDAC" box="[527,636,572,598]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="18" pageNumber="173">Fig. 13D</figureCitation>
); coxopleural process short, tip of process broken [tip of process pointed, slightly directed dorsally in
<collectionCode id="A0DBAE155F6D9D58BE43FD921DD6FD80" box="[634,694,608,634]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="18" pageNumber="173">KUZ</collectionCode>
Z4387] (
<figureCitation id="5EF12A555F6D9D58BF10FDAD1CF8FD83" box="[809,920,607,633]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="18" pageNumber="173">Fig. 13D</figureCitation>
). Surface of coxopleuron without setae, covered with coxal pores of various size; coxopleural process and dorso-posterior area of coxopleuron poreless (
<figureCitation id="5EF12A555F6D9D58BD17FD571EFBFD3A" box="[302,411,677,704]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="18" pageNumber="173">Fig. 13D</figureCitation>
).
</paragraph>
<paragraph id="C67536D05F6D9D58BC84FD151D07FCBD" blockId="18.[189,1399,743,840]" pageId="18" pageNumber="173">
Ultimate leg
<quantity id="01329B355F6D9D58BD61FD151EA1FCFB" box="[344,449,743,770]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="18" pageNumber="173" unit="mm" value="15.3">15.3 mm</quantity>
in length, 0.26
<emphasis id="F4BEEAC25F6D9D58BE4EFD1A1DEBFCFB" box="[631,651,744,769]" italics="true" pageId="18" pageNumber="173">×</emphasis>
as long as body; all articles lacking setae (
<figureCitation id="5EF12A555F6D9D58B8BEFD151B95FCFB" box="[1159,1269,743,769]" captionStart="Fig" captionStartId="19.[189,232,731,757]" captionTargetBox="[189,1398,270,686]" captionTargetId="figure-376@19.[189,1398,270,686]" captionTargetPageId="19" captionText="Fig. 14. Scolopocryptops brevisulcatus sp. nov. AB. Holotype, ♂ (KUZ Z4390). C. Paratype, ♀ (KUZ Z4389). A. Right ultimate leg, lateral view. B. Male genital segments, lamina subanalis, and anal valves, ventral view. C. Female genital segment, lamina subanalis, and anal valves, ventral view. Abbreviations: av = anal valve; ls = lamina subanalis; sgs I = sternite of genital segment 1; sgs II = sternite of genital segment 2. Scale bars: A = 2 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10202981" httpUri="https://zenodo.org/record/10202981/files/figure.png" pageId="18" pageNumber="173">Fig. 14A</figureCitation>
); prefemur with two conical and pointed spinose processes, ventral process large, dorso-medial one minute; pretarsus with two accessory spines.
</paragraph>
<paragraph id="C67536D05F6D9D59BC84FC9D1A2CFBBA" blockId="18.[189,1399,877,940]" lastBlockId="19.[189,1399,957,1089]" lastPageId="19" lastPageNumber="174" pageId="18" pageNumber="173">
Genital segments occupying approx. ½ length of sternite 23. Sternite of genital segment 1 with sparse minute setae; posterior margin weakly convex (
<figureCitation id="5EF12A555F6D9D58BED0FC601C34FC56" box="[745,852,914,940]" captionStart="Fig" captionStartId="19.[189,232,731,757]" captionTargetBox="[189,1398,270,686]" captionTargetId="figure-376@19.[189,1398,270,686]" captionTargetPageId="19" captionText="Fig. 14. Scolopocryptops brevisulcatus sp. nov. AB. Holotype, ♂ (KUZ Z4390). C. Paratype, ♀ (KUZ Z4389). A. Right ultimate leg, lateral view. B. Male genital segments, lamina subanalis, and anal valves, ventral view. C. Female genital segment, lamina subanalis, and anal valves, ventral view. Abbreviations: av = anal valve; ls = lamina subanalis; sgs I = sternite of genital segment 1; sgs II = sternite of genital segment 2. Scale bars: A = 2 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10202981" httpUri="https://zenodo.org/record/10202981/files/figure.png" pageId="18" pageNumber="173">Fig. 14B</figureCitation>
