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<mods:title id="FCD27A4A5E1F5BD3444C5468C8E893CF">Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae)</mods:title>
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<treatment id="485787BFFFB0C32DFF0BFC8FFBFDFAD1" ID-DOI="http://doi.org/10.5281/zenodo.5621738" ID-GBIF-Taxon="119417545" ID-Zenodo-Dep="5621738" LSID="urn:lsid:plazi:treatment:485787BFFFB0C32DFF0BFC8FFBFDFAD1" httpUri="http://treatment.plazi.org/id/485787BFFFB0C32DFF0BFC8FFBFDFAD1" lastPageId="15" lastPageNumber="52" pageId="13" pageNumber="50">
<subSubSection id="88E46522FFB0C32FFF0BFC8FFE32FCD7" pageId="13" pageNumber="50" type="nomenclature">
<paragraph id="C04136A9FFB0C32FFF0BFC8FFDE6FCF4" blockId="13.[151,609,835,894]" box="[151,609,835,862]" pageId="13" pageNumber="50">
<heading id="9B0981C5FFB0C32FFF0BFC8FFDE6FCF4" bold="true" box="[151,609,835,862]" fontSize="11" level="1" pageId="13" pageNumber="50" reason="1">
<taxonomicName id="07FE4D2AFFB0C32FFF0BFC8FFDE6FCF4" authority="Murray, 1884" authorityName="Murray" authorityYear="1884" box="[151,609,835,862]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="50" phylum="Chordata" rank="species" species="gleadowi">
<emphasis id="F28AEABBFFB0C32FFF0BFC8FFDE6FCF4" bold="true" box="[151,609,835,862]" pageId="13" pageNumber="50">
<emphasis id="F28AEABBFFB0C32FFF0BFC8FFE28FCF7" bold="true" box="[151,431,836,862]" italics="true" pageId="13" pageNumber="50">Hemidactylus gleadowi</emphasis>
Murray, 1884
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C04136A9FFB0C32FFF0BFCADFE32FCD7" blockId="13.[151,609,835,894]" box="[151,437,870,894]" pageId="13" pageNumber="50">
(
<figureCitation id="58C52A2CFFB0C32FFF02FCADFE81FCD7" box="[158,262,870,894]" captionStart="FIGURE 1" captionStartId="5.[151,250,1169,1192]" captionTargetBox="[151,1436,447,1147]" captionTargetId="figure@5.[151,1436,447,1147]" captionTargetPageId="5" captionText="FIGURE 1. Historical figures of the Hemidactylus brookii group taxa: A. H. brookii adult male lectotype (BMNH 1947.3.6.47) by present designation, from Gray (1867: Pl. 15, Fig. 2), later reprinted in Gray (1875: Pl. 15, Fig. 2); B. H. subtriedroides adult male paralectotype (ZSI 4135) by present designation, from Annandale (1905 b: Pl. II, Fig. 1); C. H. gleadowi not based on the original type series, from Boulenger (1890: 86, Fig. 27), later reprinted as H. brookii (Boulenger 1912: 42, Fig. 14; Smith 1935: 90, Fig. 28); D. H. gleadowi a syntype from Murray (1884 a: Fig. 3)." httpUri="https://zenodo.org/record/278833/files/figure.png" pageId="13" pageNumber="50">Figures 1</figureCitation>
D, 2B, 3C, 3D)
</paragraph>
</subSubSection>
<subSubSection id="88E46522FFB0C32FFF0BFC64FE23FBE4" pageId="13" pageNumber="50" type="reference_group">
<paragraph id="C04136A9FFB0C32FFF0BFC64FC62FC4A" blockId="13.[151,1387,942,995]" pageId="13" pageNumber="50">
<treatmentCitationGroup id="E0EE1187FFB0C32FFF0BFC64FC62FC4A" pageId="13" pageNumber="50">
