290 lines
37 KiB
XML
290 lines
37 KiB
XML
<document id="13F719B4320CB2A2426758987CED9E6C" ID-DOI="10.11646/zootaxa.4020.3.1" ID-GBIF-Dataset="09b3037d-5717-4ed1-bb48-cc0dd5daaa66" ID-ISSN="1175-5326" ID-Zenodo-Dep="236879" ID-ZooBank="9487F2FA-EC49-44F8-9A26-05CFE349CBB0" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461270290296" checkinUser="plazi" docAuthor="Esmaeili-Rineh, Somayeh, Sari, Alireza & Fišer, Cene" docDate="2015" docId="3A1187DA1128FFF8B998FF20FC7189D9" docLanguage="en" docName="zt04020p430.pdf" docOrigin="Zootaxa 4020 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Niphargus darvishi Esmaeili-Rineh, Sari & Fišer, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="418" masterDocId="C628FFA21125FFEAB90FFFB7FFAE885E" masterDocTitle="Making future taxonomy of Niphargus (Crustacea: Amphipoda: Niphargidae) in the Middle East easier: DELTA database of Middle East species with description of four new species from Iran" masterLastPageNumber="430" masterPageNumber="401" pageNumber="414" updateTime="1698611467441" updateUser="plazi">
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<mods:mods id="C6B7F1868B074021BD01D0AF88AF317B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DAED3D8A1F1CA8A21A7EBA8FB1183FE5">
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<mods:title id="6C1C5FA885DE2E3F3FA395FCDE3A1838">Making future taxonomy of Niphargus (Crustacea: Amphipoda: Niphargidae) in the Middle East easier: DELTA database of Middle East species with description of four new species from Iran</mods:title>
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</mods:titleInfo>
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<mods:name id="D0DD5F7AB5E6A88C9081DB4B0C427F3F" type="personal">
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<mods:role id="4AA14C28990030B5CDD3B4D5FC4C7D80">
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<mods:roleTerm id="8647FBD6CBCA5AE6651028516FCF13DF">Author</mods:roleTerm>
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<mods:namePart id="610188DA0344BEE53323B05A56A48698">Esmaeili-Rineh, Somayeh</mods:namePart>
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<mods:name id="A81D0F10A80C1E04F1264C9D8113FDB7" type="personal">
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<mods:role id="F5AB55F4B44048FD619C9D71787512CD">
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<mods:roleTerm id="5F24465AC6390442DBFDB5E31A49838B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="667C44FBA7A38DD30399ED6F72DDA977">Sari, Alireza</mods:namePart>
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</mods:name>
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<mods:name id="8146CD1F3DA413C04E2120EB1DD486F6" type="personal">
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<mods:roleTerm id="F9D3B8693AEE30A956BC3E15E8E02736">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F390ADC49AA25A8EB3393035354990C3">Fišer, Cene</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0DBF512A38B05AF37815BF03E24CEBBC">text</mods:typeOfResource>
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<mods:title id="064E5D6A5785FB53543F84511D222C9D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E0A9D033DB08A5BF930DB2E97433C5FA">
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<mods:date id="E8B782DE9B81691F272BCC7D153EB625">2015</mods:date>
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<mods:detail id="FB13ECD2AF570259CA0FDE4B21F41341" type="volume">
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<mods:number id="AD7AC800C7EDDA4F433E6B96F2CB2BAC">4020</mods:number>
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</mods:detail>
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<mods:detail id="C3CF605CDE58B8AA900069935CCC0F64" type="issue">
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<mods:number id="5A848C41FC4F9F9C4E3AA4DCF19DE1B1">3</mods:number>
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<mods:classification id="283AD143138073C31DBBE328CE862451">journal article</mods:classification>
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<mods:identifier id="8614F800A5404378526A56FB54EED452" type="DOI">10.11646/zootaxa.4020.3.1</mods:identifier>
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<mods:identifier id="CCB2E6F7F4EED56E49CEF9D5B4107391" type="GBIF-Dataset">09b3037d-5717-4ed1-bb48-cc0dd5daaa66</mods:identifier>
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<mods:identifier id="9BBA96FC3875B48C28A25D748D6C93C2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="09FB10C1A154A6DFF565A7CE04220033" type="Zenodo-Dep">236879</mods:identifier>
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<mods:identifier id="0378F4F4422CC145B6044FCC5852CBDD" type="ZooBank">9487F2FA-EC49-44F8-9A26-05CFE349CBB0</mods:identifier>
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</mods:mods>
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<treatment id="3A1187DA1128FFF8B998FF20FC7189D9" ID-DOI="http://doi.org/10.5281/zenodo.6092261" ID-GBIF-Taxon="119638736" ID-Zenodo-Dep="6092261" LSID="urn:lsid:plazi:treatment:3A1187DA1128FFF8B998FF20FC7189D9" httpUri="http://treatment.plazi.org/id/3A1187DA1128FFF8B998FF20FC7189D9" lastPageId="18" lastPageNumber="418" pageId="13" pageNumber="414">
