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<document id="5294F6C52F8B77012F6F1E38A17F2DCB" ID-CLB-Dataset="7182" ID-DOI="10.11646/zootaxa.4963.1.5" ID-GBIF-Dataset="ee9bfd7b-3b05-4403-9af2-e54beaae5e69" ID-ISSN="1175-5326" ID-Zenodo-Dep="4719924" ID-ZooBank="6B4B8F71-7B08-4027-A76B-7EE8C987C9BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618562743172" checkinUser="plazi" docAuthor="Zhu, Wenhui &amp; Li, Shuqiang" docDate="2021" docId="4E3C87F3FFC5FF96C6ECFCE8FC3AFF7A" docLanguage="en" docName="zootaxa.4963.1.5.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana cucphuong Zhu &amp; Li 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="122" masterDocId="B205FF8BFFC1FF91C67BFFC1FFB6FF80" masterDocTitle="Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia" masterLastPageNumber="137" masterPageNumber="115" pageNumber="119" updateTime="1698937141523" updateUser="ExternalLinkService">
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<mods:titleInfo id="3E0EA1CA481B3D0A469EB52174ADE642">
<mods:title id="2FBC85BEC0D55BCEFE030FEFDCD7EF90">Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name id="DFAE34DBD2193A8ACF0B0E133DC513AA" type="personal">
<mods:role id="64DC2FCB2CAB12914239F12FD0A6CDD4">
<mods:roleTerm id="F7EA66D1884A33BB2BF88028FFF1E79E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8D2E951718856464B26DAD9A9A0B7B13">Zhu, Wenhui</mods:namePart>
<mods:nameIdentifier id="90255A1FF7625286ACA6213CD2744FEC" type="ORCID">0000-0001-5650-1670</mods:nameIdentifier>
<mods:affiliation id="93204D80DE93C6E850A05E4BDDF23FC8">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar &amp; zhuwh 1016 @ outlook. com; https: // orcid. org / 0000 - 0001 - 5650 - 1670</mods:affiliation>
<mods:nameIdentifier id="953007050969D351A8D2A3764B80199C" type="email">zhuwh1016@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:name id="F7A25BDB82CE852BF5140ACDF8E290C7" type="personal">
<mods:role id="5B85DF6B73F4BE13EE22E2B4A736207A">
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</mods:role>
<mods:namePart id="E2325D5E663144A6A385E2D8D7A6B423">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="AF98F1F303EFB18300378190BEDFB1E1" type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="E9786D3313CA9F5F07883FFCEAF79B97">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; lisq @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 3290 - 5416</mods:affiliation>
<mods:nameIdentifier id="CD130649F8E023D61BFF4AC7ECCDA71D" type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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<mods:title id="2A66A2EA3F9B14368EB1AA70B13EFA07">Zootaxa</mods:title>
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<mods:part id="0E6CD44DA825E8FE22D896DD0747B2DA">
<mods:date id="D21B0DE01548B3D315F5040822D3E46D">2021</mods:date>
<mods:detail id="FE1AF72A707745129A57FA48F35FA0F3" type="pubDate">
<mods:number id="A2A87F339CD7B12D591D723AE695F590">2021-04-16</mods:number>
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<mods:detail id="52E6AB4C5E55EDAE75E92BD253A6C79C" type="volume">
<mods:number id="3E4C74EB4C65A9B0369A21977BFDC6A8">4963</mods:number>
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<treatment id="4E3C87F3FFC5FF96C6ECFCE8FC3AFF7A" ID-DOI="http://doi.org/10.5281/zenodo.4696998" ID-GBIF-Taxon="181600553" ID-Zenodo-Dep="4696998" LSID="urn:lsid:plazi:treatment:4E3C87F3FFC5FF96C6ECFCE8FC3AFF7A" httpUri="http://treatment.plazi.org/id/4E3C87F3FFC5FF96C6ECFCE8FC3AFF7A" lastPageId="7" lastPageNumber="122" pageId="4" pageNumber="119">
<subSubSection id="8E8F656EFFC5FF95C6ECFCE8FE43FCC3" box="[151,501,809,836]" pageId="4" pageNumber="119" type="nomenclature">
<paragraph id="C62A36E5FFC5FF95C6ECFCE8FE43FCC3" blockId="4.[151,501,809,871]" box="[151,501,809,836]" pageId="4" pageNumber="119">
<heading id="9D628189FFC5FF95C6ECFCE8FE43FCC3" bold="true" box="[151,501,809,836]" fontSize="11" level="1" pageId="4" pageNumber="119" reason="1">
<emphasis id="F4E1EAF7FFC5FF95C6ECFCE8FE43FCC3" bold="true" box="[151,501,809,836]" pageId="4" pageNumber="119">