<taxonomicName id="07FE4D2AFFB0C32FFF0BFC64FDD1FC6D" authority="Gleadowi Murray, 1884a:260" authorityName="Gleadowi Murray" authorityPageNumber="260" authorityYear="1884" box="[151,598,942,965]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="F28AEABBFFB0C32FFF0BFC64FE0CFC6C" box="[151,395,943,965]" italics="true" pageId="13" pageNumber="50">Hemidactylus Gleadowi</emphasis>
<treatmentCitation id="415F10B8FFB0C32FFE0DFC64FDD1FC6D" author="Murray" box="[401,598,942,964]" page="260" pageId="13" pageNumber="50" year="1884">
<bibRefCitation id="A46F4B58FFB0C32FFE0DFC64FDAEFC6D" author="Murray" box="[401,553,942,964]" pageId="13" pageNumber="50" refString="Murray, J. A. (1884 a) The Vertebrate Zoology of Sind. A Systematic Account with Descriptions of all the Known Species of Mammals, Birds, and Reptiles Inhabiting the Province; Observations on Their Habits, &amp; c; Tables of Their Geographic Distribution in Persia, Beloochistan, and Afghanistan; Punjab, North-West Provinces, and the Peninsula of India Generally, with Woodcuts, Lithographs, and Coloured Illustrations. Richardson &amp; Co., London. xvi + 424 pp. + Pl. I - XIII." type="book" year="1884" yearSuffix="a">Murray, 1884a</bibRefCitation>
:260
</treatmentCitation>
</taxonomicName>
. Original
<typeStatus id="1F45880BFFB0C32FFD26FC64FD61FC6D" box="[698,742,943,964]" pageId="13" pageNumber="50">type</typeStatus>
locality: “Rantah forests in Sind, (Jerruck Division)”,
<collectingCountry id="B8E97639FFB0C32FFA92FC65FAE0FC6D" box="[1294,1383,942,964]" name="Pakistan" pageId="13" pageNumber="50">Pakistan</collectingCountry>
.
<taxonomicName id="07FE4D2AFFB0C32FFF0BFC06FE01FC4A" box="[151,390,973,995]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="50" phylum="Chordata" rank="species" species="gleadovii">
<emphasis id="F28AEABBFFB0C32FFF0BFC06FE01FC4A" box="[151,390,973,995]" italics="true" pageId="13" pageNumber="50">Hemidactylus gleadovii</emphasis>
</taxonomicName>
<bibRefCitation id="A46F4B58FFB0C32FFE30FC06FDD0FC4A" author="Boulenger" box="[428,599,973,995]" pageId="13" pageNumber="50" refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Natural History). Second Ed.; Volume 1: Gekkonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. Taylor &amp; Francis, London. xii + 436 pp. + Pl. I - XXXII" type="book" year="1885">Boulenger, 1885</bibRefCitation>
:iiv, 116, 129. (Unjustified emendation)
</treatmentCitationGroup>
</paragraph>
<paragraph id="C04136A9FFB0C32FFF0BFBDBFE23FBE4" blockId="13.[151,1437,1039,2036]" pageId="13" pageNumber="50">
<emphasis id="F28AEABBFFB0C32FFF0BFBDBFD99FB81" bold="true" box="[151,542,1039,1064]" pageId="13" pageNumber="50">
<typeStatus id="1F45880BFFB0C32FFF0BFBDBFF7EFB81" box="[151,249,1040,1064]" pageId="13" pageNumber="50" type="neotype">Neotype</typeStatus>
by present designation.
</emphasis>
BMNH [18]84.7.25.8, adult male, “Sind” (= Sindh Province,
<collectingCountry id="B8E97639FFB0C32FFB69FBC4FADEFB8E" box="[1269,1369,1039,1063]" name="Pakistan" pageId="13" pageNumber="50">Pakistan</collectingCountry>
), presented by J. A. Murray.
