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<document ID-DOI="http://dx.doi.org/10.5852/ejt.2013.37" ID-GBIF-Dataset="9682b45b-5e22-4965-a293-e203660e0ed3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3806770" ID-ZooBank="ADF6394E-77B5-4309-87FE-4097FDE0A3FD" checkinTime="1588796067459" checkinUser="carolina" docAuthor="Rowson, Ben &amp; Tattersfield, Peter" docDate="2013" docId="4511E41DD832FFD1FDE8FD21F0B7F80C" docLanguage="en" docName="ejt-2013-37.pdf.imf" docOrigin="European Journal of Taxonomy 37" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article" docStyleVersion="14" docTitle="Dadagulella frontierarum Rowson &amp; Tattersfield 2013, sp. nov." docType="treatment" docUuid="9D028B17-9FCB-47E8-96B1-5901B8E3EA4E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="23" masterDocId="B9289C65D824FFC7FFA4FF80F253FFE7" masterDocTitle="Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies" masterLastPageNumber="46" masterPageNumber="1" pageNumber="23" updateTime="1683580915233" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rowson, Ben</mods:namePart>
<mods:nameIdentifier type="LSID">05D46259-DA45-45A4-9127-5AF361D566A7</mods:nameIdentifier>
<mods:affiliation>Biodiversity &amp; Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk &amp; urn: lsid: zoobank. org: author: 05 D 46259 - DA 45 - 45 A 4 - 9127 - 5 AF 361 D 566 A 7</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tattersfield, Peter</mods:namePart>
<mods:nameIdentifier type="LSID">8C55642E-B18A-461B-9BF9-31CB6DB7A070</mods:nameIdentifier>
<mods:affiliation>Biodiversity &amp; Systematic Biology, National Museum of Wales, Cathays Park, Cardiff, UK CF 10 3 NP. Email: ben. rowson @ museumwales. ac. uk &amp; urn: lsid: zoobank. org: author: 8 C 55642 E-B 18 A- 461 B- 9 BF 9 - 31 CB 6 DB 7 A 070</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-02-04</mods:number>
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<mods:detail type="issue">
<mods:number>37</mods:number>
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<mods:start>1</mods:start>
<mods:end>46</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.5852/ejt.2013.37</mods:identifier>
<mods:identifier type="GBIF-Dataset">9682b45b-5e22-4965-a293-e203660e0ed3</mods:identifier>
<mods:identifier type="ISSN">2118-9773</mods:identifier>
<mods:identifier type="Zenodo-Dep">3806770</mods:identifier>
<mods:identifier type="ZooBank">ADF6394E-77B5-4309-87FE-4097FDE0A3FD</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3815761" ID-GBIF-Taxon="163971594" ID-Zenodo-Dep="3815761" LSID="urn:lsid:zoobank.org:act:9D028B17-9FCB-47E8-96B1-5901B8E3EA4E" httpUri="http://treatment.plazi.org/id/4511E41DD832FFD1FDE8FD21F0B7F80C" lastPageNumber="23" pageId="22" pageNumber="23">
<subSubSection pageId="22" pageNumber="23" type="nomenclature">
<paragraph blockId="22.[367,1221,673,771]" box="[588,1000,673,700]" pageId="22" pageNumber="23">
<heading box="[588,1000,673,700]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="23" reason="2">
<taxonomicName ID-CoL="33XQH" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[588,902,673,700]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="frontierarum" status="sp. nov.">
<emphasis bold="true" box="[588,902,673,700]" italics="true" pageId="22" pageNumber="23">Dadagulella frontierarum</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[909,1000,674,700]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph blockId="22.[367,1221,673,771]" box="[367,1221,709,736]" pageId="22" pageNumber="23">
<uri box="[367,1221,709,736]" pageId="22" pageNumber="23">
urn:lsid:zoobank.org:act:
<uuid box="[661,1221,709,736]" pageId="22" pageNumber="23">
9D028
<collectionCode box="[743,762,709,736]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="22" pageNumber="23" type="Herbarium">B</collectionCode>
17-9FCB-47
<collectionCode box="[912,931,709,736]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="22" pageNumber="23" type="Herbarium">E</collectionCode>
8-96
<collectionCode box="[983,1002,709,736]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="22" pageNumber="23" type="Herbarium">B</collectionCode>
1-5901
<collectionCode box="[1080,1099,709,736]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="22" pageNumber="23" type="Herbarium">B</collectionCode>
8
<collectionCode box="[1112,1131,709,736]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="22" pageNumber="23" type="Herbarium">E</collectionCode>
3