). Sternite of genital segment 2 well developed, covered with sparse minute setae; posterior part of genital segment 2 overlapped by anal valves, penis not visible in ventral view (
<figureCitation id="5EF12A555F6C9D59BE34FC121D1BFC00" box="[525,635,992,1018]" captionStart="Fig" captionStartId="19.[189,232,731,757]" captionTargetBox="[189,1398,270,686]" captionTargetId="figure-376@19.[189,1398,270,686]" captionTargetPageId="19" captionText="Fig. 14. Scolopocryptops brevisulcatus sp. nov. AB. Holotype, ♂ (KUZ Z4390). C. Paratype, ♀ (KUZ Z4389). A. Right ultimate leg, lateral view. B. Male genital segments, lamina subanalis, and anal valves, ventral view. C. Female genital segment, lamina subanalis, and anal valves, ventral view. Abbreviations: av = anal valve; ls = lamina subanalis; sgs I = sternite of genital segment 1; sgs II = sternite of genital segment 2. Scale bars: A = 2 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10202981" httpUri="https://zenodo.org/record/10202981/files/figure.png" pageId="19" pageNumber="174">Fig. 14B</figureCitation>
); lamina subanalis situated between genital segment 2 and anal valves (
<figureCitation id="5EF12A555F6C9D59BD21FBF11EE1FBE7" box="[280,385,1027,1053]" captionStart="Fig" captionStartId="19.[189,232,731,757]" captionTargetBox="[189,1398,270,686]" captionTargetId="figure-376@19.[189,1398,270,686]" captionTargetPageId="19" captionText="Fig. 14. Scolopocryptops brevisulcatus sp. nov. AB. Holotype, ♂ (KUZ Z4390). C. Paratype, ♀ (KUZ Z4389). A. Right ultimate leg, lateral view. B. Male genital segments, lamina subanalis, and anal valves, ventral view. C. Female genital segment, lamina subanalis, and anal valves, ventral view. Abbreviations: av = anal valve; ls = lamina subanalis; sgs I = sternite of genital segment 1; sgs II = sternite of genital segment 2. Scale bars: A = 2 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10202981" httpUri="https://zenodo.org/record/10202981/files/figure.png" pageId="19" pageNumber="174">Fig. 14B</figureCitation>
) [in female
<typeStatus id="197188725F6C9D59BE29FBF61D15FBE4" box="[528,629,1028,1054]" pageId="19" pageNumber="174" type="paratype">paratype</typeStatus>
<collectionCode id="A0DBAE155F6C9D59BE45FBF61DD8FBE4" box="[636,696,1028,1054]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="19" pageNumber="174">KUZ</collectionCode>
Z4389, genital segment 1 as described for
<typeStatus id="197188725F6C9D59B888FBF11A7BFBE7" box="[1201,1307,1027,1053]" pageId="19" pageNumber="174" type="holotype">holotype</typeStatus>
, genital segment 2 absent; lamina subanalis situated between genital segment 1 and anal valves] (
<figureCitation id="5EF12A555F6C9D59B8EAFBD41A5FFBBA" box="[1235,1343,1062,1088]" captionStart="Fig" captionStartId="19.[189,232,731,757]" captionTargetBox="[189,1398,270,686]" captionTargetId="figure-376@19.[189,1398,270,686]" captionTargetPageId="19" captionText="Fig. 14. Scolopocryptops brevisulcatus sp. nov. AB. Holotype, ♂ (KUZ Z4390). C. Paratype, ♀ (KUZ Z4389). A. Right ultimate leg, lateral view. B. Male genital segments, lamina subanalis, and anal valves, ventral view. C. Female genital segment, lamina subanalis, and anal valves, ventral view. Abbreviations: av = anal valve; ls = lamina subanalis; sgs I = sternite of genital segment 1; sgs II = sternite of genital segment 2. Scale bars: A = 2 mm; BC = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10202981" httpUri="https://zenodo.org/record/10202981/files/figure.png" pageId="19" pageNumber="174">Fig. 14C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="92B566585F6D9D58BC84F8EC1ED0F813" ID-DOI="http://doi.org/10.5281/zenodo.10202979" ID-Zenodo-Dep="10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="18" pageNumber="173" startId="18.[189,232,1822,1848]" targetBox="[189,1398,1004,1759]" targetPageId="18">