</paragraph>
</subSubSection>
<subSubSection id="88E46522FFB0C32FFF59FB9CFB7DFBD9" box="[197,1274,1111,1136]" pageId="13" pageNumber="50" type="materials_examined">
<paragraph id="C04136A9FFB0C32FFF59FB9CFB7DFBD9" blockId="13.[151,1437,1039,2036]" box="[197,1274,1111,1136]" pageId="13" pageNumber="50">
<emphasis id="F28AEABBFFB0C32FFF59FB9CFE7DFBD9" bold="true" box="[197,506,1111,1136]" pageId="13" pageNumber="50">Other examined material.</emphasis>
<collectionCode id="A6EFAE6CFFB0C32FFD9CFB93FDD3FBC6" box="[512,596,1112,1135]" pageId="13" pageNumber="50">BMNH</collectionCode>
[18]84.7.25.9, topotype adult female, details as for
<typeStatus id="1F45880BFFB0C32FFB05FB93FB72FBD9" box="[1177,1269,1112,1136]" pageId="13" pageNumber="50" type="neotype">neotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="88E46522FFB0C32CFF59FBB7FD43FECD" lastPageId="14" lastPageNumber="51" pageId="13" pageNumber="50" type="etymology">
<paragraph id="C04136A9FFB0C32FFF59FBB7FCC7FB10" blockId="13.[151,1437,1039,2036]" pageId="13" pageNumber="50">
<emphasis id="F28AEABBFFB0C32FFF59FBB7FECCFB3D" bold="true" box="[197,331,1148,1172]" pageId="13" pageNumber="50">Etymology.</emphasis>
The specific epithet is a patronym after Mr. F. Gleadow, Deputy Conservator of Forests and collector of the original
<typeStatus id="1F45880BFFB0C32FFEFEFB6AFE15FB10" box="[354,402,1185,1209]" pageId="13" pageNumber="50">type</typeStatus>
specimens described by J. A. Murray.
</paragraph>
<paragraph id="C04136A9FFB0C32FFF59FB08FBD3F9C1" blockId="13.[151,1437,1039,2036]" pageId="13" pageNumber="50">
<emphasis id="F28AEABBFFB0C32FFF59FB08FEC3FB75" bold="true" box="[197,324,1219,1244]" pageId="13" pageNumber="50">Definition.</emphasis>
<taxonomicName id="07FE4D2AFFB0C32FFED3FB0FFDDCFB75" box="[335,603,1220,1244]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="50" phylum="Chordata" rank="species" species="gleadowi">
<emphasis id="F28AEABBFFB0C32FFED3FB0FFDDCFB75" box="[335,603,1220,1244]" italics="true" pageId="13" pageNumber="50">Hemidactylus gleadowi</emphasis>
</taxonomicName>
can be distinguished from all currently described South and Southeast Asian members of
<taxonomicName id="07FE4D2AFFB0C32FFEECFB23FD8CFAA9" box="[368,523,1256,1280]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="F28AEABBFFB0C32FFEECFB23FD8CFAA9" box="[368,523,1256,1280]" italics="true" pageId="13" pageNumber="50">Hemidactylus</emphasis>
</taxonomicName>
based on the following combination of characters: adult SVL to
<quantity id="07069B4CFFB0C32FFB68FB2CFADBFAA9" box="[1268,1372,1255,1280]" metricMagnitude="-2" metricUnit="m" metricValue="4.31" pageId="13" pageNumber="50" unit="mm" value="43.1">43.1 mm</quantity>
; TrL/ SVL 39.941.9%; HL/SVL 3030.2%; ear opening oval; primary postmental shield width is subequal to that of the first infralabial, secondary pair broadly in contact with second infralabials; tubercles of the parietal region are of subequal size to largest canthal scales; 1718 regular longitudinal rows of dorsal tubercles, largest 1214 times size of surrounding granules; two series of 1213 precloacal-femoral pores separated medially by a diastema of one non-pore-bearing scale, size subequal to pore-bearing scales, scale row bordering anteriorly the precloacal-femoral pore series of subequal size to pore-bearing scales; 45 lamellae under digit I and seven under digit IV of pes, subdigital lamellae mostly divided on digit IV of manus and pes, enlarged lamellae series under digit IV of pes absent on the basal 20% of digit length; tail oval in cross-section without lateral denticulation, TailD/TailW 71.784%, tubercles on proximal tail portion form short recurved conical spines, subcaudals completely transverse the tail width on the distal third of original tail; two very small, bluntly conical cloacal spurs.
