807 lines
123 KiB
XML
807 lines
123 KiB
XML
<document ID-DOI="http://dx.doi.org/10.5852/ejt.2013.37" ID-GBIF-Dataset="9682b45b-5e22-4965-a293-e203660e0ed3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3806770" ID-ZooBank="ADF6394E-77B5-4309-87FE-4097FDE0A3FD" checkinTime="1588796067459" checkinUser="carolina" docAuthor="Rowson, Ben & Tattersfield, Peter" docDate="2013" docId="4511E41DD83BFFE6FE7BFCBAF724F80C" docLanguage="en" docName="ejt-2013-37.pdf.imf" docOrigin="European Journal of Taxonomy 37" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article" docStyleVersion="14" docTitle="Dadagulella conoidea Rowson & Tattersfield 2013, comb. nov." docType="key" docVersion="8" lastPageNumber="34" masterDocId="B9289C65D824FFC7FFA4FF80F253FFE7" masterDocTitle="Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies" masterLastPageNumber="46" masterPageNumber="1" pageNumber="32" updateTime="1683580915233" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rowson, Ben</mods:namePart>
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<mods:nameIdentifier type="LSID">05D46259-DA45-45A4-9127-5AF361D566A7</mods:nameIdentifier>
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<mods:affiliation>Biodiversity & Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk & urn: lsid: zoobank. org: author: 05 D 46259 - DA 45 - 45 A 4 - 9127 - 5 AF 361 D 566 A 7</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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<mods:namePart>Tattersfield, Peter</mods:namePart>
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<mods:nameIdentifier type="LSID">8C55642E-B18A-461B-9BF9-31CB6DB7A070</mods:nameIdentifier>
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<mods:affiliation>Biodiversity & Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk & urn: lsid: zoobank. org: author: 8 C 55642 E-B 18 A- 461 B- 9 BF 9 - 31 CB 6 DB 7 A 070</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-02-04</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>37</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>46</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.5852/ejt.2013.37</mods:identifier>
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<mods:identifier type="GBIF-Dataset">9682b45b-5e22-4965-a293-e203660e0ed3</mods:identifier>
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<mods:identifier type="ISSN">2118-9773</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3806770</mods:identifier>
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<mods:identifier type="ZooBank">ADF6394E-77B5-4309-87FE-4097FDE0A3FD</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3815777" ID-GBIF-Taxon="163971585" ID-Zenodo-Dep="3815777" LSID="urn:lsid:plazi:treatment:4511E41DD83BFFE6FE7BFCBAF724F80C" httpUri="http://treatment.plazi.org/id/4511E41DD83BFFE6FE7BFCBAF724F80C" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
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<subSubSection box="[479,1109,826,853]" pageId="31" pageNumber="32" type="nomenclature">
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||
<paragraph blockId="31.[479,1109,826,890]" box="[479,1109,826,853]" pageId="31" pageNumber="32">
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<heading box="[479,1109,826,853]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="32" reason="2">
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<taxonomicName ID-CoL="33XQB" authority="(Verdcourt, 1996)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[479,972,826,853]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
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<emphasis bold="true" box="[479,972,826,853]" pageId="31" pageNumber="32">
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<emphasis bold="true" box="[479,744,826,853]" italics="true" pageId="31" pageNumber="32">Dadagulella conoidea</emphasis>
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(
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<bibRefCitation author="Verdcourt B." box="[759,963,826,852]" pageId="31" pageNumber="32" pagination="135 - 137" refId="ref28065" refString="Verdcourt B. 1996. A new species of Gulella Pfr. (Gastropoda Pulmonata, Streptaxidae) from NE. Tanzania. Basteria 59 (4 - 6): 135 - 137." type="journal article" year="1996">Verdcourt, 1996</bibRefCitation>
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||
)
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</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[979,1109,826,853]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[677,910,864,890]" pageId="31" pageNumber="32" type="description">
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||
<paragraph blockId="31.[479,1109,826,890]" box="[677,910,864,890]" pageId="31" pageNumber="32">
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<figureCitation box="[677,767,864,890]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="31" pageNumber="32">Figs 42</figureCitation>
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,
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<figureCitation box="[779,812,864,890]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="31" pageNumber="32">84</figureCitation>
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;
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<tableCitation box="[823,910,864,890]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="31" pageNumber="32" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="32" type="reference_group">
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<paragraph blockId="31.[189,783,932,959]" box="[189,783,932,959]" pageId="31" pageNumber="32">
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||
<treatmentCitationGroup box="[189,783,932,959]" pageId="31" pageNumber="32">
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<treatmentCitation author="Verdcourt B." box="[189,703,932,959]" page="137" pageId="31" pageNumber="32" year="1996">
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||
<taxonomicName ID-CoL="3HLPB" authority="Verdcourt, 1996: 135 - 137" authorityName="Verdcourt" authorityPageNumber="135 - 137" authorityYear="1996" box="[189,703,932,959]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="conoidea">
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<emphasis box="[189,392,932,958]" italics="true" pageId="31" pageNumber="32">Gulella conoidea</emphasis>
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<bibRefCitation author="Verdcourt B." box="[399,703,932,959]" pageId="31" pageNumber="32" pagination="135 - 137" refId="ref28065" refString="Verdcourt B. 1996. A new species of Gulella Pfr. (Gastropoda Pulmonata, Streptaxidae) from NE. Tanzania. Basteria 59 (4 - 6): 135 - 137." type="journal article" year="1996">Verdcourt, 1996: 135-137</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, fig. 1.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="31.[189,963,1000,1026]" box="[189,963,1000,1026]" pageId="31" pageNumber="32">
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<treatmentCitationGroup box="[189,963,1000,1026]" pageId="31" pageNumber="32">
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||
<emphasis box="[189,414,1000,1026]" italics="true" pageId="31" pageNumber="32">
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||
<taxonomicName ID-CoL="3HLPB" authorityName="Verdcourt" authorityYear="1996" box="[189,392,1000,1026]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="conoidea">Gulella conoidea</taxonomicName>
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–
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</emphasis>
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<treatmentCitation author="Tattersfield P." box="[421,682,1000,1026]" page="37" pageId="31" pageNumber="32" year="1998">
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<bibRefCitation author="Tattersfield P." box="[421,682,1000,1026]" pageId="31" pageNumber="32" pagination="31 - 41" refId="ref27614" refString="Tattersfield P. 1998 b. Three new species and a new subgenus of Gulella (Gastropoda: Streptaxidae) from Tanzania. Journal of Conchology 36 (2): 31 - 41." type="journal article" year="1998">Tattersfield 1998b: 37</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation author="Verdcourt B." box="[729,959,1000,1026]" page="48" pageId="31" pageNumber="32" year="2006">
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<bibRefCitation author="Verdcourt B." box="[729,959,1000,1026]" pageId="31" pageNumber="32" refId="ref28139" refString="Verdcourt B. 2006. A Revised List of the Non-marine Mollusca of East Africa (Kenya, Tanzania and Uganda, excluding Lake Malawi). Verdcourt B., Maidenhead, UK." type="book" year="2006">Verdcourt 2006: 48</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="32" type="materials_examined">