<collectionCode box="[1144,1187,709,736]" country="Kenya" lsid="urn:lsid:biocol.org:col:15088" name="National Museums of Kenya - East African Herbarium" pageId="22" pageNumber="23" type="Herbarium">EA</collectionCode>
4
<collectionCode box="[1202,1221,709,736]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="22" pageNumber="23" type="Herbarium">E</collectionCode>
</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection box="[655,932,745,771]" pageId="22" pageNumber="23" type="description">
<paragraph blockId="22.[367,1221,673,771]" box="[655,932,745,771]" pageId="22" pageNumber="23">
<figureCitation box="[655,745,745,771]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson &amp; Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="22" pageNumber="23">Figs 34</figureCitation>
,
<figureCitation box="[757,789,745,771]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="22" pageNumber="23">61</figureCitation>
,
<figureCitation box="[801,834,745,771]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="22" pageNumber="23">84</figureCitation>
;
<tableCitation box="[845,932,745,771]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="22" pageNumber="23" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="etymology">
<paragraph blockId="22.[189,324,812,838]" box="[189,324,812,838]" pageId="22" pageNumber="23">
<heading bold="true" box="[189,324,812,838]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis bold="true" box="[189,324,812,838]" pageId="22" pageNumber="23">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="22.[189,1398,859,921]" pageId="22" pageNumber="23">
After Frontier
<collectingCountry box="[359,468,859,885]" name="Tanzania" pageId="22" pageNumber="23">Tanzania</collectingCountry>
, the organisation which collected the specimens; given the ending -
<emphasis box="[1267,1329,860,885]" italics="true" pageId="22" pageNumber="23">arum</emphasis>
for a feminine noun in the genitive plural.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="materials_examined">
<paragraph blockId="22.[189,492,961,987]" box="[189,492,961,987]" pageId="22" pageNumber="23">
<heading bold="true" box="[189,492,961,987]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis bold="true" box="[189,492,961,987]" pageId="22" pageNumber="23">
<typeStatus box="[189,250,961,987]" pageId="22" pageNumber="23">Type</typeStatus>
material examined
</emphasis>
</heading>
</paragraph>
<paragraph blockId="22.[189,1399,1008,1178]" pageId="22" pageNumber="23">
<materialsCitation ID-GBIF-Occurrence="2610647311" pageId="22" pageNumber="23">
<collectingCountry box="[189,343,1008,1034]" name="Tanzania" pageId="22" pageNumber="23">TANZANIA</collectingCountry>
:
<typeStatus box="[357,460,1008,1034]" pageId="22" pageNumber="23">holotype</typeStatus>
<collectionCode box="[470,548,1008,1034]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="23">NMW</collectionCode>
.
<collectionCode box="[552,570,1008,1034]" country="Switzerland" name="Universität Zürich" pageId="22" pageNumber="23" type="Herbarium">Z</collectionCode>
.2003.074.00001:
<specimenCount box="[784,847,1008,1034]" pageId="22" pageNumber="23" type="generic">1 ad.</specimenCount>
, Mtai Forest Reserve (
<geoCoordinate box="[1126,1209,1008,1034]" degrees="4.87" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="555" value="-4.87">
4.87°
<collectionCode box="[1194,1209,1008,1034]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="22" pageNumber="23" type="Herbarium">S</collectionCode>
</geoCoordinate>
,
<geoCoordinate box="[1225,1324,1008,1034]" degrees="38.77" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="555" value="38.77">
38.77°
<collectionCode box="[1305,1324,1008,1034]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="22" pageNumber="23" type="Herbarium">E</collectionCode>
</geoCoordinate>
), East Usambara Mts, Muheza District (plot 69/3), submontane forest at
<quantity box="[938,1009,1044,1070]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="22" pageNumber="23" unit="m" value="970.0">970 m</quantity>
alt., leg. Frontier
<collectingCountry box="[1208,1317,1044,1070]" name="Tanzania" pageId="22" pageNumber="23">Tanzania</collectingCountry>
,
<date pageId="22" pageNumber="23" value="1997-03-08">
8 Mar.
<date box="[189,251,1080,1106]" pageId="22" pageNumber="23" value="1997" year="1997">1997</date>
</date>
</materialsCitation>
.
<typeStatus box="[263,377,1080,1106]" pageId="22" pageNumber="23">Paratypes</typeStatus>
<collectionCode box="[386,464,1080,1106]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="23">NMW</collectionCode>
.