</paragraph>
<paragraph id="C04136A9FFB0C32CFF59F9B8FA12FF5E" blockId="13.[151,1437,1039,2036]" lastBlockId="14.[151,1437,151,2013]" lastPageId="14" lastPageNumber="51" pageId="13" pageNumber="50">
<emphasis id="F28AEABBFFB0C32FFF59F9B8FEECF925" bold="true" box="[197,363,1651,1676]" pageId="13" pageNumber="50">Comparisons.</emphasis>
<taxonomicName id="07FE4D2AFFB0C32FFEEFF9BFFDFBF925" box="[371,636,1652,1676]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="50" phylum="Chordata" rank="species" species="gleadowi">
<emphasis id="F28AEABBFFB0C32FFEEFF9BFFDFBF925" box="[371,636,1652,1676]" italics="true" pageId="13" pageNumber="50">Hemidactylus gleadowi</emphasis>
</taxonomicName>
is here compared with the morphologically most similar Asian
<taxonomicName id="07FE4D2AFFB0C32FFACAF9BEFF73F919" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="50" phylum="Chordata" rank="genus">
<emphasis id="F28AEABBFFB0C32FFACAF9BEFF73F919" italics="true" pageId="13" pageNumber="50">Hemidactylus</emphasis>
</taxonomicName>
species occurring east of the western
<collectingCountry id="B8E97639FFB0C32FFD3FF953FC85F919" box="[675,770,1688,1712]" name="Pakistan" pageId="13" pageNumber="50">Pakistan</collectingCountry>
border. It differs from
<taxonomicName id="07FE4D2AFFB0C32FFB9BF952FB02F919" box="[1031,1157,1688,1712]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="50" phylum="Chordata" rank="species" species="treutleri">
<emphasis id="F28AEABBFFB0C32FFB9BF952FB02F919" box="[1031,1157,1688,1712]" italics="true" pageId="13" pageNumber="50">H. treutleri</emphasis>
</taxonomicName>
by its smaller size, SVL to
<quantity id="07069B4CFFB0C32FFF2FF977FE91F97D" box="[179,278,1724,1748]" metricMagnitude="-2" metricUnit="m" metricValue="4.31" pageId="13" pageNumber="50" unit="mm" value="43.1">43.1 mm</quantity>
[
<quantity id="07069B4CFFB0C32FFEBFF977FEF2F97D" box="[291,373,1724,1748]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="13" pageNumber="50" unit="mm" value="44.0">44 mm</quantity>
in original description] (vs. SVL to
<quantity id="07069B4CFFB0C32FFC95F977FCDBF97D" box="[777,860,1724,1748]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="50" unit="mm" value="70.0">70 mm</quantity>
), and total 2526 precloacal-femoral pores (vs. 14); from
<taxonomicName id="07FE4D2AFFB0C32FFF49F92AFE05F951" box="[213,386,1761,1784]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="50" phylum="Chordata" rank="species" species="gujaratensis">
<emphasis id="F28AEABBFFB0C32FFF49F92AFE05F951" box="[213,386,1761,1784]" italics="true" pageId="13" pageNumber="50">H. gujaratensis</emphasis>
</taxonomicName>
by its higher number of tubercle rows across the dorsum, 1718 (vs. 1214), and precloacalfemoral pores series separated medially by one non-pore-bearing scale (vs. at least five); from
<taxonomicName id="07FE4D2AFFB0C32FFB56F8CEFA1BF8B5" box="[1226,1436,1796,1820]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="50" phylum="Chordata" rank="species" species="parvimaculatus">
<emphasis id="F28AEABBFFB0C32FFB56F8CEFA1BF8B5" box="[1226,1436,1796,1820]" italics="true" pageId="13" pageNumber="50">H. parvimaculatus</emphasis>
</taxonomicName>
by precloacal-femoral pore series separated medially by one non-pore-bearing scale (vs. 24), and seven lamellae on digit IV of pes (vs. 810). From
<taxonomicName id="07FE4D2AFFB0C32FFDB3F886FD25F8CD" box="[559,674,1868,1892]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="50" phylum="Chordata" rank="species" species="brookii">
<emphasis id="F28AEABBFFB0C32FFDB3F886FD25F8CD" box="[559,674,1868,1892]" italics="true" pageId="13" pageNumber="50">H. brookii</emphasis>
</taxonomicName>
of which it was previously considered a synonym, by smaller size, SVL to
<quantity id="07069B4CFFB0C32FFF73F8BBFED4F821" box="[239,339,1904,1928]" metricMagnitude="-2" metricUnit="m" metricValue="4.31" pageId="13" pageNumber="50" unit="mm" value="43.1">43.1 mm</quantity>
[
<quantity id="07069B4CFFB0C32FFEFCF8BBFE32F821" box="[352,437,1904,1928]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="13" pageNumber="50" unit="mm" value="44.0">44 mm</quantity>
] (vs. SVL to
<quantity id="07069B4CFFB0C32FFDD6F8BBFD35F821" box="[586,690,1904,1928]" metricMagnitude="-2" metricUnit="m" metricValue="5.58" pageId="13" pageNumber="50" unit="mm" value="55.8">55.8 mm</quantity>
), largest dorsal tubercle size 1214 times that of surrounding granules (vs. 67 times), possessing a median non-pore-bearing scale of subequal size to pore-bearing scales (vs. a median non-pore-bearing scale &lt;50% the size of pored scales), proximal four to five segments of the original tail with transverse row of eight enlarged tubercles (vs. six), enlarged lamellae series under digit IV of pes absent on basal ~20% digit length (vs. lamellae series extend to the base of the digit), dorsal markings composed of one middorsal and two dorsolateral longitudinal rows of small darker brown spots and blotches widely separated from each other (vs. large dark longitudinally oval blotches in three rows, some merge to form incomplete transverse bands).