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<paragraph blockId="31.[189,492,1068,1094]" box="[189,492,1068,1094]" pageId="31" pageNumber="32">
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||
<heading bold="true" box="[189,492,1068,1094]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
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||
<emphasis bold="true" box="[189,492,1068,1094]" pageId="31" pageNumber="32">
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<typeStatus box="[189,250,1068,1094]" pageId="31" pageNumber="32">Type</typeStatus>
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material examined
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="31.[189,1398,1117,1219]" pageId="31" pageNumber="32">
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<materialsCitation ID-GBIF-Occurrence="2610647313" collectionCode="RMNH" collectorName="Frontier Tanzania" country="Tanzania" latitude="-4.58" location="Kwamgumi / Segoma Reserve" longLatPrecision="783" longitude="38.45" municipality="East Usambara" pageId="31" pageNumber="32" specimenCode="RMNH.MOL.57150" specimenCount="1" specimenCount-adult="1" typeStatus="holotype">
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<collectingCountry box="[189,343,1117,1143]" name="Tanzania" pageId="31" pageNumber="32">TANZANIA</collectingCountry>
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:
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<typeStatus box="[361,470,1117,1143]" pageId="31" pageNumber="32">Holotype</typeStatus>
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<specimenCode box="[486,732,1117,1144]" pageId="31" pageNumber="32">RMNH.MOL.57150</specimenCode>
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:
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<specimenCount box="[750,817,1117,1143]" pageId="31" pageNumber="32" type="adult">1 ad.</specimenCount>
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,
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<collectingMunicipality box="[837,1022,1117,1144]" pageId="31" pageNumber="32">East Usambara</collectingMunicipality>
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foothills,
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<location LSID="urn:lsid:plazi:treatment:4511E41DD83BFFE6FE7BFCBAF724F80C:C86703D0D83BFFD8FB21FBDDF34DFB7A" country="Tanzania" latitude="-4.58" longLatPrecision="783" longitude="38.45" municipality="East Usambara" name="Kwamgumi / Segoma Reserve" pageId="31" pageNumber="32">Kwamgumi/Segoma Reserve</location>
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, about 4.57°-
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<geoCoordinate box="[453,534,1155,1182]" degrees="4.58" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="555" value="-4.58">4.58°S</geoCoordinate>
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, 38.43°
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<emphasis box="[627,637,1156,1181]" italics="true" pageId="31" pageNumber="32">-</emphasis>
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<geoCoordinate box="[637,735,1155,1182]" degrees="38.45" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="555" value="38.45">38.45°E</geoCoordinate>
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, near Muzi R./Sigi R. junction, leg.
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<collectorName box="[1188,1398,1155,1181]" pageId="31" pageNumber="32">Frontier Tanzania</collectorName>
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(examined digital photograph only).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="31.[189,505,1260,1286]" box="[189,505,1260,1286]" pageId="31" pageNumber="32">
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<heading bold="true" box="[189,505,1260,1286]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
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<emphasis bold="true" box="[189,505,1260,1286]" pageId="31" pageNumber="32">Other material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="31.[189,260,1310,1336]" box="[189,260,1310,1336]" pageId="31" pageNumber="32">None.</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="32" type="description">
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<paragraph blockId="31.[189,334,1377,1403]" box="[189,334,1377,1403]" pageId="31" pageNumber="32">
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||
<heading bold="true" box="[189,334,1377,1403]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
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<emphasis bold="true" box="[189,334,1377,1403]" pageId="31" pageNumber="32">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="31.[189,1399,1426,1755]" pageId="31" pageNumber="32">
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SHELL (
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<figureCitation box="[273,360,1427,1453]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="31" pageNumber="32">Fig. 42</figureCitation>
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). Large (
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<quantity box="[460,550,1426,1453]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="31" pageNumber="32" unit="mm" value="5.5">5.5 mm</quantity>
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high x
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<quantity box="[635,738,1427,1453]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="31" pageNumber="32" unit="mm" value="2.8">2.80 mm</quantity>
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wide), of 8.0 whorls. Subconical (maximum width being in bottom third of the shell, at body whorl). Spire narrowly acuminate, almost cyrtoconoid (convex) rather than coeloconoid (spire angle 48°). Apex sharply pointed. Embryonic whorls “probably smooth but worn” (
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||
<bibRefCitation author="Verdcourt B." box="[325,515,1540,1567]" pageId="31" pageNumber="32" pagination="135 - 137" refId="ref28065" refString="Verdcourt B. 1996. A new species of Gulella Pfr. (Gastropoda Pulmonata, Streptaxidae) from NE. Tanzania. Basteria 59 (4 - 6): 135 - 137." type="journal article" year="1996">Verdcourt 1996</bibRefCitation>
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||
). Later whorls with relatively weak ribs (about 11 per mm on penultimate whorl). Sutures relatively shallow. Umbilicus narrowly open. Peristome incomplete parietally, or nearly so. Dentition 3-fold (although could be interpreted as 4-fold), consisting of: one lamella-like parietal tooth, with a swelling above it that recalls that in
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<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[781,946,1653,1679]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
|
||
<emphasis box="[781,946,1653,1679]" italics="true" pageId="31" pageNumber="32">D. rondoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[955,1089,1653,1680]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
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||
; one strongly bifid palatal tooth (or pair of teeth), not forming a parieto-palatal sinus; and a shallow columellar tooth or denticle. Shells and anatomy of juveniles unknown.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="31" pageNumber="32" type="multiple">
|
||
<paragraph blockId="31.[189,421,1797,1823]" box="[189,421,1797,1823]" pageId="31" pageNumber="32">
|
||
<heading bold="true" box="[189,421,1797,1823]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
|
||
<emphasis bold="true" box="[189,421,1797,1823]" pageId="31" pageNumber="32">Range and habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="31.[189,1398,1846,1910]" pageId="31" pageNumber="32">
|
||
Altitude and habitat not stated but probably in lowland forest at approximately
|
||
<quantity box="[1101,1174,1846,1872]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="32" unit="m" value="200.0">200 m</quantity>
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||
at the
|
||
<typeStatus box="[1249,1300,1846,1872]" pageId="31" pageNumber="32">type</typeStatus>
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||
locality, northeastern
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||
<collectingCountry box="[341,450,1884,1910]" name="Tanzania" pageId="31" pageNumber="32">Tanzania</collectingCountry>
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||
. Another species (
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||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[664,775,1884,1910]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
|
||
<emphasis box="[664,775,1884,1910]" italics="true" pageId="31" pageNumber="32">D. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[783,914,1884,1910]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