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<subSubSection id="FAA265471128FFE7B998FF20FE8B888D" pageId="13" pageNumber="414" type="nomenclature">
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<paragraph id="B20736CC1128FFE7B998FF20FE4B88EC" blockId="13.[151,485,151,211]" box="[151,485,151,178]" pageId="13" pageNumber="414">
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<heading id="E94F81A01128FFE7B998FF20FE4B88EC" bold="true" box="[151,485,151,178]" fontSize="11" level="1" pageId="13" pageNumber="414" reason="1">
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<emphasis id="80CCEADE1128FFE7B998FF20FE4B88EC" bold="true" box="[151,485,151,178]" pageId="13" pageNumber="414">
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||
<taxonomicName id="75B84D4F1128FFE7B998FF20FE2F88EF" ID-CoL="47FYW" box="[151,385,151,177]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="darvishi" status="sp. nov.">
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<emphasis id="80CCEADE1128FFE7B998FF20FE2F88EF" bold="true" box="[151,385,151,177]" italics="true" pageId="13" pageNumber="414">Niphargus darvishi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="9BFF57A51128FFE7B887FF2FFE4B88EC" box="[392,485,152,178]" pageId="13" pageNumber="414" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="B20736CC1128FFE7B998FF0DFE8B888D" blockId="13.[151,485,151,211]" box="[151,293,186,211]" pageId="13" pageNumber="414">(Figs 10–13)</paragraph>
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</subSubSection>
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||
<subSubSection id="FAA265471128FFE7B998FF4AFAE38964" pageId="13" pageNumber="414" type="materials_examined">
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<paragraph id="B20736CC1128FFE7B998FF4AFAE38964" blockId="13.[151,1437,253,2028]" pageId="13" pageNumber="414">
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<emphasis id="80CCEADE1128FFE7B998FF4AFDFE8948" bold="true" box="[151,592,253,278]" pageId="13" pageNumber="414">
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||
Material examined and
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<typeStatus id="6D03886E1128FFE7B8B7FF4AFE428948" box="[440,492,253,278]" pageId="13" pageNumber="414">type</typeStatus>
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locality.
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</emphasis>
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<typeStatus id="6D03886E1128FFE7BB57FF4AFD6D8948" box="[600,707,253,278]" pageId="13" pageNumber="414" type="holotype">Holotype</typeStatus>
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, male
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<collectionCode id="D4A9AE091128FFE7BA01FF49FCF98948" box="[782,855,254,278]" pageId="13" pageNumber="414">ZUTC</collectionCode>
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Amph. 2322. Dimeh spring (
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<figureCitation id="2A832A491128FFE7BDA8FF4AFB478948" box="[1191,1257,253,278]" captionStart="FIGURE 1" captionStartId="1.[151,250,1908,1930]" captionTargetBox="[170,1409,816,1876]" captionTargetId="figure@1.[164,1423,804,1887]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of the genus Niphargus in Iran and Middle East." httpUri="https://zenodo.org/record/236880/files/figure.png" pageId="13" pageNumber="414">Fig. 1</figureCitation>
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), Kohrang city, Chahar-mahal va Bakhteyari Province,
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<collectingCountry id="CAAF765C1128FFE7BB5EFE95FDD08967" box="[593,638,290,313]" name="Iran" pageId="13" pageNumber="414">Iran</collectingCountry>
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(N 32° 30′, E 50°13′). Five male
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<typeStatus id="6D03886E1128FFE7BAF8FE95FBCB8964" box="[1015,1125,290,314]" pageId="13" pageNumber="414" type="paratype">paratypes</typeStatus>
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,
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<collectionCode id="D4A9AE091128FFE7BD7FFE96FB178967" box="[1136,1209,289,313]" pageId="13" pageNumber="414">ZUTC</collectionCode>
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Amph.2347.
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</paragraph>
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</subSubSection>
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<subSubSection id="FAA265471128FFE7B9C8FEF2FD9A8994" pageId="13" pageNumber="414" type="diagnosis">
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<paragraph id="B20736CC1128FFE7B9C8FEF2FD9A8994" blockId="13.[151,1437,253,2028]" pageId="13" pageNumber="414">
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<emphasis id="80CCEADE1128FFE7B9C8FEF2FEEF8900" bold="true" box="[199,321,325,350]" pageId="13" pageNumber="414">Diagnosis.</emphasis>
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Maxilla I with pluritooth robust setae on outer lobe. Propods of gnathopods large, broader than long. Bases of pereopods V–VII mid-sized (neither narrow nor broad). Dactyli of pereopods III–VII with more than two robust setae along inner margin. Urosomite I with two robust straight postero-distal setae. Epimeral plate III distinctly produced disto-posteriorly.