<taxonomicName id="01954D66FFC5FF95C6ECFCE8FE39FCC4" ID-CoL="98YXV" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[151,399,809,836]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="cucphuong" status="sp. nov.">
<emphasis id="F4E1EAF7FFC5FF95C6ECFCE8FE39FCC4" bold="true" box="[151,399,809,836]" italics="true" pageId="4" pageNumber="119">Belisana cucphuong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFD2578CFFC5FF95C7ECFCE8FE43FCC3" box="[407,501,809,835]" pageId="4" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC5FF95C6ECFC8DFE45FCE7" box="[151,499,844,871]" pageId="4" pageNumber="119" type="description">
<paragraph id="C62A36E5FFC5FF95C6ECFC8DFE45FCE7" blockId="4.[151,501,809,871]" box="[151,499,844,871]" pageId="4" pageNumber="119">
<figureCitation id="5EAE2A60FFC5FF95C6ECFC8DFF4DFCE6" box="[151,251,844,870]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1806,1830]" captionStartId-1="6.[151,250,1875,1899]" captionTargetBox-0="[249,1339,180,1783]" captionTargetBox-1="[226,1361,181,1851]" captionTargetId-0="figure-17@5.[249,1339,180,1783]" captionTargetId-1="figure-19@6.[226,1361,181,1851]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." captionText-1="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi-0="http://doi.org/10.5281/zenodo.4719932" figureDoi-1="http://doi.org/10.5281/zenodo.4719936" httpUri-0="https://zenodo.org/record/4719932/files/figure.png" httpUri-1="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 3–4</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C77DFC8DFEECFCE6" box="[262,346,844,870]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13C–D</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C71EFC8DFE0FFCE6" box="[357,441,844,870]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14C–D</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C7BFFC8DFE45FCE7" box="[452,499,844,871]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="4" pageNumber="119">15B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC5FF95C6ECFC50FE45FC71" pageId="4" pageNumber="119" type="materials_examined">
<paragraph id="C62A36E5FFC5FF95C6ECFC50FE45FC71" blockId="4.[151,1437,913,2015]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6ECFC50FEF3FC2B" bold="true" box="[151,325,913,939]" pageId="4" pageNumber="119">Type material.</emphasis>
<materialsCitation id="76FD3CB8FFC5FF95C736FC50FC73FC4E" ID-GBIF-Occurrence="3091104310" collectingDate="2015-08-20" collectionCode="IZCAS" collectorName="D. S. Pham" country="Vietnam" elevation="437" latitude="20.349533" location="Male" longLatPrecision="1" longitude="105.59334" municipality="Cuc Phuong National Park" pageId="4" pageNumber="119" specimenCount="1" stateProvince="Ninh Binh" typeStatus="holotype">
<emphasis id="F4E1EAF7FFC5FF95C736FC50FE75FC2B" bold="true" box="[333,451,913,939]" pageId="4" pageNumber="119">
<typeStatus id="192E8847FFC5FF95C736FC50FE08FC2B" box="[333,446,913,939]" pageId="4" pageNumber="119" type="holotype">Holotype</typeStatus>
:
</emphasis>
<location id="C34A603EFFC5FF95C7B1FC50FDB2FC2B" LSID="urn:lsid:plazi:treatment:4E3C87F3FFC5FF96C6ECFCE8FC3AFF7A:C34A603EFFC5FF95C7B1FC50FDB2FC2B" box="[458,516,913,939]" country="Vietnam" latitude="20.349533" longLatPrecision="1" longitude="105.59334" municipality="Cuc Phuong National Park" name="Male" pageId="4" pageNumber="119" stateProvince="Ninh Binh">Male</location>
(
<collectionCode id="A084AE20FFC5FF95C468FC50FDD0FC2B" box="[531,614,913,939]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="119">IZCAS</collectionCode>
Ar41904),
<collectingMunicipality id="264EAC9FFFC5FF95C49DFC50FBA5FC2B" box="[742,1043,913,939]" pageId="4" pageNumber="119">Cuc Phuong National Park</collectingMunicipality>
(
<geoCoordinate id="A3A15022FFC5FF95C259FC50FB64FC2C" box="[1058,1234,913,940]" degrees="20" direction="north" minutes="20" orientation="latitude" pageId="4" pageNumber="119" precision="1" seconds="58.32" value="20.349533">20°20’58.32”N</geoCoordinate>
,
<geoCoordinate id="A3A15022FFC5FF95C2A5FC50FA2EFC2C" box="[1246,1432,913,940]" degrees="105" direction="east" minutes="35" orientation="longitude" pageId="4" pageNumber="119" precision="1" seconds="36.00" value="105.59334">105°35’36.00”E</geoCoordinate>