</paragraph>
<paragraph id="C04136A9FFB3C32CFF59FECEFA0CFEB5" blockId="14.[151,1437,151,2013]" box="[197,1419,260,284]" pageId="14" pageNumber="51">
For diagnosis from
<taxonomicName id="07FE4D2AFFB3C32CFE3DFECEFDDDFEB5" box="[417,602,260,284]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="51" phylum="Chordata" rank="species" species="kushmorensis">
<emphasis id="F28AEABBFFB3C32CFE3DFECEFDDDFEB5" box="[417,602,260,284]" italics="true" pageId="14" pageNumber="51">H. kushmorensis</emphasis>
</taxonomicName>
and
<taxonomicName id="07FE4D2AFFB3C32CFD0DFECEFC8CFEB5" box="[657,779,260,284]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="51" phylum="Chordata" rank="species" species="tenkatei">
<emphasis id="F28AEABBFFB3C32CFD0DFECEFC8CFEB5" box="[657,779,260,284]" italics="true" pageId="14" pageNumber="51">H. tenkatei</emphasis>
</taxonomicName>
see the respective comparison sections for these species.
</paragraph>
<paragraph id="C04136A9FFB3C32CFF59FEECFD43FECD" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="51">
<emphasis id="F28AEABBFFB3C32CFF59FEECFE25FEE9" bold="true" box="[197,418,295,320]" pageId="14" pageNumber="51">
Condition of
<typeStatus id="1F45880BFFB3C32CFEFBFEE3FE1BFEE9" box="[359,412,296,320]" pageId="14" pageNumber="51">type</typeStatus>
.
</emphasis>
<typeStatus id="1F45880BFFB3C32CFE37FEE3FD8DFEE9" box="[427,522,296,320]" pageId="14" pageNumber="51" type="neotype">Neotype</typeStatus>
is complete and undamaged with the exception of the posterior half of the tail which is detached, but present with the specimen.
</paragraph>
</subSubSection>
<subSubSection id="88E46522FFB3C32DFF59FEA4FE70FB41" lastPageId="15" lastPageNumber="52" pageId="14" pageNumber="51" type="description">
<paragraph id="C04136A9FFB3C32CFF59FEA4FBA1FB9D" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="51">
<emphasis id="F28AEABBFFB3C32CFF59FEA4FE5BFE21" bold="true" box="[197,476,367,392]" pageId="14" pageNumber="51">
Description of
<typeStatus id="1F45880BFFB3C32CFEEBFEBBFE50FE21" box="[375,471,368,392]" pageId="14" pageNumber="51" type="neotype">neotype</typeStatus>
.