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</paragraph>
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</subSubSection>
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<subSubSection id="FAA265471128FFFBB9C8FE62FC758F64" lastPageId="17" lastPageNumber="418" pageId="13" pageNumber="414" type="description">
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||
<paragraph id="B20736CC1128FFE7B9C8FE62FAFF89B0" blockId="13.[151,1437,253,2028]" box="[199,1361,469,494]" pageId="13" pageNumber="414">
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<emphasis id="80CCEADE1128FFE7B9C8FE62FE4A89B0" bold="true" box="[199,484,469,494]" pageId="13" pageNumber="414">
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Description of
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<typeStatus id="6D03886E1128FFE7B879FE62FE7189B0" box="[374,479,469,494]" pageId="13" pageNumber="414" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Body length
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<quantity id="75409B291128FFE7BB73FE61FD6089B3" box="[636,718,470,494]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="414" unit="mm" value="13.0">13 mm</quantity>
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. Body strong and stout. Head length 11% of body length.
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</paragraph>
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<paragraph id="B20736CC1128FFE7B9C8FE4EFD048A20" blockId="13.[151,1437,253,2028]" pageId="13" pageNumber="414">Antennae I (fig. 10A). Antenna I 0.51 of body length. Peduncular articles 1–3 progressively shorter; length of peduncular article 3 less than one half of peduncular article 2; main flagellum with 29 articles (most of which with short setae); accessory flagellum bi-articulated and reaching beyond two-third of article 1 of main flagellum; second article with two simple setae (fig. 10A).</paragraph>
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<paragraph id="B20736CC1128FFE7B9C8FD3DFB978AB4" blockId="13.[151,1437,253,2028]" pageId="13" pageNumber="414">Antennae II (fig. 10B). Antenna II as long as 0.48 antenna I. Peduncular article 5 slightly shorter than article 4, ratio between articles 4:5 equal to 1.25: 1, articles 4 and 5 with five and eight groups of simple setae, respectively. Flagellum with 10 articles, approximately 0.58 of length of peduncle articles 4 + 5.</paragraph>
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<paragraph id="B20736CC1128FFE7B9C8FD42FDFB8B50" blockId="13.[151,1437,253,2028]" box="[199,597,757,782]" pageId="13" pageNumber="414">Labium (fig. 11B) with inner lobes.</paragraph>
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<paragraph id="B20736CC1128FFE7B9C8FCAEFDFD8B27" blockId="13.[151,1437,253,2028]" pageId="13" pageNumber="414">Maxilla I (fig. 10C & D). Inner plate with two long simple setae, outer plate with seven robust setae of which two with three, two with two, one with five, one with four and one without lateral projections; palp bi-articulated, long, with four long distal simple setae.</paragraph>
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<paragraph id="B20736CC1128FFE7B9C8FC32FA388BC3" blockId="13.[151,1437,253,2028]" box="[199,1430,901,926]" pageId="13" pageNumber="414">Maxilla II (fig. 10H). Both plates with numerous long distal simple setae, inner lobe with one lateral simple seta.</paragraph>
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<paragraph id="B20736CC1128FFE7B9C8FC1EFC6F8C57" blockId="13.[151,1437,253,2028]" pageId="13" pageNumber="414">Mandibular palp (fig. 10G). Ratio of mandibular palp articles 1: 2: 3 equal to 1: 1.51: 1.88, no setae on first article; second article with six medial and two disto-lateral long simple setae. Distal article with one group of four A-setae, five groups of B-setae, no C-setae, 25 D-setae and five E-setae.</paragraph>
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<paragraph id="B20736CC1128FFE7B9C8FBA2FBED8C70" blockId="13.[151,1437,253,2028]" box="[199,1091,1045,1070]" pageId="13" pageNumber="414">Left mandible (fig. 10F). Incisor with five teeth, lacinia mobilis with four teeth.</paragraph>
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<paragraph id="B20736CC1128FFE7B9C8FB8EFD138C2B" blockId="13.[151,1437,253,2028]" pageId="13" pageNumber="414">Right mandible (fig. 10E). Incisor with four teeth, lacinia mobilis pluritooth, seven setae with lateral projections between lacinia and triturative molar.</paragraph>
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<paragraph id="B20736CC1128FFE7B9C8FB37FAF08D5D" blockId="13.[151,1437,253,2028]" pageId="13" pageNumber="414">Maxilliped (fig. 11A). Inner plate short; distally three robust setae intermixed with three simple setae, subdistally one simple long lateral seta; outer plate exceeding half of posterior margin of palp article 2, with 11 marginal robust setae and six distal simple setae; palp article 3 at outer margin with one proximal, distal and medial group of long simple setae; distal article with five simple setae along outer margin, nail shorter than pedestal.</paragraph>
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<paragraph id="B20736CC1128FFE7B9C8FAB9FF4B8ED7" blockId="13.[151,1437,253,2028]" pageId="13" pageNumber="414">Gnathopod I (fig. 11C). Coxa of gnathopod I smaller than gnathopod II. Coxa I trapezoid, broader than long, ventral and anterior margins with one and six simple setae, respectively. Basis with long setae on anterior and posterior margins; ischium and merus with one posterior group of setae. Carpus with one group of setae anterodistally and many subdistal setae along anterior margin. Posterior part of carpus with setae on proximal bulbous and along postero-medial margin. Carpus 0.67 of basis length and 0.86 of propodus length. Propodus large, broader than long, posterior margin slightly convex. Anterior margin of propodus with 11 simple setae in two groups in addition to antero-distal group with six setae; palm slightly convex, with one strong long palmar robust seta, one short supporting robust seta on inner surface and two short robust setae with lateral projections on outer surface. In addition, two setae under supporting robust seta in palmar corner. Dactylus not reaching posterior margin of propodus, outer and inner margins each with a row of four simple setae, nail short, length 0.23 of total dactylus length.</paragraph>