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||
) also occurs at Kwamgumi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[189,624,1952,2027]" pageId="31" pageNumber="32" type="discussion">
|
||
<paragraph blockId="31.[189,303,1952,1978]" box="[189,303,1952,1978]" pageId="31" pageNumber="32">
|
||
<heading bold="true" box="[189,303,1952,1978]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
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||
<emphasis bold="true" box="[189,303,1952,1978]" pageId="31" pageNumber="32">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="31.[189,624,2001,2027]" box="[189,624,2001,2027]" pageId="31" pageNumber="32">
|
||
See
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[239,403,2001,2027]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
|
||
<emphasis box="[239,403,2001,2027]" italics="true" pageId="31" pageNumber="32">D. rondoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[410,539,2001,2027]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
|
||
above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSection lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="key">
|
||
<subSubSection box="[189,822,267,299]" pageId="32" pageNumber="33" type="nomenclature">
|
||
<paragraph blockId="32.[189,822,267,299]" box="[189,822,267,299]" pageId="32" pageNumber="33">
|
||
<heading bold="true" box="[189,822,267,299]" fontSize="13" level="1" pageId="32" pageNumber="33" reason="1">
|
||
<emphasis bold="true" box="[189,822,267,299]" pageId="32" pageNumber="33">Key to species and subspecies (adult shells)</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="key">
|
||
<keyStep pageId="32" pageNumber="33">
|
||
<paragraph blockId="32.[189,1399,322,423]" pageId="32" pageNumber="33">
|
||
<key pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
1. Dentition simple, 2-fold to 4-fold (
|
||
<figureCitation box="[636,726,322,349]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="32" pageNumber="33">Figs 35</figureCitation>
|
||
,
|
||
<figureCitation box="[737,769,323,349]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="32" pageNumber="33">38</figureCitation>
|
||
,
|
||
<figureCitation box="[780,855,323,349]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="32" pageNumber="33">41, 42</figureCitation>
|
||
,
|
||
<figureCitation box="[866,984,322,349]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="32" pageNumber="33">51, 56, 62</figureCitation>
|
||
), never including a parietal denticle to the left of parietal tooth ............................................................................................................... 2
|
||
</keyLead>
|
||
</key>
|
||
</paragraph>
|
||
<key pageId="32" pageNumber="33">
|
||
<paragraph blockId="32.[189,1399,322,423]" box="[193,1399,396,423]" pageId="32" pageNumber="33">
|
||
<keyLead box="[193,1399,396,423]" pageId="32" pageNumber="33">
|
||
– Dentition complex, 5-fold to 8-fold (
|
||
<figureCitation box="[665,965,396,423]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="32" pageNumber="33">Figs 52-55, 57-61, 63, 64</figureCitation>
|
||
) (if in doubt key from here) ......... 8
|
||
</keyLead>
|
||
</paragraph>
|
||
<keyStep pageId="32" pageNumber="33">
|
||
<paragraph blockId="32.[189,1399,464,565]" box="[189,1399,464,490]" pageId="32" pageNumber="33">
|
||
<keyLead box="[189,1399,464,490]" pageId="32" pageNumber="33">2. Basal denticle absent ........................................................................................................................ 3</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="32.[189,1399,464,565]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
<key pageId="32" pageNumber="33">
|
||
– Basal denticle present (may be partly or wholly hidden behind palatal tooth, e.g.
|
||
<figureCitation box="[1237,1327,501,528]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="32" pageNumber="33">Fig. 56</figureCitation>
|
||
, so if alternative fails return here) ............................................................................................................. 6
|
||
</key>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</keyStep>
|
||
<keyStep pageId="32" pageNumber="33">
|
||
<paragraph blockId="32.[189,1399,605,743]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
<key pageId="32" pageNumber="33">
|
||
3. Shell small (up to
|
||
<quantity box="[445,534,605,632]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="32" pageNumber="33" unit="mm" value="3.1">3.1 mm</quantity>
|
||
high) with very fine, very numerous ribs (about 27 per mm on penultimate whorl) (
|
||
<figureCitation box="[329,413,642,668]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="32" pageNumber="33">Fig. 37</figureCitation>
|
||
) ......................................................
|
||
<taxonomicName authority="(van Bruggen, 2000)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="2000" box="[840,1262,642,669]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="meredithae" status="comb. nov.">
|
||
<emphasis bold="true" box="[840,1011,642,669]" italics="true" pageId="32" pageNumber="33">D. meredithae</emphasis>
|
||
(van
|
||
<bibRefCitation author="Bruggen A. C. van" box="[1078,1253,642,669]" pageId="32" pageNumber="33" pagination="225 - 235" refId="ref26131" refString="Bruggen A. C. van. 2000. Studies on the Streptaxidae (Mollusca: Gastropoda Pulmonata) of Malawi. 5. Description of Gulella meredithae spec. nov. Zoologische Mededelingen 74 (15): 225 - 235." type="journal article" year="2000">Bruggen, 2000</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1269,1399,642,669]" pageId="32" pageNumber="33" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</key>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="32.[189,1399,605,743]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
<key pageId="32" pageNumber="33">
|
||
– Shell large (
|
||
<quantity box="[376,513,679,706]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.5" metricValueMin="4.1" pageId="32" pageNumber="33" unit="mm" value="4.8" valueMax="5.5" valueMin="4.1">
|
||
4.1
|
||
<emphasis box="[414,424,680,705]" italics="true" pageId="32" pageNumber="33">-</emphasis>
|
||
5.5 mm
|
||
</quantity>
|
||
high) and with 13 or fewer ribs per mm on penultimate whorl (
|
||
<figureCitation box="[1256,1347,679,706]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="32" pageNumber="33">Figs 35</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="32" pageNumber="33">41- 42</figureCitation>
|
||
) .................................................................................................................................................... 4
|
||
</key>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<key lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
|
||
<keyStep pageId="32" pageNumber="33">
|
||
<paragraph blockId="32.[189,1399,783,921]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
4. Shell ovate-acuminate (maximum width approximately at middle of shell); spire broadly acuminate (spire angle 56° or more); ribs sinuous (
|
||
<figureCitation box="[702,787,820,847]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="32" pageNumber="33">Fig. 35</figureCitation>
|
||
) .....................................................
|
||
<taxonomicName authority="Rowson & Tattersfield, 2013" authorityName="Rowson & Tattersfield" authorityYear="2013" box="[1206,1301,820,847]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="delta" status="sp. nov.">
|
||
<emphasis bold="true" box="[1206,1301,820,847]" italics="true" pageId="32" pageNumber="33">D. delta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1308,1399,821,847]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="32.[189,1399,783,921]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
– Shell subconical (maximum width in bottom third of shell); spire narrowly acuminate (spire angle 52° or less); ribs not sinuous (
|
||
<figureCitation box="[585,714,894,920]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="32" pageNumber="33">Figs 41-42</figureCitation>
|
||
) ....................................................................................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="32" pageNumber="33">
|
||
<paragraph blockId="32.[189,1399,961,1099]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
5. One palatal tooth, columellar dentition weak or absent;
|
||
<quantity box="[887,992,962,988]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="32" pageNumber="33" unit="mm" value="4.1">4.10 mm</quantity>
|
||
high (
|
||
<figureCitation box="[1067,1152,961,988]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="32" pageNumber="33">Fig. 41</figureCitation>
|
||
) .............................. ......................................................................................