,
<elevation id="4DB8D1D6FFC5FF95C6ECFC75FEE7FC4F" box="[151,337,948,975]" metricMagnitude="2" metricUnit="m" metricValue="4.37" pageId="4" pageNumber="119" unit="m" value="437.0">
elevation
<quantity id="016D9B00FFC5FF95C77DFC75FEE7FC4F" box="[262,337,948,975]" metricMagnitude="2" metricUnit="m" metricValue="4.37" pageId="4" pageNumber="119" unit="m" value="437.0">437 m</quantity>
</elevation>
),
<collectingRegion id="0451F807FFC5FF95C71DFC75FE68FC4E" box="[358,478,948,974]" country="Vietnam" name="Ninh Binh" pageId="4" pageNumber="119">Ninh Binh</collectingRegion>
,
<collectingCountry id="BE827675FFC5FF95C791FC75FDFAFC4E" box="[490,588,948,974]" name="Vietnam" pageId="4" pageNumber="119">Vietnam</collectingCountry>
,
<date id="B22B1025FFC5FF95C423FC75FCB8FC4F" box="[600,782,948,975]" pageId="4" pageNumber="119" value="2015-08-20">
<collectingDate id="A26FE9CDFFC5FF95C423FC75FCB8FC4F" box="[600,782,948,975]" pageId="4" pageNumber="119" value="2015-08-20">20 August 2015</collectingDate>
</date>
,
<collectorName id="6B605333FFC5FF95C561FC75FC22FC4E" box="[794,916,948,974]" pageId="4" pageNumber="119">D.S. Pham</collectorName>
leg.
</materialsCitation>
<materialsCitation id="76FD3CB8FFC5FF95C5B7FC75FE59FC71" ID-GBIF-Occurrence="3091104301" collectingDate="2015-08-20" collectionCode="IZCAS" collectorName="D. S. Pham" country="Vietnam" elevation="437" latitude="20.349533" location="Male" longLatPrecision="1" longitude="105.59334" municipality="Cuc Phuong National Park" pageId="4" pageNumber="119" specimenCount="2" specimenCount-female="2" stateProvince="Ninh Binh" typeStatus="paratype">
<emphasis id="F4E1EAF7FFC5FF95C5B7FC75FBFBFC4E" bold="true" box="[972,1101,948,974]" pageId="4" pageNumber="119">
<typeStatus id="192E8847FFC5FF95C5B7FC75FBFEFC4E" box="[972,1096,948,974]" pageId="4" pageNumber="119" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="D093FD6CFFC5FF95C22EFC75FB74FC4E" box="[1109,1218,948,974]" pageId="4" pageNumber="119" type="female">2 females</specimenCount>
(
<collectionCode id="A084AE20FFC5FF95C2A9FC75FA93FC4E" box="[1234,1317,948,974]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="119">IZCAS</collectionCode>
Ar41905– 41906), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC5FF95C6BCFC38FBCAFB93" box="[199,1148,1017,1044]" pageId="4" pageNumber="119" type="etymology">
<paragraph id="C62A36E5FFC5FF95C6BCFC38FBCAFB93" blockId="4.[151,1437,913,2015]" box="[199,1148,1017,1044]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFC38FEF8FB93" bold="true" box="[199,334,1017,1043]" pageId="4" pageNumber="119">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="192E8847FFC5FF95C4CEFC3BFD53FB94" box="[693,741,1018,1044]" pageId="4" pageNumber="119">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC5FF95C6BCFBDDFE8EFA87" pageId="4" pageNumber="119" type="diagnosis">
<paragraph id="C62A36E5FFC5FF95C6BCFBDDFE8EFA87" blockId="4.[151,1437,913,2015]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFBDDFEF4FBB6" bold="true" box="[199,322,1052,1078]" pageId="4" pageNumber="119">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="01954D66FFC5FF95C434FBDCFBB6FBB7" authority="Yao, Pham &amp; Li, 2015" authorityName="Yao, Pham &amp; Li" authorityYear="2015" box="[591,1024,1052,1079]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="F4E1EAF7FFC5FF95C434FBDCFD49FBB6" box="[591,767,1053,1078]" italics="true" pageId="4" pageNumber="119">B. vietnamensis</emphasis>
<bibRefCitation id="A2044B14FFC5FF95C57FFBDCFBB6FBB7" author="Yao, Z. &amp; Pham, D. S. &amp; Li, S." box="[772,1024,1052,1079]" journalOrPublisher="Zootaxa" pageId="4" pageNumber="119" pagination="1 - 82" part="3909" refId="ref8999" refString="Yao, Z., Pham, D. S. &amp; Li, S. (2015) Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species. Zootaxa, 3909 (1), 1 - 82. https: // doi. org / 10.11646 / zootaxa. 3909.1.1" title="Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species" type="journal article" year="2015">Yao, Pham &amp; Li, 2015</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="A2044B14FFC5FF95C241FBDCFAA6FBB6" author="Yao, Z. &amp; Pham, D. S. &amp; Li, S." box="[1082,1296,1052,1079]" journalOrPublisher="Zootaxa" pageId="4" pageNumber="119" pagination="1 - 82" part="3909" refId="ref8999" refString="Yao, Z., Pham, D. S. &amp; Li, S. (2015) Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species. Zootaxa, 3909 (1), 1 - 82. https: // doi. org / 10.11646 / zootaxa. 3909.1.1" title="Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species" type="journal article" year="2015">