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<paragraph id="B20736CC1128FFE7B9C8F923FCB88FB2" blockId="13.[151,1437,253,2028]" pageId="13" pageNumber="414">Gnathopod II (fig. 11D). Coxa II rectangular, with 13 simple setae along antero-ventral margin. Posterior margin of basis with two groups of setae and two single setae, anterior margin of basis with single seta. Posterior margin of ischium and merus with one posterior group of setae. Carpus 0.49 of basis length and 0.84 propodus length; carpus with single antero-distal group of setae, with setae on posterior-proximal bulbous and with a row of setae along postero-medial margin. Propodus in gnathopod II larger than gnathopod I, broader than long, anterior margin with two simple setae in one group in addition to five antero-distal setae. Medial surface of propodus with few setae. Palm slightly convex, with one strong palmar robust seta, one short supporting robust seta on inner surface and two short robust setae with lateral projections on outer surface. Dactylus not reaching posterior margin of propodus, outer margin of dactylus with two groups of two and three simple setae, and with seven simple setae on inner margin, nail length 0.16 of total dactylus length.</paragraph>
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<paragraph id="B20736CC112BFFE4B9C8FF2FFADE891E" blockId="14.[151,1436,151,320]" pageId="14" pageNumber="415">
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Coxae III–VII (fig. 12A–F). Length to width ratio in coxa
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<date id="C606100C112BFFE4BA6FFF2FFC3B88EE" box="[864,917,152,176]" pageId="14" pageNumber="415">III 2</date>
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: 1, antero-ventral margins of coxa III with 12 simple setae and one robust seta. Length to width ratio in coxa IV 1.12: 1, ventro- posterior margins of coxa IV with eight simple setae, posterior concavity shallow and approximately 0.07 of coxa width (fig. 12A–B). Coxa V with posteriorly developed lobe, with four simple setae in anterior lobe and four setae posteriorly. Coxa VI with anteriorly developed lobe, and single seta posteriorly. Coxa VII with one seta in posterior margin (fig. 12C–F).
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</paragraph>
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<paragraph id="B20736CC112BFFE4B998F823FB0A8F95" blockId="14.[151,1436,1940,1995]" pageId="14" pageNumber="415">
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<emphasis id="80CCEADE112BFFE4B998F823FE8E8FF7" bold="true" box="[151,288,1940,1962]" pageId="14" pageNumber="415">FIGURE 10.</emphasis>
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<taxonomicName id="75B84D4F112BFFE4B828F823FE428FF4" box="[295,492,1940,1962]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="darvishi" status="sp. nov.">
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<emphasis id="80CCEADE112BFFE4B828F823FE428FF4" box="[295,492,1940,1962]" italics="true" pageId="14" pageNumber="415">Niphargus darvishi</emphasis>
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</taxonomicName>
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,
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<emphasis id="80CCEADE112BFFE4B8F6F822FDE48FF4" bold="true" box="[505,586,1941,1962]" pageId="14" pageNumber="415">
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||
<taxonomicNameLabel id="9BFF57A5112BFFE4B8F6F822FDE48FF4" box="[505,586,1941,1962]" pageId="14" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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, Dimeh Spring, male
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||
<quantity id="75409B29112BFFE4BA21F823FCD88FF4" box="[814,886,1940,1962]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="415" unit="mm" value="13.0">13 mm</quantity>
|
||
(
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||
<typeStatus id="6D03886E112BFFE4BA8BF823FC4C8FF4" box="[900,994,1940,1962]" pageId="14" pageNumber="415" type="holotype">holotype</typeStatus>
|
||
). A, A1; B, A2; C–D,
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<collectingCountry id="CAAF765C112BFFE4BDC6F822FB5C8FF4" box="[1225,1266,1941,1962]" name="Mexico" pageId="14" pageNumber="415">MX</collectingCountry>
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||
1; E, LMND; F, RMND; G, MNDP; H,
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<collectingCountry id="CAAF765C112BFFE4B88EF801FE048F95" box="[385,426,1974,1995]" name="Mexico" pageId="14" pageNumber="415">MX</collectingCountry>
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||
2. Scale bars: 1=
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<quantity id="75409B29112BFFE4BB6FF802FD178F95" box="[608,697,1973,1995]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="415" unit="mm" value="0.25">0.25 mm</quantity>
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||
(E–F); 2=
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<quantity id="75409B29112BFFE4BA27F802FCDB8F95" box="[808,885,1973,1995]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="415" unit="mm" value="0.5">0.5 mm</quantity>
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(C–D, G–H); 3=
|
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<quantity id="75409B29112BFFE4BD2BF802FBF78F95" box="[1060,1113,1973,1995]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="415" unit="mm" value="1.0">1mm</quantity>
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(A–B).