|
||
<taxonomicName authority="(Verdcourt, 1994)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[878,1262,998,1025]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
|
||
<emphasis bold="true" box="[878,1046,998,1025]" italics="true" pageId="32" pageNumber="33">D. rondoensis</emphasis>
|
||
(
|
||
<bibRefCitation author="Verdcourt B." box="[1061,1253,998,1025]" pageId="32" pageNumber="33" pagination="137 - 139" refId="ref28031" refString="Verdcourt B. 1994. A new species of Gulella from SE. Tanzania (Pulmonata: Streptaxidae). Archiv fur Molluskenkunde 123 (1 / 6): 137 - 139." type="journal article" year="1994">Verdcourt, 1994</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1269,1399,998,1025]" pageId="32" pageNumber="33" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="32.[189,1399,961,1099]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
– Two palatal teeth; one strong shallow columellar tooth;
|
||
<quantity box="[889,979,1035,1062]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="32" pageNumber="33" unit="mm" value="5.5">5.5 mm</quantity>
|
||
high (
|
||
<figureCitation box="[1054,1140,1035,1062]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="32" pageNumber="33">Fig. 42</figureCitation>
|
||
) ................................ .........................................................................................
|
||
<taxonomicName authority="(Verdcourt, 1996)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[901,1262,1072,1099]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
|
||
<emphasis bold="true" box="[901,1046,1072,1099]" italics="true" pageId="32" pageNumber="33">D. conoidea</emphasis>
|
||
(
|
||
<bibRefCitation author="Verdcourt B." box="[1061,1253,1072,1099]" pageId="32" pageNumber="33" pagination="135 - 137" refId="ref28065" refString="Verdcourt B. 1996. A new species of Gulella Pfr. (Gastropoda Pulmonata, Streptaxidae) from NE. Tanzania. Basteria 59 (4 - 6): 135 - 137." type="journal article" year="1996">Verdcourt, 1996</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1269,1399,1072,1099]" pageId="32" pageNumber="33" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="32" pageNumber="33">
|
||
<paragraph blockId="32.[189,1399,1140,1277]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
6. Deep-set columellar baffle clearly visible through aperture (if not so clearly visible, then apex pointed and not rounded) (e.g.
|
||
<figureCitation box="[597,808,1177,1204]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="32" pageNumber="33">Figs 51-57, 62-64</figureCitation>
|
||
) .......................................................................... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="32.[189,1399,1140,1277]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
– Columellar baffle barely visible through aperture; apex rounded (
|
||
<collectingRegion box="[998,1058,1214,1240]" country="Kenya" name="Taita/Taveta" pageId="32" pageNumber="33">Taita</collectingRegion>
|
||
Hills,
|
||
<collectingCountry box="[1137,1215,1214,1240]" name="Kenya" pageId="32" pageNumber="33">Kenya</collectingCountry>
|
||
) (
|
||
<figureCitation box="[1240,1325,1214,1240]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="32" pageNumber="33">Fig. 33</figureCitation>
|
||
) ...... .............................................................................
|
||
<taxonomicName authority="(Rowson & Lange, 2007)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Rowson & Lange" baseAuthorityYear="2007" box="[815,1262,1251,1277]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="nictitans" status="comb. nov.">
|
||
<emphasis bold="true" box="[815,954,1251,1277]" italics="true" pageId="32" pageNumber="33">D. nictitans</emphasis>
|
||
(
|
||
<bibRefCitation author="Rowson B. & Lange C. N." box="[970,1253,1251,1277]" pageId="32" pageNumber="33" pagination="21 - 32" refId="ref27291" refString="Rowson B. & Lange C. N. 2007. Two new species of Gulella (Mollusca: Pulmonata: Streptaxidae) from the Taita Hills, Kenya. African Invertebrates 48 (2): 21 - 32." type="journal article" year="2007">Rowson & Lange, 2007</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1269,1399,1251,1277]" pageId="32" pageNumber="33" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="32" pageNumber="33">
|
||
<paragraph blockId="32.[189,1399,1318,1493]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
7. No columellar dentition other than deep-set baffle; basal denticle present, either completely hidden by a slab-like palatal tooth or visible only when shell is turned (Kimboza, Uluguru Mts,
|
||
<collectingCountry box="[1283,1391,1355,1382]" name="Tanzania" pageId="32" pageNumber="33">Tanzania</collectingCountry>
|
||
) (
|
||
<figureCitation box="[243,334,1392,1418]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="32" pageNumber="33">Figs 28</figureCitation>
|
||
,
|
||
<figureCitation box="[346,377,1392,1419]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="32" pageNumber="33">56</figureCitation>
|
||
) .....................................................................................................
|
||
<taxonomicName authority="Rowson & Tattersfield, 2013" authorityName="Rowson & Tattersfield" authorityYear="2013" box="[1148,1301,1392,1419]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="ecclesiola" status="sp. nov.">
|
||
<emphasis bold="true" box="[1148,1301,1392,1419]" italics="true" pageId="32" pageNumber="33">D. ecclesiola</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1308,1399,1393,1419]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="32.[189,1399,1318,1493]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">– Usually with shallow columellar dentition as well as deep-set baffle; basal denticle clearly visible in normal apertural view .................................................................................................................... 19</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="32" pageNumber="33">
|
||
<paragraph blockId="32.[189,1399,1533,1634]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
8. Parietal tooth strongly V-shaped, flaring, directed outwards; three palatal teeth; shell of 5.0 or fewer whorls (
|
||
<figureCitation box="[330,421,1570,1596]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="32" pageNumber="33">Figs 34</figureCitation>
|
||
,
|
||
<figureCitation box="[433,463,1570,1597]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="32" pageNumber="33">61</figureCitation>
|
||
) ....................................................................................
|
||
<taxonomicName authority="Rowson & Tattersfield, 2013" authorityName="Rowson & Tattersfield" authorityYear="2013" box="[1107,1301,1570,1597]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="frontierarum" status="sp. nov.">
|
||
<emphasis bold="true" box="[1107,1301,1570,1597]" italics="true" pageId="32" pageNumber="33">D. frontierarum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1308,1399,1571,1597]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="32.[189,1399,1533,1634]" box="[193,1399,1607,1634]" pageId="32" pageNumber="33">
|
||
<keyLead box="[193,1399,1607,1634]" pageId="32" pageNumber="33">– Shell without this combination of characters ................................................................................... 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="32" pageNumber="33">
|
||
<paragraph blockId="32.[189,1398,1674,1812]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
9. Upper shallow columellar tooth strong, squarish, projecting into aperture as far or further than any deeper columellar baffle (
|
||
<figureCitation box="[535,666,1711,1738]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="32" pageNumber="33">Figs 30-31</figureCitation>
|
||
,
|
||
<figureCitation box="[677,709,1711,1737]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="32" pageNumber="33">36</figureCitation>
|
||
,
|
||
<figureCitation box="[721,791,1711,1738]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="32" pageNumber="33">58-59</figureCitation>
|
||
) .......................................................................... 10
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="32.[189,1398,1674,1812]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
– Shallow columellar dentition projecting less far into aperture than any deeper columellar baffle (
|
||
<figureCitation captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="32" pageNumber="33">Figs 51-57, 60-64</figureCitation>
|
||
) ................................................................................................................................. 12
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="32" pageNumber="33">
|
||
<paragraph blockId="32.[189,1399,1853,2027]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
10. Apex rounded; spire broadly acuminate, spire angle 70°; ribs weak, almost disappearing in the middle of the whorl (Virunga NP,
|
||
<collectingCountry box="[634,762,1890,1916]" name="Democratic Republic of the Congo" pageId="32" pageNumber="33">DR Congo</collectingCountry>
|
||
) (
|
||
<figureCitation box="[786,872,1890,1917]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="32" pageNumber="33">Fig. 36</figureCitation>
|
||
) .................................................................... ..............................................................