Yao
<emphasis id="F4E1EAF7FFC5FF95C216FBDCFB28FBB6" box="[1133,1182,1052,1078]" italics="true" pageId="4" pageNumber="119">et al</emphasis>
. 2015: 10
</bibRefCitation>
, figs 29–31) with the elongated bulbal apophyses (
<figureCitation id="5EAE2A60FFC5FF95C433FBFEFD17FBD9" box="[584,673,1087,1113]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Fig. 4D</figureCitation>
) and the male chelicerae (
<figureCitation id="5EAE2A60FFC5FF95C5AAFBFEFB83FBD9" box="[977,1077,1087,1113]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs. 4C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C238FBFFFBC2FBD9" box="[1091,1140,1086,1113]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="4" pageNumber="119">15B</figureCitation>
), but males can be distinguished by the long distal spine, 7 times as long as wide (arrow
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<figureCitation id="5EAE2A60FFC5FF95C5E6FBA0FC4EFBFB" box="[925,1016,1121,1147]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C27FFBA0FB83FBFB" box="[1028,1077,1121,1147]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C23AFBA0FBF9FBFB" box="[1089,1103,1121,1147]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">4</figureCitation>
times in
<taxonomicName id="01954D66FFC5FF95C2C3FBA3FADFFBFB" authorityName="Yao, Pham &amp; Li" authorityYear="2015" box="[1208,1385,1122,1147]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="F4E1EAF7FFC5FF95C2C3FBA3FADFFBFB" box="[1208,1385,1122,1147]" italics="true" pageId="4" pageNumber="119">B. vietnamensis</emphasis>
</taxonomicName>
), on the procursus, and by the presence of a long ventral apophysis on the palpal trochanter (arrow
<quantity id="016D9B00FFC5FF95C2ADFB45FAB5FB1E" box="[1238,1283,1156,1182]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="119" unit="in" value="1.0">1 in</quantity>
<figureCitation id="5EAE2A60FFC5FF95C377FB45FAD4FB1E" box="[1292,1378,1156,1182]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Fig. 3B</figureCitation>
) and distinct ventral apophysis on palpal femur (arrow
<quantity id="016D9B00FFC5FF95C4A9FB66FD49FB41" box="[722,767,1191,1217]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="119" unit="in" value="3.0">3 in</quantity>
<figureCitation id="5EAE2A60FFC5FF95C57CFB66FCE9FB41" box="[775,863,1191,1217]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Fig. 3B</figureCitation>
). Females can be distinguished by the vulval pore plates (arrow pp in
<figureCitation id="5EAE2A60FFC5FF95C70EFB08FE66FB63" box="[373,464,1225,1251]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C7A6FB08FDB9FB63" box="[477,527,1225,1251]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14D</figureCitation>
) being longer than wide (vulval pore plates approximately as wide as long in
<taxonomicName id="01954D66FFC5FF95C3FFFB0BFE9EFA86" authorityName="Yao, Pham &amp; Li" authorityYear="2015" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="F4E1EAF7FFC5FF95C3FFFB0BFE9EFA86" italics="true" pageId="4" pageNumber="119">B. vietnamensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC5FF96C6BCFACFFB67FF55" lastPageId="7" lastPageNumber="122" pageId="4" pageNumber="119" type="description">
<paragraph id="C62A36E5FFC5FF95C6BCFACFFE23F96C" blockId="4.[151,1437,913,2015]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFACFFE29FAA9" bold="true" box="[199,415,1294,1321]" pageId="4" pageNumber="119">Description. Male</emphasis>
(
<emphasis id="F4E1EAF7FFC5FF95C7C9FACEFDAFFAA9" bold="true" box="[434,537,1295,1321]" pageId="4" pageNumber="119">