<paragraph id="C67536D05F6D9D58BC84F8EC1ED0F813" blockId="18.[188,1399,1822,2025]" pageId="18" pageNumber="173">
<emphasis id="F4BEEAC25F6D9D58BC84F8EC1E7CF8C2" bold="true" box="[189,284,1822,1848]" pageId="18" pageNumber="173">Fig. 13.</emphasis>
<taxonomicName id="01CA4D535F6D9D58BD1FF8EC1DEEF8C2" authority="Jonishi &amp; Nakano, 2023" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[294,654,1822,1848]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="18" pageNumber="173" phylum="Arthropoda" rank="species" species="brevisulcatus" status="sp. nov.">
<emphasis id="F4BEEAC25F6D9D58BD1FF8EC1DEEF8C2" box="[294,654,1822,1848]" italics="true" pageId="18" pageNumber="173">Scolopocryptops brevisulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF8D57B95F6D9D58BEA1F8ED1D95F8C3" box="[664,757,1823,1849]" pageId="18" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="F4BEEAC25F6D9D58BF39F8EC1C1DF8C2" bold="true" box="[768,893,1822,1848]" pageId="18" pageNumber="173">AB, DE</emphasis>
. Holotype, ♂ (KUZ Z4390).
<emphasis id="F4BEEAC25F6D9D58B8DBF8EC1B97F8C2" bold="true" box="[1250,1271,1822,1848]" pageId="18" pageNumber="173">C</emphasis>
. Paratype, sex undetermined (KUZ Z4387).
<emphasis id="F4BEEAC25F6D9D58BE68F8B31D04F8A1" bold="true" box="[593,612,1857,1883]" pageId="18" pageNumber="173">A</emphasis>
. Cephalic plate and tergites 18, dorsal view.
<emphasis id="F4BEEAC25F6D9D58B8AAF8B01BC4F8A6" bold="true" box="[1171,1188,1858,1884]" pageId="18" pageNumber="173">B</emphasis>
. Sternites 1017, ventral view.
<emphasis id="F4BEEAC25F6D9D58BD60F8961E0EF884" bold="true" box="[345,366,1892,1918]" pageId="18" pageNumber="173">C</emphasis>
. Sternite 23, ventral view.
<emphasis id="F4BEEAC25F6D9D58BE9CF8971DDAF885" bold="true" box="[677,698,1893,1919]" pageId="18" pageNumber="173">D</emphasis>
. Right coxopleuron, lateral view.
<emphasis id="F4BEEAC25F6D9D58B87CF8971B39F885" bold="true" box="[1093,1113,1893,1919]" pageId="18" pageNumber="173">E</emphasis>
. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm.
</paragraph>
</caption>
<caption id="92B566585F6C9D59BC84FD291C66FC79" ID-DOI="http://doi.org/10.5281/zenodo.10202981" ID-Zenodo-Dep="10202981" httpUri="https://zenodo.org/record/10202981/files/figure.png" pageId="19" pageNumber="174" startId="19.[189,232,731,757]" targetBox="[189,1398,270,686]" targetPageId="19">
<paragraph id="C67536D05F6C9D59BC84FD291C66FC79" blockId="19.[188,1399,731,899]" pageId="19" pageNumber="174">
<emphasis id="F4BEEAC25F6C9D59BC84FD291E7AFD0F" bold="true" box="[189,282,731,757]" pageId="19" pageNumber="174">Fig. 14.</emphasis>
<taxonomicName id="01CA4D535F6C9D59BD19FD291DE5FD0F" authority="Jonishi &amp; Nakano, 2023" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[288,645,731,757]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="brevisulcatus" status="sp. nov.">
<emphasis id="F4BEEAC25F6C9D59BD19FD291DE5FD0F" box="[288,645,731,757]" italics="true" pageId="19" pageNumber="174">Scolopocryptops brevisulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF8D57B95F6C9D59BEB4FD2E1D87FD0C" box="[653,743,732,758]" pageId="19" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="F4BEEAC25F6C9D59BED5FD291C44FD0F" bold="true" box="[748,804,731,757]" pageId="19" pageNumber="174">AB</emphasis>
. Holotype, ♂ (KUZ Z4390).