|
||
<taxonomicName authority="(van Bruggen & Van Goethem, 1999)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen & Van Goethem" baseAuthorityYear="1999" box="[703,1262,1926,1953]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="selene" status="comb. nov.">
|
||
<emphasis bold="true" box="[703,814,1926,1953]" italics="true" pageId="32" pageNumber="33">D. selene</emphasis>
|
||
(van
|
||
<bibRefCitation author="Bruggen A. C. van & Van Goethem J. L." box="[880,1253,1927,1953]" pageId="32" pageNumber="33" pagination="31 - 45" refId="ref26326" refString="Bruggen A. C. van & Van Goethem J. L. 1999. Dr. William Adam's iconography of Central and West African Gulella species (Gastropoda Pulmonata: Streptaxidae). Part 3: Nine new species from the D. R. Congo. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Biologie 69: 31 - 45." type="journal article" year="1999">Bruggen & Van Goethem, 1999</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1269,1399,1927,1953]" pageId="32" pageNumber="33" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="32.[189,1399,1853,2027]" pageId="32" pageNumber="33">
|
||
<keyLead pageId="32" pageNumber="33">
|
||
– Apex sharply pointed; spire acuminate or coeloconoid, spire angle 65° or less; ribs prominent, running from suture to suture; apex finely radially striate in fresh specimens (
|
||
<figureCitation box="[1140,1269,2001,2027]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="32" pageNumber="33">Figs 30-31</figureCitation>
|
||
) .......... 11
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="33" pageNumber="34">
|
||
<paragraph blockId="33.[189,1399,267,401]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
11. Shell more than 4.0 mm high; parietal tooth simple, lamella-like; two palatal teeth (W. Usambara Mts,
|
||
<collectingCountry box="[294,402,303,330]" name="Tanzania" pageId="33" pageNumber="34">Tanzania</collectingCountry>
|
||
) (
|
||
<figureCitation box="[425,516,303,329]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="33" pageNumber="34">Figs 31</figureCitation>
|
||
,
|
||
<figureCitation box="[528,559,303,330]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">59</figureCitation>
|
||
) ..........................................
|
||
<taxonomicName authority="(Verdcourt, 1962)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1962" box="[894,1262,303,330]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="cuspidata" status="comb. nov.">
|
||
<emphasis bold="true" box="[894,1046,303,330]" italics="true" pageId="33" pageNumber="34">D. cuspidata</emphasis>
|
||
(
|
||
<bibRefCitation author="Verdcourt B." box="[1061,1253,303,330]" pageId="33" pageNumber="34" refId="ref27856" refString="Verdcourt B. 1962. Preliminary Keys for the Identification of the Species of the Genus Gulella Pfr. Occurring in East Africa. Annales du Musee Royal de l'Afrique Centrale 106, Musee Royal de l'Afrique Centrale, Tervuren." type="journal volume" year="1962">Verdcourt, 1962</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1269,1399,303,330]" pageId="33" pageNumber="34" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="33.[189,1399,267,401]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
– Shell up to 4.0 mm high; parietal tooth V-shaped; one palatal tooth (elsewhere in
|
||
<collectingCountry box="[1174,1282,339,366]" name="Tanzania" pageId="33" pageNumber="34">Tanzania</collectingCountry>
|
||
) (
|
||
<figureCitation box="[1303,1394,339,365]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="33" pageNumber="34">Figs 30</figureCitation>
|
||
,
|
||
<figureCitation box="[234,265,374,401]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">58</figureCitation>
|
||
) .....................................................................................................................
|
||
<taxonomicName authority="Rowson & Tattersfield, 2013" authorityName="Rowson & Tattersfield" authorityYear="2013" box="[1156,1301,375,401]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="minareta" status="sp. nov.">
|
||
<emphasis bold="true" box="[1156,1301,375,401]" italics="true" pageId="33" pageNumber="34">D. minareta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1308,1399,375,401]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="33" pageNumber="34">
|
||
<paragraph blockId="33.[189,1399,433,567]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
12. Outer palatal surface of aperture with very deep, long, furrow-like depression; basal denticle absent or not visible in apertural view (
|
||
<figureCitation box="[611,746,469,495]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="33" pageNumber="34">Figs 29, 33</figureCitation>
|
||
,
|
||
<figureCitation box="[758,789,469,496]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">57</figureCitation>
|
||
) ........................................................................... 13
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="33.[189,1399,433,567]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">– Outer palatal surface of aperture with shallow, not furrow-like depression; basal denticle clearly visible in apertural view ................................................................................................................. 14</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="33" pageNumber="34">
|
||
<paragraph blockId="33.[189,1399,598,661]" box="[189,1399,598,626]" pageId="33" pageNumber="34">
|
||
<keyLead box="[189,1399,598,626]" pageId="33" pageNumber="34">
|
||
13. Apex sharply pointed; spire coeloconoid (Ngorongoro,
|
||
<collectingCountry box="[861,965,599,626]" name="Tanzania" pageId="33" pageNumber="34">Tanzania</collectingCountry>
|
||
) (
|
||
<figureCitation box="[987,1076,599,625]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="33" pageNumber="34">Figs 29</figureCitation>
|
||
,
|
||
<figureCitation box="[1087,1117,599,626]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">57</figureCitation>
|
||
) ..
|
||
<taxonomicName authority="Rowson & Tattersfield, 2013" authorityName="Rowson & Tattersfield" authorityYear="2013" box="[1157,1305,598,625]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="cresswelli" status="sp. nov.">
|
||
<emphasis bold="true" box="[1157,1305,598,625]" italics="true" pageId="33" pageNumber="34">D. cresswelli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1311,1399,599,625]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="33.[189,1399,598,661]" box="[193,1399,635,661]" pageId="33" pageNumber="34">
|
||
<keyLead box="[193,1399,635,661]" pageId="33" pageNumber="34">
|
||
– Apex rounded (
|
||
<collectingRegion box="[416,476,635,661]" country="Kenya" name="Taita/Taveta" pageId="33" pageNumber="34">Taita</collectingRegion>
|
||
Hills,
|
||
<collectingCountry box="[556,634,635,661]" name="Kenya" pageId="33" pageNumber="34">Kenya</collectingCountry>
|
||
) (
|
||
<figureCitation box="[658,743,635,661]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="33" pageNumber="34">Fig. 33</figureCitation>
|
||
) ......