<collectionCode box="[467,485,1080,1106]" country="Switzerland" name="Universität Zürich" pageId="22" pageNumber="23" type="Herbarium">Z</collectionCode>
.2003.074.00002: 2 ads, data as
<typeStatus box="[865,972,1080,1106]" pageId="22" pageNumber="23">holotype</typeStatus>
.
<typeStatus box="[984,1087,1080,1106]" pageId="22" pageNumber="23">Paratype</typeStatus>
<collectionCode box="[1095,1173,1080,1106]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="23">NMW</collectionCode>
.
<collectionCode box="[1176,1194,1080,1106]" country="Switzerland" name="Universität Zürich" pageId="22" pageNumber="23" type="Herbarium">Z</collectionCode>
.2003.074.00003:
<specimenCount box="[189,251,1116,1142]" pageId="22" pageNumber="23" type="generic">1 ad.</specimenCount>
, data as
<typeStatus box="[361,468,1116,1142]" pageId="22" pageNumber="23">holotype</typeStatus>
.
<typeStatus box="[482,585,1116,1142]" pageId="22" pageNumber="23">Paratype</typeStatus>
<collectionCode box="[595,673,1116,1142]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="22" pageNumber="23">NMW</collectionCode>
.
<collectionCode box="[676,694,1116,1142]" country="Switzerland" name="Universität Zürich" pageId="22" pageNumber="23" type="Herbarium">Z</collectionCode>
.2003.074.00004:
<specimenCount box="[909,971,1116,1142]" pageId="22" pageNumber="23" type="generic">1 ad.</specimenCount>
, data as
<typeStatus box="[1081,1184,1116,1142]" pageId="22" pageNumber="23">holotype</typeStatus>
but
<date box="[1242,1394,1116,1142]" pageId="22" pageNumber="23" value="1997-03-07">
7 Mar.
<date box="[1332,1394,1116,1142]" pageId="22" pageNumber="23" value="1997" year="1997">1997</date>
</date>
.
<typeStatus box="[189,303,1152,1178]" pageId="22" pageNumber="23">Paratypes</typeStatus>
<collectionCode box="[310,378,1152,1178]" pageId="22" pageNumber="23">NMT</collectionCode>
: 2 ads, data as
<typeStatus box="[553,660,1152,1178]" pageId="22" pageNumber="23">holotype</typeStatus>
.
</paragraph>
<paragraph blockId="22.[189,505,1218,1244]" box="[189,505,1218,1244]" pageId="22" pageNumber="23">
<heading bold="true" box="[189,505,1218,1244]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis bold="true" box="[189,505,1218,1244]" pageId="22" pageNumber="23">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph blockId="22.[189,260,1266,1292]" box="[189,260,1266,1292]" pageId="22" pageNumber="23">None.</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="description">
<paragraph blockId="22.[189,334,1331,1357]" box="[189,334,1331,1357]" pageId="22" pageNumber="23">
<heading bold="true" box="[189,334,1331,1357]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis bold="true" box="[189,334,1331,1357]" pageId="22" pageNumber="23">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="22.[189,1399,1378,1692]" pageId="22" pageNumber="23">
<collectionCode box="[189,257,1379,1405]" pageId="22" pageNumber="23">SHELL</collectionCode>
(
<figureCitation box="[273,362,1379,1405]" captionStart="Figs 27-35" captionStartId="16.[189,241,1824,1850]" captionTargetBox="[189,1399,348,1784]" captionTargetId="figure@16.[189,1399,348,1784]" captionTargetPageId="16" captionText="Figs 27-35. Adult shells of Dadagulella gen. nov. species. 27. D. delgada (Muratov, 2010) comb. nov., syntype, Cabo Delgado (after Muratov 2010). 28. D. ecclesiola sp. nov., holotype, Kimboza. 29. D. cresswelli sp. nov., holotype, Ngorongoro. 30. D. minareta sp. nov., holotype, Kimboza. 31. D. cuspidata (Verdcourt, 1962) comb. nov., holotype, Shume. 32. D. pembensis sp. nov., holotype, Ngezi. 33. D. nictitans (Rowson &amp; Lange, 2007) comb. nov., holotype, Macha. 34. D. frontierarum sp. nov., holotype, Mtai. 35. D. delta sp. nov., holotype, Mwanihana." figureDoi="http://doi.org/10.5281/zenodo.3806780" httpUri="https://zenodo.org/record/3806780/files/figure.png" pageId="22" pageNumber="23">Figs 34</figureCitation>
,
<figureCitation box="[372,405,1378,1405]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="22" pageNumber="23">61</figureCitation>
). Medium-sized (