<emphasis id="F4BEEAC25F6C9D59B845FD291BF1FD0F" bold="true" box="[1148,1169,731,757]" pageId="19" pageNumber="174">C</emphasis>
. Paratype, ♀ (KUZ Z4389).
<emphasis id="F4BEEAC25F6C9D59BD18FD0C1E56FCE2" bold="true" box="[289,310,766,792]" pageId="19" pageNumber="174">A</emphasis>
. Right ultimate leg, lateral view.
<emphasis id="F4BEEAC25F6C9D59BE8FFD0D1DAAFCE3" bold="true" box="[694,714,767,793]" pageId="19" pageNumber="174">B</emphasis>
. Male genital segments, lamina subanalis, and anal valves, ventral view.
<emphasis id="F4BEEAC25F6C9D59BD61FCD31E0DFCC1" bold="true" box="[344,365,801,827]" pageId="19" pageNumber="174">C</emphasis>
. Female genital segment, lamina subanalis, and anal valves, ventral view. Abbreviations: av = anal valve; ls = lamina subanalis; sgs I = sternite of genital segment 1; sgs II = sternite of genital segment 2. Scale bars: A = 2 mm; BC = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="8ED0655B5F6C9D59BC84FB9A1DA8F9AD" pageId="19" pageNumber="174" type="discussion">
<paragraph id="C67536D05F6C9D59BC84FB9A1E4FFB78" blockId="19.[189,303,1128,1154]" box="[189,303,1128,1154]" pageId="19" pageNumber="174">
<heading id="9D3D81BC5F6C9D59BC84FB9A1E4FFB78" bold="true" box="[189,303,1128,1154]" fontSize="11" level="3" pageId="19" pageNumber="174" reason="3">
<emphasis id="F4BEEAC25F6C9D59BC84FB9A1E4FFB78" bold="true" box="[189,303,1128,1154]" pageId="19" pageNumber="174">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C67536D05F6C9D59BC84FB641DA8F9AD" blockId="19.[187,1400,1174,1623]" pageId="19" pageNumber="174">
This species has previously been identified as
<taxonomicName id="01CA4D535F6C9D59BF3AFB651CE0FB4A" box="[771,896,1174,1201]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="F4BEEAC25F6C9D59BF3AFB651CE0FB4A" box="[771,896,1174,1201]" italics="true" pageId="19" pageNumber="174">S. elegans</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="A25B4B215F6C9D59BFECFB651B0FFB4A" author="Omine T." box="[981,1135,1174,1203]" pageId="19" pageNumber="174" pagination="37 - 39" refId="ref13336" refString="Omine T. 1987. [Chilopods of Kunigami district]. In: Okinawa Prefectural Board of Education (ed.) Special Animals of Kunigami District in Okinawa Island: 37 - 39. Okinawa Prefectural Board of Education, Naha. [In Japanese.]" type="book chapter" year="1987">Ômine 1987</bibRefCitation>
,
<bibRefCitation id="A25B4B215F6C9D59B8B8FB641BDDFB4A" author="Omine T." box="[1153,1213,1174,1200]" pageId="19" pageNumber="174" pagination="151 - 162" refId="ref13385" refString="Omine T. 2002. [Characteristics of the distribution of myriapods in the Ryukyu Archipelago]. In: Kimura M. (ed.) [Formation and Introduction of Organisms of the Ryukyu Archipelago]: 151 - 162. Okinawa Times, Naha. [In Japanese.]" type="book chapter" year="2002">2002</bibRefCitation>
) and
<taxonomicName id="01CA4D535F6C9D59B933FB651ECEFB2E" authority="(Omine &amp; Ito 1998)" baseAuthorityName="Omine &amp; Ito" baseAuthorityYear="1998" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="F4BEEAC25F6C9D59B933FB651A16FB4A" box="[1290,1398,1175,1201]" italics="true" pageId="19" pageNumber="174">S. curtus</emphasis>
(
<bibRefCitation id="A25B4B215F6C9D59BCFCFB4B1EC6FB2E" author="Omine T. &amp; Ito Y." box="[197,422,1209,1236]" pageId="19" pageNumber="174" pagination="131 - 159" refId="ref13438" refString="Omine T. &amp; Ito Y. 1998. Abundance and diversity of soil macrofauna of forests in Yanbaru, northern montane part of Okinawa Island, with special reference to removal of undergrowth. Okinawa Daigaku Kiyo 15: 131 - 159." type="journal article" year="1998">Omine &amp; Ito 1998</bibRefCitation>