|
||
<taxonomicName authority="(Rowson & Lange, 2007)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Rowson & Lange" baseAuthorityYear="2007" box="[815,1262,635,661]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="nictitans" status="comb. nov.">
|
||
<emphasis bold="true" box="[815,954,635,661]" italics="true" pageId="33" pageNumber="34">D. nictitans</emphasis>
|
||
(
|
||
<bibRefCitation author="Rowson B. & Lange C. N." box="[970,1253,635,661]" pageId="33" pageNumber="34" pagination="21 - 32" refId="ref27291" refString="Rowson B. & Lange C. N. 2007. Two new species of Gulella (Mollusca: Pulmonata: Streptaxidae) from the Taita Hills, Kenya. African Invertebrates 48 (2): 21 - 32." type="journal article" year="2007">Rowson & Lange, 2007</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1269,1399,635,661]" pageId="33" pageNumber="34" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="33" pageNumber="34">
|
||
<paragraph blockId="33.[189,1399,693,792]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
14. Sculpture of widely-spaced, flaring, subtriangular, lamella-like ribs; spire narrowly acuminate (spire angle <50°) (
|
||
<collectingRegion box="[388,561,729,755]" country="Mozambique" name="Cabo Delgado" pageId="33" pageNumber="34">Cabo Delgado</collectingRegion>
|
||
,
|
||
<collectingCountry box="[573,730,729,756]" name="Mozambique" pageId="33" pageNumber="34">Mozambique</collectingCountry>
|
||
) (
|
||
<figureCitation box="[754,839,729,755]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="33" pageNumber="34">Fig. 27</figureCitation>
|
||
) .........
|
||
<taxonomicName authority="(Muratov, 2010)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Muratov" baseAuthorityYear="2010" box="[932,1262,729,756]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="delgada" status="comb. nov.">
|
||
<emphasis bold="true" box="[932,1063,729,756]" italics="true" pageId="33" pageNumber="34">D. delgada</emphasis>
|
||
(
|
||
<bibRefCitation author="Muratov I. V." box="[1078,1253,729,755]" pageId="33" pageNumber="34" pagination="255 - 288" refId="ref26968" refString="Muratov I. V. 2010. Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique. African Invertebrates 51 (2): 255 - 288. http: // dx. doi. org / 10.5733 / afin. 051.0203" type="journal article" year="2010">Muratov, 2010</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1269,1399,729,755]" pageId="33" pageNumber="34" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="33.[189,1399,693,792]" box="[193,1398,765,792]" pageId="33" pageNumber="34">
|
||
<keyLead box="[193,1398,765,792]" pageId="33" pageNumber="34">– Shell without this combination of characters ................................................................................. 15</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="33" pageNumber="34">
|
||
<paragraph blockId="33.[189,1399,827,889]" box="[189,1399,827,854]" pageId="33" pageNumber="34">
|
||
<keyLead box="[189,1399,827,854]" pageId="33" pageNumber="34">
|
||
15. Shell
|
||
<quantity box="[302,392,827,853]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="33" pageNumber="34" unit="mm" value="4.8">4.8 mm</quantity>
|
||
<emphasis box="[392,402,828,853]" italics="true" pageId="33" pageNumber="34">-</emphasis>
|
||
<quantity box="[401,491,827,854]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="33" pageNumber="34" unit="mm" value="5.5">5.5 mm</quantity>
|
||
high (
|
||
<collectingRegion box="[566,648,827,853]" country="Tanzania" name="Kusini Pemba" pageId="33" pageNumber="34">Pemba</collectingRegion>
|
||
I.,
|
||
<collectingCountry box="[686,794,827,854]" name="Tanzania" pageId="33" pageNumber="34">Tanzania</collectingCountry>
|
||
) (
|
||
<figureCitation box="[817,908,827,853]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson & Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="33" pageNumber="34">Figs 32</figureCitation>
|
||
,
|
||
<figureCitation box="[920,951,827,854]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">60</figureCitation>
|
||
) .....................
|
||
<taxonomicName authority="Rowson & Tattersfield, 2013" authorityName="Rowson & Tattersfield" authorityYear="2013" box="[1140,1300,827,854]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="pembensis" status="sp. nov.">
|
||
<emphasis bold="true" box="[1140,1300,827,854]" italics="true" pageId="33" pageNumber="34">D. pembensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1308,1399,827,853]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="33.[189,1399,827,889]" box="[193,1398,863,889]" pageId="33" pageNumber="34">
|
||
<keyLead box="[193,1398,863,889]" pageId="33" pageNumber="34">
|
||
– Shell up to
|
||
<quantity box="[369,459,863,889]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="33" pageNumber="34" unit="mm" value="4.6">4.6 mm</quantity>
|
||
high ................................................................................................................. 16
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="33" pageNumber="34">
|
||
<paragraph blockId="33.[189,1399,921,1091]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
16. Shell up to
|
||
<quantity box="[375,467,921,948]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="33" pageNumber="34" unit="mm" value="3.3">3.3 mm</quantity>
|
||
high; parieto-palatal sinus narrow, appearing parallel-sided in apertural view; nearly always with an additional, small parietal denticle to the left of parietal lamella; nearly always with two basal denticles (
|
||
<figureCitation box="[533,662,993,1019]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="33" pageNumber="34">Figs 21-26</figureCitation>
|
||
,
|
||
<figureCitation box="[675,745,993,1020]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">54-55</figureCitation>
|
||
) ... 17 (
|
||
<taxonomicName authority="(van Bruggen, 1969)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[839,1210,992,1020]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
|
||
<emphasis bold="true" box="[839,960,992,1019]" italics="true" pageId="33" pageNumber="34">D. browni</emphasis>
|
||
(van
|
||
<bibRefCitation author="Bruggen A. C. van" box="[1026,1202,993,1020]" pageId="33" pageNumber="34" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">Bruggen, 1969</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1218,1348,993,1019]" pageId="33" pageNumber="34" rank="species">comb. nov.</taxonomicNameLabel>
|
||
<emphasis box="[1354,1389,993,1019]" italics="true" pageId="33" pageNumber="34">s.l.</emphasis>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="33.[189,1399,921,1091]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
– Shell up to
|
||
<quantity box="[360,446,1028,1055]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="33" pageNumber="34" unit="mm" value="4.6">4.6 mm</quantity>
|
||
high; parieto-palatal sinus wide, not parallel-sided in apertural view; always without a small parietal denticle to the left of parietal lamella, even if shell less than
|
||
<quantity box="[1105,1192,1064,1091]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="33" pageNumber="34" unit="mm" value="3.3">3.3 mm</quantity>
|
||
high .............. 19
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="33" pageNumber="34">
|
||
<paragraph blockId="33.[189,1399,1123,1257]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
17. Only one basal denticle present; no columellar dentition other than a deep-set baffle (Mafia I.,
|
||
<collectingCountry box="[234,342,1158,1185]" name="Tanzania" pageId="33" pageNumber="34">Tanzania</collectingCountry>
|
||
) (
|
||
<figureCitation box="[365,457,1158,1185]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="33" pageNumber="34">Figs 21</figureCitation>
|
||
,
|
||
<figureCitation box="[469,500,1158,1185]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">54</figureCitation>
|
||
) ...................................................................
|
||
<taxonomicName box="[1018,1260,1158,1185]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="subSpecies" species="browni" subSpecies="mafiensis">
|
||
<emphasis bold="true" box="[1018,1260,1158,1185]" italics="true" pageId="33" pageNumber="34">D. browni mafiensis</emphasis>
|
||
</taxonomicName>
|
||
subsp. nov.