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||
</paragraph>
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<paragraph id="B20736CC112AFFE5B998FA72FD9E8DA2" blockId="15.[151,1436,1477,1532]" pageId="15" pageNumber="416">
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||
<emphasis id="80CCEADE112AFFE5B998FA72FEB18D84" bold="true" box="[151,287,1477,1499]" pageId="15" pageNumber="416">FIGURE 11.</emphasis>
|
||
<taxonomicName id="75B84D4F112AFFE5B829FA72FE458D85" box="[294,491,1477,1499]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="darvishi" status="sp. nov.">
|
||
<emphasis id="80CCEADE112AFFE5B829FA72FE458D85" box="[294,491,1477,1499]" italics="true" pageId="15" pageNumber="416">Niphargus darvishi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="80CCEADE112AFFE5B8F6FA72FDE48D84" bold="true" box="[505,586,1477,1498]" pageId="15" pageNumber="416">
|
||
<taxonomicNameLabel id="9BFF57A5112AFFE5B8F6FA72FDE48D84" box="[505,586,1477,1498]" pageId="15" pageNumber="416" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Dimeh Spring, male
|
||
<quantity id="75409B29112AFFE5BA20FA72FCD98D85" box="[815,887,1477,1499]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="416" unit="mm" value="13.0">13 mm</quantity>
|
||
(
|
||
<typeStatus id="6D03886E112AFFE5BA89FA72FC4B8D85" box="[902,997,1477,1499]" pageId="15" pageNumber="416" type="holotype">holotype</typeStatus>
|
||
). A, MXP; B, L; C, GN 1; D, GN 2. Scale bars: 1=
|
||
<quantity id="75409B29112AFFE5B9FFFA51FE928DA2" box="[240,316,1510,1532]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="416" unit="mm" value="0.5">0.5 mm</quantity>
|
||
(A–B); 2=
|
||
<quantity id="75409B29112AFFE5B8BEFA51FE4B8DA2" box="[433,485,1510,1532]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="416" unit="mm" value="1.0">1mm</quantity>
|
||
(C–D).
|
||
</paragraph>
|
||
<paragraph id="B20736CC112AFFE5B9C8F99CFB4B8ED2" blockId="15.[151,1437,1579,2036]" pageId="15" pageNumber="416">Pereopods III–VII (fig. 12A–F). Pereopod III longer than IV with length ratio of 1.06: 1 (fig. 8A–B). Dactyli of pereopods III–IV short, simple seta along outer margin and with two robust setae and one simple setae at inner margin. Length of dactylus in pereopod IV 0.32 of propodus. Nail shorter than pedestal (fig. 12B).</paragraph>
|
||
<paragraph id="B20736CC112AFFE5B9C8F920FC028FF2" blockId="15.[151,1437,1579,2036]" pageId="15" pageNumber="416">
|
||
Pereopods V–VII. Length ratios of perepods V: VI: VII equal to 1: 1.48: 1.51. Pereopod VII 0.52 of body length. Bases in pereopods V–VII with respective nine, six and six gropus of anterior setae and with 12, nine, and eight short single setae along posterior margin (fig. 12C–F). Postero-ventral lobe of ischium in pereopods V–VII developed. Ischium, merus and carpus in pereopods V–VII with several groups of robust and simple setae along anterior and posterior margin. Propodus of pereopod VII slightly longer than these in V and VI. Dactyli of pereopods V–VII with three (on
|
||
<typeStatus id="6D03886E112AFFE5BB05F8FBFDC08F3A" box="[522,622,1868,1892]" pageId="15" pageNumber="416" type="paratype">paratype</typeStatus>
|
||
, appendage broken in
|
||
<typeStatus id="6D03886E112AFFE5BA62F8FCFC7D8F3A" box="[877,979,1867,1892]" pageId="15" pageNumber="416" type="holotype">holotype</typeStatus>
|
||
), three and four robust setae along inner margin, respectively. Outer margin with single seta in pereopods V–VI and with two such setae in pereopod VII. Nail length of pereopod VII 0.28 of total dactylus length (fig. 12C–E).