</emphasis>
BMNH [18]84.7.25.8, adult male. A summary of mensural and meristic data is provided in table 1. A small sized species of
<taxonomicName id="07FE4D2AFFB3C32CFDFEFE5FFD7BFE05" box="[610,764,404,428]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="51" phylum="Chordata" rank="genus">
<emphasis id="F28AEABBFFB3C32CFDFEFE5FFD7BFE05" box="[610,764,404,428]" italics="true" pageId="14" pageNumber="51">Hemidactylus</emphasis>
</taxonomicName>
(SVL 43.0 mm); head distinct from neck, lores rounded and interorbital region flat, forehead flat; snout longer than orbit diameter; scales on snout circular, smallest are domed, largest on the canthal region are bluntly conical, size subequal to enlarged tubercles on the parietal, becoming mixed over the frontal with small granular scales; supraoculars primarily covered with homogenous small granular scales with a row of slightly enlarged, domed tubercles running parallel to the supraciliaries; dorsal and lateral surfaces of the head are covered with small granular scales densely mixed with enlarged domed tubercles, size increasing laterally and posteriorly; twelve interorbital scales across the narrowest point of the frontal; canthus rostralis rounded; pupil vertically elliptical, with crenellated edges; supraciliaries rounded, size increasing slightly anterodorsally, all lacking spines; ear opening deep, oval, obliquely orientated, lacking enlarged tubercles on anterior edge; orbit to ear distance slightly greater than diameter of orbits; rostral subrectangular, with medial groove dorsally, extending to half of the rostral depth; rostral depth more than half its width, contacted by nostrils, supralabial I, one internasal and two oval, slightly enlarged supranasals; nostrils circular, oriented dorsolaterally, nostril in contact with supralabial I, two postnasals, supranasal and rostral; three scale rows separating eye from supralabials; 9/9 (left/right) supralabials; 7/7 (left/right) infralabials; mental subtriangular, wider than its length (MenL/MenW 85.7%); two paired postmentals, primary pair subequally in contact with each other and the mental, secondary pair not in contact with each other, ~80% the size of the first and rounded posteriorly, primary postmental pair is bordered posteriorly by two enlarged smooth, rounded chin shields three times larger than adjacent granular scales; 2 3 rows of enlarged elongated scales border the lower edge of the infralabials, size increasing gradually anteriorly and laterally from the small throat granular scales; endolymphatic sac not visible.
</paragraph>
<paragraph id="C04136A9FFB3C32CFF59FB8BFD26FAD1" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="51">
Body slightly compressed dorsally, ventrolateral fold absent; dorsum covered with uniform, small granular scales interspersed with large tubercles, those of the nape are smallest, circular and domed to bluntly conical, size increasing posteriorly, dorsal most tubercles with a weak anterior median keel, laterally and posteriorly becoming more conical, all circular, largest are 1214 times the size of surrounding granular scales; 18 distinctly linear longitudinal rows at midbody,
<quantity id="07069B4CFFB3C32CFE28FB1BFE6AFB41" box="[436,493,1232,1256]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" pageId="14" pageNumber="51" unit="in" value="29.0">29 in</quantity>
a paravertebral line from the back of the skull to the area above the vent, intertubercle distance varies randomly; ventrolateral and gular granular scales grade suddenly into large, smooth, imbricate ventrals; preanal depression absent; precloacal-femoral pores number 13/12 (left/right), a single non-pore-bearing scale of subequal size to pored scales separates pore-bearing rows; precloacal scales between the pore series and the cloaca are not enlarged relative to ventrals.