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="33.[189,1399,1123,1257]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
– Two basal denticles almost always present; one or two columellar denticles present in addition to a deep-set baffle (
|
||
<figureCitation box="[422,552,1230,1256]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="33" pageNumber="34">Figs 22-26</figureCitation>
|
||
,
|
||
<figureCitation box="[564,594,1230,1257]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">55</figureCitation>
|
||
) ..................................................................................................... 18
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="33" pageNumber="34">
|
||
<paragraph blockId="33.[189,1399,1288,1494]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
18. Shell
|
||
<quantity box="[308,479,1288,1315]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.25" metricValueMin="3.15" pageId="33" pageNumber="34" unit="mm" value="3.2" valueMax="3.25" valueMin="3.15">
|
||
3.15
|
||
<emphasis box="[359,369,1289,1314]" italics="true" pageId="33" pageNumber="34">-</emphasis>
|
||
3.25 mm
|
||
</quantity>
|
||
high; sutures relatively shallow; peristome complete; parieto-palatal sinus longer and narrower, not widening appreciably towards its inner end (Semuliki NP,
|
||
<collectingCountry box="[1229,1322,1324,1351]" name="Uganda" pageId="33" pageNumber="34">Uganda</collectingCountry>
|
||
) (
|
||
<figureCitation captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="33" pageNumber="34">Figs 22</figureCitation>
|
||
,
|
||
<figureCitation box="[278,310,1360,1387]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">55</figureCitation>
|
||
; compare with
|
||
<figureCitation box="[491,618,1360,1387]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="33" pageNumber="34">Figs 23-26</figureCitation>
|
||
) ............................................
|
||
<taxonomicName box="[969,1260,1360,1387]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="subSpecies" species="browni" subSpecies="semulikiensis">
|
||
<emphasis bold="true" box="[969,1260,1360,1387]" italics="true" pageId="33" pageNumber="34">D. browni semulikiensis</emphasis>
|
||
</taxonomicName>
|
||
subsp. nov.
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="33.[189,1399,1288,1494]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
– Shell (
|
||
<quantity box="[311,480,1396,1423]" metricMagnitude="-3" metricUnit="m" metricValue="2.875" metricValueMax="3.2" metricValueMin="2.55" pageId="33" pageNumber="34" unit="mm" value="2.875" valueMax="3.2" valueMin="2.55">
|
||
2.55
|
||
<emphasis box="[364,374,1397,1422]" italics="true" pageId="33" pageNumber="34">-</emphasis>
|
||
3.20 mm
|
||
</quantity>
|
||
high; sutures relatively deep; peristome incomplete parietally; parieto-palatal sinus shorter and broader, widening towards its inner end (
|
||
<collectingCountry box="[919,1072,1432,1458]" name="South Africa" pageId="33" pageNumber="34">South Africa</collectingCountry>
|
||
,
|
||
<collectingCountry box="[1084,1242,1432,1458]" name="Mozambique" pageId="33" pageNumber="34">Mozambique</collectingCountry>
|
||
, S.
|
||
<collectingCountry box="[1283,1391,1432,1459]" name="Tanzania" pageId="33" pageNumber="34">Tanzania</collectingCountry>
|
||
) (
|
||
<figureCitation box="[243,374,1467,1493]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="33" pageNumber="34">Figs 23-26</figureCitation>
|
||
; compare with
|
||
<figureCitation box="[554,644,1467,1494]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="33" pageNumber="34">Figs 22</figureCitation>
|
||
,
|
||
<figureCitation box="[656,686,1467,1494]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">55</figureCitation>
|
||
) ............
|
||
<taxonomicName authority="browni (van Bruggen, 1969)" authorityName="browni (van Bruggen" authorityYear="1969" box="[798,1262,1467,1494]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="subSpecies" species="browni" status="comb. nov." subSpecies="browni">
|
||
<emphasis bold="true" box="[798,1011,1467,1494]" italics="true" pageId="33" pageNumber="34">D. browni browni</emphasis>
|
||
(van
|
||
<bibRefCitation author="Bruggen A. C. van" box="[1078,1253,1467,1494]" pageId="33" pageNumber="34" pagination="3 - 116" refId="ref26021" refString="Bruggen A. C. van 1969. Studies on the land molluscs of Zululand, with notes on the distribution of land molluscs in Southern Africa. Zoologische Verhandelingen 103: 3 - 116." type="journal article" year="1969">Bruggen, 1969</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1269,1399,1467,1494]" pageId="33" pageNumber="34" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="33" pageNumber="34">
|
||
<paragraph blockId="33.[189,1399,1526,1624]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
19. Sculpture of numerous, fine ribs (15 or more per mm on penultimate whorl); spire coeloconoid, elongate (
|
||
<collectingCountry box="[349,450,1562,1588]" name="Mayotte" pageId="33" pageNumber="34">Mayotte</collectingCountry>
|
||
) (
|
||
<figureCitation box="[474,565,1562,1588]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="33" pageNumber="34">Figs 40</figureCitation>
|
||
,
|
||
<figureCitation box="[577,608,1562,1589]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">64</figureCitation>
|
||
) ............................................
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" box="[964,1261,1561,1588]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="subSpecies" species="minuscula" subSpecies="mahorana">
|
||
<emphasis bold="true" box="[964,1261,1561,1588]" italics="true" pageId="33" pageNumber="34">D. minuscula mahorana</emphasis>
|
||
</taxonomicName>
|
||
subsp. nov.
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="33.[189,1399,1526,1624]" box="[193,1398,1597,1624]" pageId="33" pageNumber="34">
|
||
<keyLead box="[193,1398,1597,1624]" pageId="33" pageNumber="34">
|
||
– Shell without this combination of characters (e.g.
|
||
<figureCitation box="[820,934,1597,1624]" captionStart-0="Figs 1-5" captionStart-1="Figs 6-26" captionStartId-0="6.[189,242,1930,1956]" captionStartId-1="9.[189,241,1754,1780]" captionTargetBox-0="[363,1200,672,1890]" captionTargetBox-1="[295,1292,265,1714]" captionTargetId-0="figure@6.[363,1200,672,1890]" captionTargetId-1="figure@9.[295,1292,265,1714]" captionTargetPageId-0="6" captionTargetPageId-1="9" captionText-0="Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view." captionText-1="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi-0="http://doi.org/10.5281/zenodo.3806774" figureDoi-1="http://doi.org/10.5281/zenodo.3806778" httpUri-0="https://zenodo.org/record/3806774/files/figure.png" httpUri-1="https://zenodo.org/record/3806778/files/figure.png" pageId="33" pageNumber="34">Figs 1-20</figureCitation>
|
||
,
|
||
<figureCitation box="[946,1016,1598,1624]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="33" pageNumber="34">38-39</figureCitation>
|
||
) ............................................ 20
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="33" pageNumber="34">
|
||
<paragraph blockId="33.[189,1399,1656,1790]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
20. Shell 4.0 mm or more high, sculpture of few, coarse, widely-spaced ribs (5
|
||
<emphasis box="[1087,1097,1657,1682]" italics="true" pageId="33" pageNumber="34">-</emphasis>
|
||
10 per mm on penultimate whorl); dentition 4-fold to 5-fold (Taru Desert and elsewhere in SE
|
||
<collectingCountry box="[1028,1106,1692,1718]" name="Kenya" pageId="33" pageNumber="34">Kenya</collectingCountry>
|
||
) (
|
||
<figureCitation box="[1130,1231,1692,1718]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="33" pageNumber="34">Figs 6-8</figureCitation>
|
||
,
|
||
<figureCitation box="[1243,1273,1692,1719]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">51</figureCitation>
|
||
). ............ .........................................................................