)
</taxonomicName>
and is quite similar to
<taxonomicName id="01CA4D535F6C9D59BEF1FB481C5CFB29" authority="Jonishi &amp; Nakano, 2023" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[712,828,1210,1236]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="miyosii" status="sp. nov.">
<emphasis id="F4BEEAC25F6C9D59BEF1FB481C5CFB29" box="[712,828,1210,1236]" italics="true" pageId="19" pageNumber="174">S. miyosii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF8D57B95F6C9D59BF7AFB481CC0FB2E" box="[835,928,1210,1236]" pageId="19" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
(see above). However,
<taxonomicName id="01CA4D535F6C9D59B887FB481A18FB29" authority="Jonishi &amp; Nakano, 2023" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[1214,1400,1209,1236]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="brevisulcatus" status="sp. nov.">
<emphasis id="F4BEEAC25F6C9D59B887FB481A18FB29" box="[1214,1400,1209,1236]" italics="true" pageId="19" pageNumber="174">S. brevisulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF8D57B95F6C9D59BC84FB2F1E78FB0D" box="[189,280,1245,1271]" pageId="19" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished by the combination of the following features: basal 4 antennal articles of adults with sparse setae (
<figureCitation id="5EF12A555F6C9D59BDE5FAF21D25FAE0" box="[476,581,1280,1306]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[210,1387,1436,1868]" captionTargetId="figure-392@15.[189,1398,1421,1882]" captionTargetPageId="15" captionText="Fig. 10. Scolopocryptops brevisulcatus sp. nov., holotype, ♂ (KUZ Z4390). A. Basal articles of left antenna, dorsal view. B. Cephalic plate, dorsal view; arrowhead indicates left cephalic marginal sulcus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202971" httpUri="https://zenodo.org/record/10202971/files/figure.png" pageId="19" pageNumber="174">Fig. 10A</figureCitation>
) (vs only basalmost article sparsely setose or all articles densely setose in
<taxonomicName id="01CA4D535F6C9D59BCE2FAD11E33FAC7" box="[219,339,1315,1341]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="F4BEEAC25F6C9D59BCE2FAD11E33FAC7" box="[219,339,1315,1341]" italics="true" pageId="19" pageNumber="174">S. elegans</emphasis>
</taxonomicName>
, and basal 2 or 3 articles with sparse setae in
<taxonomicName id="01CA4D535F6C9D59BF56FAD11C81FAC7" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[879,993,1315,1341]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="miyosii">
<emphasis id="F4BEEAC25F6C9D59BF56FAD11C81FAC7" box="[879,993,1315,1341]" italics="true" pageId="19" pageNumber="174">S. miyosii</emphasis>
</taxonomicName>
), the presence of short sulci along the lateral margin of the cephalic plate in adults (
<figureCitation id="5EF12A555F6C9D59BF34FAB41C18FA9A" box="[781,888,1350,1376]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[210,1387,1436,1868]" captionTargetId="figure-392@15.[189,1398,1421,1882]" captionTargetPageId="15" captionText="Fig. 10. Scolopocryptops brevisulcatus sp. nov., holotype, ♂ (KUZ Z4390). A. Basal articles of left antenna, dorsal view. B. Cephalic plate, dorsal view; arrowhead indicates left cephalic marginal sulcus. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202971" httpUri="https://zenodo.org/record/10202971/files/figure.png" pageId="19" pageNumber="174">Fig. 10B</figureCitation>