|
||
</paragraph>
|
||
<paragraph id="B20736CC112AFFE5B9C8F800FE598F91" blockId="15.[151,1437,1579,2036]" box="[199,503,1975,2000]" pageId="15" pageNumber="416">Pereonites I–VII. No setae.</paragraph>
|
||
<paragraph id="B20736CC112AFFE5B9C8F86CFC2E8FAA" blockId="15.[151,1437,1579,2036]" box="[199,896,2011,2036]" pageId="15" pageNumber="416">Pleonites I–III. With one simple seta at postero-dorsal margin.</paragraph>
|
||
<caption id="E6C766441135FFFAB998FA86FDC68D38" httpUri="https://zenodo.org/record/236883/files/figure.png" pageId="16" pageNumber="417" targetBox="[151,1436,195,1305]" targetPageId="16">
|
||
<paragraph id="B20736CC1135FFFAB998FA86FDC68D38" blockId="16.[151,1435,1329,1382]" pageId="16" pageNumber="417">
|
||
<emphasis id="80CCEADE1135FFFAB998FA86FE8F8D18" bold="true" box="[151,289,1329,1351]" pageId="16" pageNumber="417">FIGURE 12.</emphasis>
|
||
<taxonomicName id="75B84D4F1135FFFAB826FA86FE418D19" box="[297,495,1329,1351]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="darvishi" status="sp. nov.">
|
||
<emphasis id="80CCEADE1135FFFAB826FA86FE418D19" box="[297,495,1329,1351]" italics="true" pageId="16" pageNumber="417">Niphargus darvishi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="9BFF57A51135FFFAB8F2FA85FDFE8D19" box="[509,592,1330,1351]" pageId="16" pageNumber="417" rank="species">
|
||
<emphasis id="80CCEADE1135FFFAB8F2FA85FDFE8D19" bold="true" box="[509,592,1330,1351]" pageId="16" pageNumber="417">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, Dimeh Spring, male 13 mm (holotype). A, P 3; B, P 4; C, P 5; D, distal part of P5(paratype); E, P 6; F, P 7. Scale bars: 1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B20736CC1135FFFAB9C8FA24FD758DAA" blockId="16.[151,1437,1427,2028]" pageId="16" pageNumber="417">Pleopods I–III (fig. 13A–C). Peduncle of pleopods I–III with two hooked retinacula at distal part of inner margin; peduncle of pleopods I–II each with one simple seta. Rami of pleopod I broken, Rami of pleopod II–III with 13–15 articles and 12–14 articles, respectively.</paragraph>
|
||
<paragraph id="B20736CC1135FFFAB9C8FA49FEC48EF9" blockId="16.[151,1437,1427,2028]" pageId="16" pageNumber="417">Epimeral plates I–III (fig. 13D). Epimeral plates with postero-ventral corner angular; posterior and ventral margins slightly convex. Epimeral plate III with distinctly produced posterodistal corner, posterior and ventral margins concave and convex, respectively. All plates with one long robust seta at posterior margin; with two, three and three simple setae at posterior margin, respectively; ventral margin of plates II and III with three and one robust setae, respectively.</paragraph>
|
||
<paragraph id="B20736CC1135FFFAB9C8F904FCC38EB1" blockId="16.[151,1437,1427,2028]" pageId="16" pageNumber="417">Urosomites I–II. Postero-dorsally each with two robust setae. Dorsally with one and two simple setae, respectively. With one strong robust seta, at the base of uropod I.</paragraph>
|
||
<paragraph id="B20736CC1135FFFBB9C8F94CFEF4888A" blockId="16.[151,1437,1427,2028]" lastBlockId="17.[151,1436,151,285]" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">Uropods I–III (fig. 13E–F). Peduncle in uropod I with five and three large robust setae in dorso-lateral and dorso-medial margins, respectively. Ratio of inner to outer ramus length 1.15: 1.05. Inner ramus with five groups of robust setae and five robust setae distally. Outer ramus with four groups of robust and simple setae laterally and five single robust setae distally (fig. 13E). Inner ramus in uropod II slightly longer than outer, both rami with lateral and distal long robust setae (fig. 13E). Uropod III with no sexual dimorphism, almost 0.24 of body length. Peduncle of uropod III with five robust setae distally, proximal article of outer ramus with five and six groups of robust setae along outer and inner margin, respectively (fig. 13F). Distal to proximal article ratio of outer ramus 1: 12, distal article with two simple lateral setae and one distal simple seta. Inner ramus of uropod III short, with one distal robust seta.</paragraph>
|
||
<paragraph id="B20736CC1134FFFBB9C8FF68FDD08943" blockId="17.[151,1436,151,285]" pageId="17" pageNumber="418">Telson (fig. 13G) as long as broad, deeply cleft, each lobe with three distal long robust setae and one long simple seta, with two simple setae laterally.</paragraph>
|
||
<paragraph id="B20736CC1134FFFBB998F8B5FC758F64" blockId="17.[151,1436,1794,1850]" pageId="17" pageNumber="418">
|
||
<emphasis id="80CCEADE1134FFFBB998F8B5FE8E8F49" bold="true" box="[151,288,1794,1816]" pageId="17" pageNumber="418">FIGURE 13.</emphasis>