<quantity box="[608,773,1378,1405]" metricMagnitude="-3" metricUnit="m" metricValue="3.275" metricValueMax="3.4" metricValueMin="3.15" pageId="22" pageNumber="23" unit="mm" value="3.275" valueMax="3.4" valueMin="3.15">
3.15
<emphasis box="[661,671,1379,1404]" italics="true" pageId="22" pageNumber="23">-</emphasis>
3.40 mm
</quantity>
high x
<quantity box="[857,1021,1378,1405]" metricMagnitude="-3" metricUnit="m" metricValue="1.9249999999999998" metricValueMax="2.05" metricValueMin="1.8" pageId="22" pageNumber="23" unit="mm" value="1.9249999999999998" valueMax="2.05" valueMin="1.8">
1.80
<emphasis box="[908,918,1379,1404]" italics="true" pageId="22" pageNumber="23">-</emphasis>
2.05 mm
</quantity>
wide), of 4.5
<emphasis box="[1173,1183,1379,1404]" italics="true" pageId="22" pageNumber="23">-</emphasis>
5.0 whorls. Ovateacuminate, spire narrowly acuminate (spire angle 68
<emphasis box="[824,834,1415,1440]" italics="true" pageId="22" pageNumber="23">-</emphasis>
77°). Apex sharply pointed. Embryonic whorls smoothly granulate. Later whorls with few, widely-spaced, coarse ribs (5
<emphasis box="[1078,1088,1451,1476]" italics="true" pageId="22" pageNumber="23">-</emphasis>
7 per mm on penultimate whorl). Sutures very deep. Umbilicus narrowly open. Peristome complete, almost detached. Outer palatal surface of aperture with a depression corresponding to the lower palatal teeth. Dentition 6-fold, consisting of: one complex, V-shaped and flaring parietal tooth; three palatal teeth, the lower two larger and set low down on the palatal surface, not forming a parieto-palatal sinus; one basal, in-running denticle; and one shallow but strong, in-running columellar tooth. Shells and anatomy of juveniles unknown.
</paragraph>
<paragraph blockId="22.[189,421,1732,1758]" box="[189,421,1732,1758]" pageId="22" pageNumber="23">
<heading bold="true" box="[189,421,1732,1758]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis bold="true" box="[189,421,1732,1758]" pageId="22" pageNumber="23">Range and habitat</emphasis>
</heading>
</paragraph>
<paragraph blockId="22.[189,1296,1779,1806]" box="[189,1296,1779,1806]" pageId="22" pageNumber="23">
Submontane forest (
<quantity box="[425,501,1780,1806]" metricMagnitude="2" metricUnit="m" metricValue="9.7" pageId="22" pageNumber="23" unit="m" value="970.0">970 m</quantity>
elevation) in the East Usambara Mountains, northeastern
<collectingCountry box="[1182,1291,1779,1805]" name="Tanzania" pageId="22" pageNumber="23">Tanzania</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="discussion">
<paragraph blockId="22.[189,303,1846,1872]" box="[189,303,1846,1872]" pageId="22" pageNumber="23">
<heading bold="true" box="[189,303,1846,1872]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis bold="true" box="[189,303,1846,1872]" pageId="22" pageNumber="23">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="22.[189,1398,1893,2027]" pageId="22" pageNumber="23">
The complex dentition and detached peristome of
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[783,970,1893,1919]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="frontierarum" status="sp. nov.">
<emphasis box="[783,970,1893,1919]" italics="true" pageId="22" pageNumber="23">
D.
<taxonomicName box="[820,970,1893,1919]" pageId="22" pageNumber="23" rank="species" species="frontierarum" status="sp. nov.">frontierarum</taxonomicName>
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[979,1070,1893,1919]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
are unlike that of any other
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[189,336,1929,1955]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[189,336,1929,1955]" italics="true" pageId="22" pageNumber="23">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[344,452,1929,1955]" pageId="22" pageNumber="23" rank="genus">gen. nov.</taxonomicNameLabel>
species. It is also characteristic for its few whorls, few ribs and sharply pointed apex. Biogeographically a close relationship with other species in and around the East Usambaras would seem likely but there is no strong resemblance.
</paragraph>
</subSubSection>
</treatment>
</document>