) (vs sulci absent in
<taxonomicName id="01CA4D535F6C9D59B854FAB51BB3FA9A" baseAuthorityName="Omine &amp; Ito" baseAuthorityYear="1998" box="[1133,1235,1351,1377]" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="F4BEEAC25F6C9D59B854FAB51BB3FA9A" box="[1133,1235,1351,1377]" italics="true" pageId="19" pageNumber="174">S. curtus</emphasis>
</taxonomicName>
), the anterior margin of coxosternite almost lacking median diastema (
<figureCitation id="5EF12A555F6C9D59BF63FA9B1CAAFA7E" box="[858,970,1385,1412]" captionStart="Fig" captionStartId="16.[189,232,1636,1662]" captionTargetBox="[189,1398,265,1570]" captionTargetId="figure-17@16.[189,1398,265,1570]" captionTargetPageId="16" captionText="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="19" pageNumber="174">Figs 11D</figureCitation>
,
<figureCitation id="5EF12A555F6C9D59BFEFFA981B6AFA7E" box="[982,1034,1386,1412]" captionStart="Fig" captionStartId="17.[189,232,1773,1799]" captionTargetBox="[230,1369,322,1671]" captionTargetId="figure-11@17.[189,1398,280,1693]" captionTargetPageId="17" captionText="Fig. 12. Scolopocryptops brevisulcatus sp. nov. A. Holotype, ♂ (KUZ Z4390). B. Paratype, subadult, sex undetermined (KUZ Z4386). Forcipules and forcipular coxosternite, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; md = median diastema; op = outer part of anterior margin of forcipular coxosternite; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A = 1 mm; B = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202977" httpUri="https://zenodo.org/record/10202977/files/figure.png" pageId="19" pageNumber="174">12A</figureCitation>
) (vs median diastema apparent in
<taxonomicName id="01CA4D535F6C9D59BCE7FA7F1E37FA5C" box="[222,343,1420,1447]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="F4BEEAC25F6C9D59BCE7FA7F1E37FA5C" box="[222,343,1420,1447]" italics="true" pageId="19" pageNumber="174">S. elegans</emphasis>
</taxonomicName>
,
<taxonomicName id="01CA4D535F6C9D59BD51FA7F1EBDFA5C" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[360,477,1421,1447]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="miyosii">
<emphasis id="F4BEEAC25F6C9D59BD51FA7F1EBDFA5C" box="[360,477,1421,1447]" italics="true" pageId="19" pageNumber="174">S. miyosii</emphasis>
</taxonomicName>
, and
<taxonomicName id="01CA4D535F6C9D59BE1DFA7F1DEBFA5C" baseAuthorityName="Omine &amp; Ito" baseAuthorityYear="1998" box="[548,651,1421,1447]" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="F4BEEAC25F6C9D59BE1DFA7F1DEBFA5C" box="[548,651,1421,1447]" italics="true" pageId="19" pageNumber="174">S. curtus</emphasis>
</taxonomicName>
), the sclerotized bands of coxosternite not reaching outer part of anterior margin (
<figureCitation id="5EF12A555F6C9D59BD91FA421D79FA30" box="[424,537,1456,1482]" captionStart="Fig" captionStartId="16.[189,232,1636,1662]" captionTargetBox="[189,1398,265,1570]" captionTargetId="figure-17@16.[189,1398,265,1570]" captionTargetPageId="16" captionText="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="19" pageNumber="174">Figs 11D</figureCitation>
,
<figureCitation id="5EF12A555F6C9D59BE1EFA421D3AFA30" box="[551,602,1456,1482]" captionStart="Fig" captionStartId="17.[189,232,1773,1799]" captionTargetBox="[230,1369,322,1671]" captionTargetId="figure-11@17.[189,1398,280,1693]" captionTargetPageId="17" captionText="Fig. 12. Scolopocryptops brevisulcatus sp. nov. A. Holotype, ♂ (KUZ Z4390). B. Paratype, subadult, sex undetermined (KUZ Z4386). Forcipules and forcipular coxosternite, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; md = median diastema; op = outer part of anterior margin of forcipular coxosternite; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A = 1 mm; B = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202977" httpUri="https://zenodo.org/record/10202977/files/figure.png" pageId="19" pageNumber="174">12A</figureCitation>