|
||
<taxonomicName id="75B84D4F1134FFFBB828F8B5FE458F46" box="[295,491,1794,1816]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="darvishi" status="sp. nov.">
|
||
<emphasis id="80CCEADE1134FFFBB828F8B5FE458F46" box="[295,491,1794,1816]" italics="true" pageId="17" pageNumber="418">Niphargus darvishi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="80CCEADE1134FFFBB8F6F8B4FDE48F46" bold="true" box="[505,586,1795,1816]" pageId="17" pageNumber="418">
|
||
<taxonomicNameLabel id="9BFF57A51134FFFBB8F6F8B4FDE48F46" box="[505,586,1795,1816]" pageId="17" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Dimeh Spring, male
|
||
<quantity id="75409B291134FFFBBA21F8B5FCDB8F46" box="[814,885,1794,1816]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="17" pageNumber="418" unit="mm" value="13.0">13 mm</quantity>
|
||
(
|
||
<typeStatus id="6D03886E1134FFFBBA8BF8B5FC4C8F46" box="[900,994,1794,1816]" pageId="17" pageNumber="418" type="holotype">holotype</typeStatus>
|
||
). A, PL 1; B, PL 2; C, PL 3; D, EP 1–3; E, USM; F, U 3; G, T. Scale bars: 1=
|
||
<quantity id="75409B291134FFFBB8FDF893FD918F64" box="[498,575,1828,1850]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="418" unit="mm" value="0.5">0.5 mm</quantity>
|
||
(D, G); 2=
|
||
<quantity id="75409B291134FFFBBBB9F893FD458F64" box="[694,747,1828,1850]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="418" unit="mm" value="1.0">1mm</quantity>
|
||
(A–C, E); 3=
|
||
<quantity id="75409B291134FFFBBA74F893FC008F64" box="[891,942,1828,1850]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="418" unit="mm" value="2.0">2mm</quantity>
|
||
(F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FAA265471134FFFBB9C8F8DEFC7C8FFB" pageId="17" pageNumber="418" type="etymology">
|
||
<paragraph id="B20736CC1134FFFBB9C8F8DEFC7C8FFB" blockId="17.[151,1436,1897,2030]" pageId="17" pageNumber="418">
|
||
<emphasis id="80CCEADE1134FFFBB9C8F8DEFEE08FDC" bold="true" box="[199,334,1897,1922]" pageId="17" pageNumber="418">Etymology.</emphasis>
|
||
The species is named in honor of Professor Jamshid Darvish, Ferdowsi University of Mashhad who is one of the leading experts in animal biosystematics studies in
|
||
<collectingCountry id="CAAF765C1134FFFBBA91F839FC608FFB" box="[926,974,1934,1957]" name="Iran" pageId="17" pageNumber="418">Iran</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FAA265471134FFF8B9C8F806FC7189D9" lastPageId="18" lastPageNumber="419" pageId="17" pageNumber="418" type="discussion">
|
||
<paragraph id="B20736CC1134FFF8B9C8F806FC7189D9" blockId="17.[151,1436,1897,2030]" lastBlockId="18.[151,1437,151,392]" lastPageId="18" lastPageNumber="419" pageId="17" pageNumber="418">
|
||
<emphasis id="80CCEADE1134FFFBB9C8F806FE718F94" bold="true" box="[199,479,1969,1994]" pageId="17" pageNumber="418">Remarks and affinities.</emphasis>
|
||
<taxonomicName id="75B84D4F1134FFFBB8E9F806FD118F97" box="[486,703,1969,1993]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="darvishi">
|
||
<emphasis id="80CCEADE1134FFFBB8E9F806FD118F97" box="[486,703,1969,1993]" italics="true" pageId="17" pageNumber="418">Niphargus darvishi</emphasis>
|
||
</taxonomicName>
|
||
in many aspects (pectinate dactyli, produced epimeral plates and broad gnathopods) resembles to
|
||
<taxonomicName id="75B84D4F1134FFFBBB08F861FD0F8FB3" box="[519,673,2005,2029]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="valachicus">
|
||
<emphasis id="80CCEADE1134FFFBBB08F861FD0F8FB3" box="[519,673,2005,2029]" italics="true" pageId="17" pageNumber="418">N. valachicus</emphasis>
|
||
</taxonomicName>
|
||
; however, the two species differ from each other by ornamentation of urosomite I (
|
||
<taxonomicName id="75B84D4F1137FFF8B85EFF2EFE4388EE" box="[337,493,152,176]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="valachicus">
|
||
<emphasis id="80CCEADE1137FFF8B85EFF2EFE4388EE" box="[337,493,152,176]" italics="true" pageId="18" pageNumber="419">N. valachicus</emphasis>
|
||
</taxonomicName>
|
||
with single strong and curved seta on tiny protuberance) and male uropod I (
|
||
<taxonomicName id="75B84D4F1137FFF8BC8DFF2EFEA1888A" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="valachicus">
|
||
<emphasis id="80CCEADE1137FFF8BC8DFF2EFEA1888A" italics="true" pageId="18" pageNumber="419">N. valachicus</emphasis>
|
||
</taxonomicName>
|
||
with finger-like protuberance distally on peduncle). Some similarities (pectinate dactyli, broad gnathopods) are found in species from West Balkan (
|
||
<taxonomicName id="75B84D4F1137FFF8BBEBFF56FC9088A9" box="[740,830,224,248]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="rejici">
|
||