|
||
<taxonomicName authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[787,900,1727,1754]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="radius">
|
||
<emphasis bold="true" box="[787,900,1727,1754]" italics="true" pageId="33" pageNumber="34">D. radius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName authority="(Connolly, 1922)" authorityName="Rowson & Tattersfield" authorityYear="2013" baseAuthorityName="Connolly" baseAuthorityYear="1922" box="[907,1178,1727,1754]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="calva" status="comb. et stat. nov.">
|
||
<emphasis bold="true" box="[907,971,1727,1754]" italics="true" pageId="33" pageNumber="34">calva</emphasis>
|
||
(
|
||
<bibRefCitation author="Connolly M." box="[986,1169,1727,1754]" pageId="33" pageNumber="34" pagination="485 - 517" refId="ref26391" refString="Connolly M. 1922. Notes on African Non-marine Mollusca, with Descriptions of Many New Species. Annals and Magazine of Natural History Series 9 10 (59): 485 - 517, pl. XIV. http: // dx. doi. org / 10.1080 / 00222932208632797" type="journal article" year="1922">Connolly, 1922</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1185,1399,1727,1754]" pageId="33" pageNumber="34">comb. et stat. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="33.[189,1399,1656,1790]" box="[193,1398,1763,1790]" pageId="33" pageNumber="34">
|
||
<keyLead box="[193,1398,1763,1790]" pageId="33" pageNumber="34">– Shell without this combination of characters ................................................................................. 21</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="33" pageNumber="34">
|
||
<paragraph blockId="33.[189,1399,1822,2027]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
21. Dentition 5-fold or more, although when shallow columellar dentition is weak, might be considered 4-fold (lowlands of SE.
|
||
<collectingCountry box="[517,594,1858,1884]" name="Kenya" pageId="33" pageNumber="34">Kenya</collectingCountry>
|
||
and NE.
|
||
<collectingCountry box="[704,812,1857,1884]" name="Tanzania" pageId="33" pageNumber="34">Tanzania</collectingCountry>
|
||
) (
|
||
<figureCitation box="[835,935,1857,1884]" captionStart="Figs 1-5" captionStartId="6.[189,242,1930,1956]" captionTargetBox="[363,1200,672,1890]" captionTargetId="figure@6.[363,1200,672,1890]" captionTargetPageId="6" captionText="Figs 1-5. Lectotype of Ennea radius Preston, 1910, type species of Dadagulella gen. nov. 1. Apical view. 2. Apertural view. 3. Last two whorls, from left side. 4. Last two whorls, from right side. 5. Umbilical view." figureDoi="http://doi.org/10.5281/zenodo.3806774" httpUri="https://zenodo.org/record/3806774/files/figure.png" pageId="33" pageNumber="34">Figs 1-5</figureCitation>
|
||
,
|
||
<figureCitation box="[948,1004,1858,1884]" captionStart="Figs 6-26" captionStartId="9.[189,241,1754,1780]" captionTargetBox="[295,1292,265,1714]" captionTargetId="figure@9.[295,1292,265,1714]" captionTargetPageId="9" captionText="Figs 6-26.Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969)." figureDoi="http://doi.org/10.5281/zenodo.3806778" httpUri="https://zenodo.org/record/3806778/files/figure.png" pageId="33" pageNumber="34">9-20</figureCitation>
|
||
,
|
||
<figureCitation box="[1016,1086,1857,1884]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">52-53</figureCitation>
|
||
) ....................................... .....................................................................................
|
||
<taxonomicName authority="radius (Preston, 1910)" authorityName="radius (Preston" authorityYear="1910" box="[877,1262,1893,1920]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="variety" species="radius" status="comb. nov." variety="radius">
|
||
<emphasis bold="true" box="[877,1074,1893,1920]" italics="true" pageId="33" pageNumber="34">D. radius radius</emphasis>
|
||
(
|
||
<bibRefCitation author="Preston H. B." box="[1089,1253,1893,1919]" pageId="33" pageNumber="34" pagination="526 - 536" refId="ref27068" refString="Preston H. B. 1910. Additions to the non-Marine Molluscan Fauna of British and German East Africa and Lake Albert Edward. Annals and Magazine of Natural History, Series 8 6: 526 - 536. http: // dx. doi. org / 10.1080 / 00222931008692885" type="journal article" year="1910">Preston, 1910</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1269,1399,1893,1920]" pageId="33" pageNumber="34" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="33.[189,1399,1822,2027]" pageId="33" pageNumber="34">
|
||
<keyLead pageId="33" pageNumber="34">
|
||
– Dentition 4-fold, lacking a shallow columellar tooth (as in the
|
||
<typeStatus box="[979,1088,1929,1956]" pageId="33" pageNumber="34">lectotype</typeStatus>
|
||
) or 5-fold, with a shallow columellar tooth (as in the
|
||
<typeStatus box="[550,710,1965,1992]" pageId="33" pageNumber="34">paralectotype</typeStatus>
|
||
) (
|
||
<collectingRegion box="[733,826,1965,1991]" country="Comoros" name="Anjouan" pageId="33" pageNumber="34">Nzwani</collectingRegion>
|
||
[
|
||
<collectingRegion box="[840,947,1964,1991]" country="Comoros" name="Anjouan" pageId="33" pageNumber="34">Anjouan</collectingRegion>
|
||
] I.,
|
||
<collectingCountry box="[993,1102,1965,1991]" name="Comoros" pageId="33" pageNumber="34">Comoros</collectingCountry>
|
||
) (
|
||
<figureCitation box="[1126,1256,1965,1991]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen & Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen & Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="33" pageNumber="34">Figs 38-39</figureCitation>
|
||
,
|
||
<figureCitation box="[1268,1338,1965,1992]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="33" pageNumber="34">62-63</figureCitation>
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) ..... .......................................................................
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<taxonomicName authority="minuscula (Morelet, 1877)" authorityName="minuscula (Morelet" authorityYear="1877" box="[771,1261,2000,2027]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="subSpecies" species="minuscula" status="comb. nov." subSpecies="minuscula">
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||
<emphasis bold="true" box="[771,1069,2000,2027]" italics="true" pageId="33" pageNumber="34">D. minuscula minuscula</emphasis>
|
||
(
|
||
<bibRefCitation author="Morelet A." box="[1084,1253,2001,2027]" pageId="33" pageNumber="34" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet, 1877</bibRefCitation>
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||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1269,1399,2001,2027]" pageId="33" pageNumber="34" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
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||
</subSection>
|
||
</treatment>
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||
</document> |