) (vs sclerotized bands almost reaching outer part in
<taxonomicName id="01CA4D535F6C9D59B8F5FA421A25FA30" box="[1228,1349,1456,1482]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="F4BEEAC25F6C9D59B8F5FA421A25FA30" box="[1228,1349,1456,1482]" italics="true" pageId="19" pageNumber="174">S. elegans</emphasis>
</taxonomicName>
and
<taxonomicName id="01CA4D535F6C9D59BC87FA211E4EFA17" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[190,302,1491,1517]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="miyosii">
<emphasis id="F4BEEAC25F6C9D59BC87FA211E4EFA17" box="[190,302,1491,1517]" italics="true" pageId="19" pageNumber="174">S. miyosii</emphasis>
</taxonomicName>
), the almost-reduced teeth on coxosternite (
<figureCitation id="5EF12A555F6C9D59BF16FA211CFEFA17" box="[815,926,1491,1517]" captionStart="Fig" captionStartId="16.[189,232,1636,1662]" captionTargetBox="[189,1398,265,1570]" captionTargetId="figure-17@16.[189,1398,265,1570]" captionTargetPageId="16" captionText="Fig. 11. Scolopocryptops brevisulcatus sp. nov. AD. Holotype, ♂ (KUZ Z4390). E. Paratype, subadult, sex undetermined (KUZ Z4386). A. Cephalic plate and tergite 1, dorsal view. B. Distal part of article 2, article 3, and pretarsus of left second maxilla, medial view. C. Article 3 and pretarsus of left second maxilla, lateral view. DE. Head, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; db = dorsal brush on article 3 of second maxilla; ds = dorsal spur on article 2 of second maxilla; pt = pretarsus of second maxilla; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A, DE = 1 mm; BC = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202975" httpUri="https://zenodo.org/record/10202975/files/figure.png" pageId="19" pageNumber="174">Figs 11D</figureCitation>
,
<figureCitation id="5EF12A555F6C9D59BF90FA211CBDFA17" box="[937,989,1491,1517]" captionStart="Fig" captionStartId="17.[189,232,1773,1799]" captionTargetBox="[230,1369,322,1671]" captionTargetId="figure-11@17.[189,1398,280,1693]" captionTargetPageId="17" captionText="Fig. 12. Scolopocryptops brevisulcatus sp. nov. A. Holotype, ♂ (KUZ Z4390). B. Paratype, subadult, sex undetermined (KUZ Z4386). Forcipules and forcipular coxosternite, ventral view. Abbreviations: am = anterior margin of forcipular coxosternite; bs = basal suture on forcipular trochanteroprefemoral process; md = median diastema; op = outer part of anterior margin of forcipular coxosternite; ptr = process of forcipular trochanteroprefemur; t = tooth on anterior margin of forcipular coxosternite. Scale bars: A = 1 mm; B = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10202977" httpUri="https://zenodo.org/record/10202977/files/figure.png" pageId="19" pageNumber="174">12A</figureCitation>
) (vs teeth prominent in
<taxonomicName id="01CA4D535F6C9D59B8CAFA211A05FA17" authorityName="Jonishi &amp; Nakano" authorityYear="2023" box="[1267,1381,1491,1517]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="miyosii">
<emphasis id="F4BEEAC25F6C9D59B8CAFA211A05FA17" box="[1267,1381,1491,1517]" italics="true" pageId="19" pageNumber="174">S. miyosii</emphasis>
</taxonomicName>