<typeStatus id="192E8847FFC5FF95C7C9FACEFDAFFAA9" box="[434,537,1295,1321]" pageId="4" pageNumber="119" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar41904): Total length 2.22 (2.35 with clypeus), carapace 0.94 long, 0.78 wide, opisthosoma 1.41 long, 0.78 wide. Leg I: – (5.70 + 0.38 + 5.77 + – + –), leg II: 13.00 (3.97 + 0.32 + 3.33 + 4.10 + 1.28), leg III: 8.28 (2.50 + 0.26 + 1.99 + 2.82 + 0.71), leg IV: 14.15 (6.03 + 0.30 + 2.88 + 3.85 + 1.09); tibia I L/d: 38. Distance PME-PME 0.09, diameter PME 0.09, distance PME-ALE 0.09, AME absent. Habitus as in
<figureCitation id="5EAE2A60FFC5FF95C6ECFA58FEB8FA33" box="[151,270,1433,1459]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4E–F</figureCitation>
. Carapace pale, with brown lateral margins. Ocular area not elevated. Clypeus unmodified. Chelicerae (
<figureCitation id="5EAE2A60FFC5FF95C6E4FA7DFEB5FA56" box="[159,259,1468,1494]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs. 4C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C774FA7DFE88FA57" box="[271,318,1468,1495]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="4" pageNumber="119">15B</figureCitation>
) with pair of small proximo-lateral apophyses and pair of short distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.04). Sternum pale, approximately as wide as long (0.50). Thoracic furrow absent. Legs pale. Opisthosoma pale, with large, dark spots. Palps as in
<figureCitation id="5EAE2A60FFC5FF95C362F9C0FA2EF99B" box="[1305,1432,1537,1563]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3A–B</figureCitation>
; coxa unmodified; trochanter with long ventral apophysis (arrow
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<figureCitation id="5EAE2A60FFC5FF95C5C9F9E5FBBDF9BE" box="[946,1035,1572,1598]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Fig. 3B</figureCitation>
); femur with small proximo-dorsal apophysis (arrow
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<figureCitation id="5EAE2A60FFC5FF95C7E9F986FE51F9E1" box="[402,487,1607,1633]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Fig. 3B</figureCitation>
) and distinct ventral apophysis (arrow
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<figureCitation id="5EAE2A60FFC5FF95C5B4F986FB93F9E1" box="[975,1061,1607,1633]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Fig. 3B</figureCitation>
); procursus (
<figureCitation id="5EAE2A60FFC5FF95C2CCF986FA80F9E1" box="[1207,1334,1607,1633]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3A–D</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C339F986FA22F9E1" box="[1346,1428,1607,1633]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13C–D</figureCitation>
) simple proximally but complex distally, with large subdisto-prolateral sclerite (arrow
<quantity id="016D9B00FFC5FF95C212F9A8FB20F903" box="[1129,1174,1641,1667]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="119" unit="in" value="1.0">1 in</quantity>
<figureCitation id="5EAE2A60FFC5FF95C2E5F9A8FB4CF903" box="[1182,1274,1641,1667]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C373F9A8FA8FF903" box="[1288,1337,1641,1667]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13C</figureCitation>
), curved distal spine (arrow
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<figureCitation id="5EAE2A60FFC5FF95C7E4F94DFE4FF926" box="[415,505,1676,1702]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C47EF94DFD82F926" box="[517,564,1676,1702]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13C</figureCitation>
) and subdisto-retrolateral membranous process (arrow in
<figureCitation id="5EAE2A60FFC5FF95C2C0F94DFAA1F926" box="[1211,1303,1676,1702]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3D</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C359F94DFAE0F926" box="[1314,1366,1676,1702]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13D</figureCitation>
); bulb (
<figureCitation id="5EAE2A60FFC5FF95C6DBF96EFF40F949" box="[160,246,1711,1737]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Fig. 4D</figureCitation>
) with hooked apophysis and simple embolus. Retrolateral trichobothria of tibia Iat 5% proximally; without spines or curved setae.