</paragraph>
<paragraph id="C04136A9FFB3C32CFF59FA4FFEF6F881" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="51">Forelimbs slender; dorsal surface of the upper forelimb covered with smooth, slightly imbricate scales grading to small granular scales posteriorly and ventrally, dorsal surface of the lower forelimb covered with small granular scales of subequal size to those on the dorsum and intermixed with enlarged tubercles of subequal size to those on the nape, subimbricate scales of the upper forelimb extend anteriorly along the lower forelimb onto the dorsal surface of the manus; hind limbs relatively short; dorsal surface of the hind limbs and posterior thigh covered with small granular scales, dorsally widely interspersed with larger conical tubercles, largest being of equal size to the largest dorsal tubercles, ventral surfaces of hind limbs with flat imbricate to subimbricate scales; ventral surface of the manus and pes covered with smooth, rounded granular scales; digits relatively short, flattened, a small curved claw on all digital tips; distal phalanges elevated; lamellae numbering on left manus (total: divided) I (5: 2), II (6: 5), III (7: 5), IV (7: 4) and V (6: 3); and on left pes I (4: 2), II (7: 4), III (7: 5), IV (7: 5) and V (6: 4); basal subdigital lamellae narrow, enlarged lamellae series under digit IV of pes absent on basal ~20% digit length; interdigital webbing absent.</paragraph>
<paragraph id="C04136A9FFB3C32DFF59F8FFFC8EFE96" blockId="14.[151,1437,151,2013]" lastBlockId="15.[151,1437,151,1401]" lastPageId="15" lastPageNumber="52" pageId="14" pageNumber="51">Tail completely original, distal half is detached but present; dorsally compressed and oval in cross section, longer than snout to vent length; slightly constricted at the base and tapering gradually to a narrow tip; post-cloacal hemipenal bulge distinct; two very small, conical cloacal spurs on each side; longitudinal middorsal furrow present on the tail, lateral furrow absent; median subcaudal series begin on the third tail segment consisting of transversely enlarged, smooth, subimbricate scales, approximately 30% tail width on the fourth segment only reaching approximately 80% tail width from the distal quarter of the tail length, laterally bordered by large posteriorly rounded subimbricate scales, size rapidly decreasing laterally and dorsally; tail segments are distinct for at least the proximal two thirds, becoming less distinct distally, first four segments with a transverse row of eight tubercles, followed by rows of six, dorsal most tail tubercles are domed to conical, laterally tail tubercles are erect conical spines anteriorly becoming obliquely pointed conical spines distally.</paragraph>
<paragraph id="C04136A9FFB2C32DFF59FE87FBC6FD0E" blockId="15.[151,1437,151,1401]" pageId="15" pageNumber="52">Colouration in preservative: Dorsal surfaces of the head, body, limbs and tail primarily mid brown. Dorsally with one middorsal and two dorsolateral longitudinal rows of barely distinguishable darker brown spots and blotches; dorsal surface of the head also with brown blotches and spots; two parallel stripes extend from the posterior edge of the orbit, the lower extending to the lower edge of the ear opening, the upper stripe reaching to half way between the orbit and the upper border of the ear opening; a brown canthal stripe from the supranasal to the anterior border of the orbit; the rostral, supra and infralabials with a large brown blotch on each scale; the upper surface of the forelimb manus and pes with fine brown mottling, the hind limb with scattered brown blotches; anteriorly the tail appears faintly mottled and faintly banded distally; entire ventral surface of the head, body, limbs and tail appear plain light beige, however under magnification individual scales have varying numbers of minute dark brown specks, most notably on the tail; precloacal-femoral pores are orange-brown.</paragraph>
<paragraph id="C04136A9FFB2C32DFF59FD78FE26FC0D" blockId="15.[151,1437,151,1401]" pageId="15" pageNumber="52">
<emphasis id="F28AEABBFFB2C32DFF59FD78FEBEFD65" bold="true" box="[197,313,691,716]" pageId="15" pageNumber="52">Variation.</emphasis>
Variation of major mensural and meristic characters are presented in table 1. Further difference from the
<typeStatus id="1F45880BFFB2C32DFE9CFD1CFEE1FD46" box="[256,358,727,751]" pageId="15" pageNumber="52" type="lectotype">lectotype</typeStatus>
include: contacting edges of first postmentals with each other are slightly longer than edges in contact with the mental, enlarged chin shields contacting the posterior edge of the first postmentals are replaced by slightly enlarged granular scales, secondary postmental pair in contact with first infralabials; first and third to fifth tail segments with a transverse row of eight tubercles, second segment with a row of nine and distal rows with six tubercles, lateral tail tubercles are not as pointed and more acutely angled posteriorly; no precloacal-femoral porebearing or pitted scales.