), the dorsal margin of the ultimate pleuron protruding from the lateral margin of tergite 23 (
<figureCitation id="5EF12A555F6C9D59B93DFA041A0EF9EA" box="[1284,1390,1526,1552]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="19" pageNumber="174">Fig. 13E</figureCitation>
) (vs dorsal margin flattened in
<taxonomicName id="01CA4D535F6C9D59BE1FF9E81DEFF9C9" baseAuthorityName="Omine &amp; Ito" baseAuthorityYear="1998" box="[550,655,1562,1588]" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="curtus">
<emphasis id="F4BEEAC25F6C9D59BE1FF9E81DEFF9C9" box="[550,655,1562,1588]" italics="true" pageId="19" pageNumber="174">S. curtus</emphasis>
</taxonomicName>
) and the short coxopleural process (
<figureCitation id="5EF12A555F6C9D59B87CF9EB1BD2F9CE" box="[1093,1202,1561,1588]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[189,1398,1004,1759]" captionTargetId="figure-235@18.[189,1398,1004,1759]" captionTargetPageId="18" captionText="Fig. 13. Scolopocryptops brevisulcatus sp. nov. AB, DE. Holotype, ♂ (KUZ Z4390). C. Paratype, sex undetermined (KUZ Z4387). A. Cephalic plate and tergites 18, dorsal view. B. Sternites 1017, ventral view.C. Sternite 23, ventral view. D. Right coxopleuron, lateral view. E. Tergite 23, dorsal view. Abbreviations: cxp = coxopleural process; dm = dorsal margin of ultimate pleuron; mds = minute dark spine on ultimate pleuron; tm = tergal margination; tps = tergal paramedian suture. Scale bars: AB = 5 mm; CE = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202979" httpUri="https://zenodo.org/record/10202979/files/figure.png" pageId="19" pageNumber="174">Fig. 13D</figureCitation>
) (vs the process moderately long in
<taxonomicName id="01CA4D535F6C9D59BD9BF9CF1D79F9AC" box="[418,537,1596,1623]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="19" pageNumber="174" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="F4BEEAC25F6C9D59BD9BF9CF1D79F9AC" box="[418,537,1596,1623]" italics="true" pageId="19" pageNumber="174">S. elegans</emphasis>
</taxonomicName>
) (see
<tableCitation id="8B48036B5F6C9D59BE59F9CE1DD8F9AD" box="[608,696,1596,1623]" captionStart="Table 3" captionStartId="20.[185,255,622,650]" captionText="Table 3. Morphological comparison of the four closely related species of Scolopocryptops Newport, 1844 [variations are indicated in brackets]. Diagnostic characters of S. elegans (Takakuwa, 1937) and S. curtus (Takakuwa, 1939) mainly follow Jonishi &amp; Nakano (2022)." pageId="19" pageNumber="174">Table 3</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8ED0655B5F6C9D59BC84F98C1BE0F93C" box="[189,1152,1662,1735]" pageId="19" pageNumber="174" type="distribution">
<paragraph id="C67536D05F6C9D59BC84F98C1E36F962" blockId="19.[189,342,1662,1688]" box="[189,342,1662,1688]" pageId="19" pageNumber="174">
<heading id="9D3D81BC5F6C9D59BC84F98C1E36F962" bold="true" box="[189,342,1662,1688]" fontSize="11" level="3" pageId="19" pageNumber="174" reason="3">
<emphasis id="F4BEEAC25F6C9D59BC84F98C1E36F962" bold="true" box="[189,342,1662,1688]" pageId="19" pageNumber="174">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C67536D05F6C9D59BC84F95F1BE0F93C" blockId="19.[189,1152,1708,1735]" box="[189,1152,1708,1735]" pageId="19" pageNumber="174">
Known from
<collectingRegion id="040EF8325F6C9D59BD62F95E1EA5F93C" box="[347,453,1708,1734]" country="Japan" name="Okinawa" pageId="19" pageNumber="174">Okinawa</collectingRegion>
Island and its adjacent islet (Izena Island), Ryukyu Islands.
</paragraph>
</subSubSection>
</treatment>
</document>