</paragraph>
<paragraph id="C62A36E5FFC5FF95C6BCF935FC88F85E" blockId="4.[151,1437,913,2015]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCF935FEA8F88E" bold="true" box="[199,286,1780,1806]" pageId="4" pageNumber="119">Female</emphasis>
(IZCAS Ar41906): Similar to male, habitus as in
<figureCitation id="5EAE2A60FFC5FF95C51BF935FC57F88E" box="[864,993,1780,1806]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4G–H</figureCitation>
. Total length 1.90 (2.00 with clypeus), carapace 0.72 long, 0.66 wide, opisthosoma 1.28 long, 0.66 wide. Tibia I: 4.04; tibia I L/d: 51. Distance PME-PME 0.13, diameter PME 0.06, distance PME-ALE 0.09, AME absent. Sternum slightly wider than long (0.50/0.40). Opisthosoma with indistinct dark spots. External female genitalia (
<figureCitation id="5EAE2A60FFC5FF95C5DBF89DFBB7F8F6" box="[928,1025,1884,1910]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4A</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C26BF89DFBF6F8F6" box="[1040,1088,1884,1910]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14C</figureCitation>
) simple and flat, with pair of frontal “epigynal” pockets on sclerotized plate (arrow ep in
<figureCitation id="5EAE2A60FFC5FF95C542F8BEFC00F819" box="[825,950,1919,1945]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4A–B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C5B9F8BEFBA0F819" box="[962,1046,1919,1945]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14C–D</figureCitation>
), 0.03 apart. Vulva (
<figureCitation id="5EAE2A60FFC5FF95C280F8BEFAE0F819" box="[1275,1366,1919,1945]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C318F8BEFA22F819" box="[1379,1428,1919,1945]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14D</figureCitation>
) with wavy anterior arch (median ridge strongly protruding, arrow in
<figureCitation id="5EAE2A60FFC5FF95C5E9F860FC5AF83B" box="[914,1004,1953,1979]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C58CF860FB9FF83B" box="[1015,1065,1953,1979]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14D</figureCitation>
) and pair of nearly elliptical pore plates (arrow pp in
<figureCitation id="5EAE2A60FFC5FF95C70AF805FE7DF85E" box="[369,459,1988,2014]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (C–F) and paratype female (A–B, G–H).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C7ACF805FDBCF85E" box="[471,522,1988,2014]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. A–B. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; C–D. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov., arrow points at extending part of anterior arch; I–J. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14D</figureCitation>
), without serrated sclerites.