</paragraph>
<paragraph id="C04136A9FFB2C32DFF59FC64FE70FB41" blockId="15.[151,1437,151,1401]" pageId="15" pageNumber="52">
The original description is based on an unknown number of
<typeStatus id="1F45880BFFB2C32DFCE7FC7AFC5AFC60" box="[891,989,945,969]" pageId="15" pageNumber="52" type="syntype">syntypes</typeStatus>
and characters which vary from specimens examined here are as follows: 13 precloacal-femoral pores on each thigh; 1012 supralabials and 810 infralabials; 1516 tubercle rows at midbody; a transverse row of 810 tubercles on the first segment of the tail; and 3839 ventral scale rows. For colouration, it was not mentioned if the description provided was for live or preserved specimens (
<bibRefCitation id="A46F4B58FFB2C32DFEF6FB8BFD8AFBF1" author="Murray" box="[362,525,1088,1112]" pageId="15" pageNumber="52" refString="Murray, J. A. (1884 a) The Vertebrate Zoology of Sind. A Systematic Account with Descriptions of all the Known Species of Mammals, Birds, and Reptiles Inhabiting the Province; Observations on Their Habits, &amp; c; Tables of Their Geographic Distribution in Persia, Beloochistan, and Afghanistan; Punjab, North-West Provinces, and the Peninsula of India Generally, with Woodcuts, Lithographs, and Coloured Illustrations. Richardson &amp; Co., London. xvi + 424 pp. + Pl. I - XIII." type="book" year="1884" yearSuffix="a">Murray 1884a</bibRefCitation>
) and so is provided here “Colour pinkish grey, a greyish-white line from the nostril to the orbit, bordered by a dark line above and below; three dark lines radiating from behind the orbits, the uppermost curving behind and nearly meeting on the occiput, the next or central one extends from above the ear opening to the shoulder, and the third or lowest to below the ear opening. Back with 5 imperfect transverse bands. Tail with from 12 to 15 bands.”
</paragraph>
</subSubSection>
<subSubSection id="88E46522FFB2C32DFF59FB38FBFDFAD1" pageId="15" pageNumber="52" type="distribution">
<paragraph id="C04136A9FFB2C32DFF59FB38FBFDFAD1" blockId="15.[151,1437,151,1401]" pageId="15" pageNumber="52">
<emphasis id="F28AEABBFFB2C32DFF59FB38FED9FAA5" bold="true" box="[197,350,1267,1292]" pageId="15" pageNumber="52">Distribution.</emphasis>
This species is currently known only with certainty from the original
<typeStatus id="1F45880BFFB2C32DFB15FB3EFB3EFAA4" box="[1161,1209,1269,1293]" pageId="15" pageNumber="52">type</typeStatus>
locality as “Rantah forests in Sind, (Jerruck Division)”. Jerruck (
<geoCoordinate id="A5CA506EFFB2C32DFD3FFAD3FC94FA99" box="[675,787,1304,1328]" direction="north" orientation="latitude" pageId="15" pageNumber="52" precision="15" value="25.05">25°30”N</geoCoordinate>
,
<geoCoordinate id="A5CA506EFFB2C32DFCBEFAD3FC1CFA99" box="[802,923,1304,1328]" direction="east" orientation="longitude" pageId="15" pageNumber="52" precision="15" value="68.25">68°150”E</geoCoordinate>
) is currently situated in Hyderabad District, Sindh Province in southeastern
<collectingCountry id="B8E97639FFB2C32DFD99FAF7FDEFFAFD" box="[517,616,1340,1364]" name="Pakistan" pageId="15" pageNumber="52">Pakistan</collectingCountry>
. They were collected under the bark of Babool (Acacia) trees (
<bibRefCitation id="A46F4B58FFB2C32DFADAFAF7FF67FAD1" author="Murray" pageId="15" pageNumber="52" refString="Murray, J. A. (1884 a) The Vertebrate Zoology of Sind. A Systematic Account with Descriptions of all the Known Species of Mammals, Birds, and Reptiles Inhabiting the Province; Observations on Their Habits, &amp; c; Tables of Their Geographic Distribution in Persia, Beloochistan, and Afghanistan; Punjab, North-West Provinces, and the Peninsula of India Generally, with Woodcuts, Lithographs, and Coloured Illustrations. Richardson &amp; Co., London. xvi + 424 pp. + Pl. I - XIII." type="book" year="1884" yearSuffix="a">Murray 1884a</bibRefCitation>
). The considerably less informative locality associated with the
<typeStatus id="1F45880BFFB2C32DFC31FAAAFB81FAD0" box="[941,1030,1377,1401]" pageId="15" pageNumber="52" type="neotype">neotype</typeStatus>
is “Sind”.
</paragraph>
</subSubSection>
</treatment>
</document>