</paragraph>
<caption id="92EA666DFFC4FF94C6ECF8CFFC26F837" ID-DOI="http://doi.org/10.5281/zenodo.4719932" ID-Zenodo-Dep="4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="5" pageNumber="120" startId="5.[151,250,1806,1830]" targetBox="[249,1339,180,1783]" targetPageId="5">
<paragraph id="C62A36E5FFC4FF94C6ECF8CFFC26F837" blockId="5.[151,1437,1806,1975]" pageId="5" pageNumber="120">
<emphasis id="F4E1EAF7FFC4FF94C6ECF8CFFEA5F8A7" bold="true" box="[151,275,1806,1831]" pageId="5" pageNumber="120">FIGURE 3.</emphasis>
<taxonomicName id="01954D66FFC4FF94C762F8CEFE51F8A7" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[281,487,1807,1831]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="120" phylum="Arthropoda" rank="species" species="cucphuong" status="sp. nov.">
<emphasis id="F4E1EAF7FFC4FF94C762F8CEFE51F8A7" box="[281,487,1807,1831]" italics="true" pageId="5" pageNumber="120">Belisana cucphuong</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFC4FF94C796F8CEFD88F8A7" bold="true" box="[493,574,1807,1831]" pageId="5" pageNumber="120">
<taxonomicNameLabel id="EFD2578CFFC4FF94C796F8CEFD88F8A7" box="[493,574,1807,1831]" pageId="5" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A–B. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D).
</paragraph>
</caption>
<caption id="92EA666DFFC7FF97C6ECF892FE7FF87B" ID-DOI="http://doi.org/10.5281/zenodo.4719936" ID-Zenodo-Dep="4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="6" pageNumber="121" startId="6.[151,250,1875,1899]" targetBox="[226,1361,181,1851]" targetPageId="6">
<paragraph id="C62A36E5FFC7FF97C6ECF892FE7FF87B" blockId="6.[151,1437,1875,2043]" pageId="6" pageNumber="121">
<emphasis id="F4E1EAF7FFC7FF97C6ECF892FEA4F8EB" bold="true" box="[151,274,1875,1899]" pageId="6" pageNumber="121">FIGURE 4.</emphasis>
<taxonomicName id="01954D66FFC7FF97C763F892FE50F8EB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[280,486,1875,1899]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="121" phylum="Arthropoda" rank="species" species="cucphuong" status="sp. nov.">
<emphasis id="F4E1EAF7FFC7FF97C763F892FE50F8EB" box="[280,486,1875,1899]" italics="true" pageId="6" pageNumber="121">Belisana cucphuong</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFC7FF97C790F892FD8DF8EB" bold="true" box="[491,571,1875,1899]" pageId="6" pageNumber="121">
<taxonomicNameLabel id="EFD2578CFFC7FF97C790F892FD8DF8EB" box="[491,571,1875,1899]" pageId="6" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.30 (E–H).
</paragraph>
</caption>
<paragraph id="C62A36E5FFC6FF96C6BCFF57FCCEFF31" blockId="7.[199,1233,150,250]" box="[199,888,150,177]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCFF57FEF7FF30" bold="true" box="[199,321,150,176]" pageId="7" pageNumber="122">Variation:</emphasis>
Tibia Iin another female (IZCAS Ar39905): 3.99.
</paragraph>
<paragraph id="C62A36E5FFC6FF96C6BCFF7AFB67FF55" blockId="7.[199,1233,150,250]" box="[199,1233,186,213]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCFF7AFE33FF54" bold="true" box="[199,389,186,213]" pageId="7" pageNumber="122">Natural history.</emphasis>
The species was found on domed webs attached to the underside of leaves.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC6FF96C6BCFF1FFC3AFF7A" box="[199,908,222,250]" pageId="7" pageNumber="122" type="distribution">
<paragraph id="C62A36E5FFC6FF96C6BCFF1FFC3AFF7A" blockId="7.[199,1233,150,250]" box="[199,908,222,250]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCFF1FFED7FF78" bold="true" box="[199,353,222,248]" pageId="7" pageNumber="122">Distribution.</emphasis>
<collectingCountry id="BE827675FFC6FF96C71CFF1EFE70FF79" box="[359,454,223,249]" name="Vietnam" pageId="7" pageNumber="122">Vietnam</collectingCountry>
(
<collectingRegion id="0451F807FFC6FF96C7AFFF1EFDF8FF79" box="[468,590,223,249]" country="Vietnam" name="Ninh Binh" pageId="7" pageNumber="122">Ninh Binh</collectingRegion>
,
<typeStatus id="192E8847FFC6FF96C422FF1EFD3FFF79" box="[601,649,223,249]" pageId="7" pageNumber="122">type</typeStatus>
locality only;
<figureCitation id="5EAE2A60FFC6FF96C557FF1EFCC9FF7A" box="[812,895,223,250]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="7" pageNumber="122">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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