<emphasis id="80CCEADE1137FFF8BBEBFF56FC9088A9" box="[740,830,224,248]" italics="true" pageId="18" pageNumber="419">N. rejici</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="75B84D4F1137FFF8BA43FF56FC7F88A6" box="[844,977,224,248]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="jadranko">
|
||
<emphasis id="80CCEADE1137FFF8BA43FF56FC7F88A6" box="[844,977,224,248]" italics="true" pageId="18" pageNumber="419">N. jadranko</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="75B84D4F1137FFF8BAD1FF56FB0488A6" box="[990,1194,225,248]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="pectencoronate">
|
||
<emphasis id="80CCEADE1137FFF8BAD1FF56FB0488A6" box="[990,1194,225,248]" italics="true" pageId="18" pageNumber="419">N. pectencoronate</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="D6294B3D1137FFF8BDEEFF68FF7E8942" author="Sket" pageId="18" pageNumber="419" refString="Sket, B. & Karaman, G. (1990) Niphargus rejici (Amphipoda), its relatives in the Adriatic islands, and its possible relations to S. W. Asian taxa. Stygologia, 5,153 - 172." type="book" year="1990">Sket & Karaman 1990</bibRefCitation>
|
||
) and from Middle East (
|
||
<taxonomicName id="75B84D4F1137FFF8B8FCFEB2FDD98942" box="[499,631,260,284]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="nadarini">
|
||
<emphasis id="80CCEADE1137FFF8B8FCFEB2FDD98942" box="[499,631,260,284]" italics="true" pageId="18" pageNumber="419">N. nadarini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="80CCEADE1137FFF8BB87FEB2FC368942" box="[648,920,260,284]" italics="true" pageId="18" pageNumber="419">
|
||
<taxonomicName id="75B84D4F1137FFF8BB87FEB2FD558942" box="[648,763,261,284]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="sertaci">N. sertaci</taxonomicName>
|
||
,
|
||
<taxonomicName id="75B84D4F1137FFF8BA06FEB2FC368942" box="[777,920,260,284]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="khayyami">N. khayyami</taxonomicName>
|
||
</emphasis>
|
||
see
|
||
<bibRefCitation id="D6294B3D1137FFF8BADFFEB3FBF88942" author="Alouf" box="[976,1110,260,285]" pageId="18" pageNumber="419" refString="Alouf, N. (1972) Description d'un nouvel amphipode Gammaride´des eaux souterraines du Liban, Niphargus nadarini n. sp .. Annales de Speleologie, 27, 547 - 558." type="journal article" year="1972">Alouf 1972</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D6294B3D1137FFF8BD6AFEB2FABF8943" author="Karaman" box="[1125,1297,260,285]" pageId="18" pageNumber="419" refString="Karaman, G. S. (1986) First discovery of genus Niphargus Sch. in Iraq, Israel and adjacent regions, with description of N. itus, new Species (Fam. Niphargidae) (contribution to the knowledge of the Amphipoda 153). Poljoprivreda i sumarstvo, 32, 13 - 36." type="journal article" year="1986">Karaman 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D6294B3D1137FFF8BC10FEB3FF4E891E" author="Fiser" pageId="18" pageNumber="419" refString="Fiser, C., Camur-Elipek, B. & Ozbek, M. (2009 b) The subterranean genus Niphargus (Crustacea, Amphipoda) in the Middle East: a faunistic overview with descriptions of two new species. Zoologischer Anzeiger, 248, 137 - 150. http: // dx. doi. org / 10.1016 / j. jcz. 2009.03.003" type="journal article" year="2009" yearSuffix="b">
|
||
Fišer
|
||
<emphasis id="80CCEADE1137FFF8BC6EFEB2FA3B8942" box="[1377,1429,260,284]" italics="true" pageId="18" pageNumber="419">et al</emphasis>
|
||
. 2009b
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D6294B3D1137FFF8B9E2FE90FE40891E" author="Hekmatara" box="[237,494,295,320]" pageId="18" pageNumber="419" refString="Hekmatara, M., Zaksek, V., Heidari, M. B. & Fiser, C. (2013) Two new species of Niphargus (Crustacea: Amphipoda) from Iran. Journal of Natural History, 47, 1421 - 1449. http: // dx. doi. org / 10.1080 / 00222933.2012.743616" type="journal article" year="2013">
|
||
Hekmatara
|
||
<emphasis id="80CCEADE1137FFF8B87FFE9EFE0C891E" box="[368,418,296,320]" italics="true" pageId="18" pageNumber="419">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
). Moreover, Middle East species share with
|
||
<taxonomicName id="75B84D4F1137FFF8BAEAFE9EFBCD891E" box="[997,1123,296,320]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="darvishi">
|
||
<emphasis id="80CCEADE1137FFF8BAEAFE9EFBCD891E" box="[997,1123,296,320]" italics="true" pageId="18" pageNumber="419">N. darvishi</emphasis>
|
||
</taxonomicName>
|
||
another trait, i.e. pluritooth robust setae on outer lobe of maxilla I. None of these species, however, has disto-posteriorly produced epimeral plates or narrow bases of pereopods V–VII as herein described
|
||
<taxonomicName id="75B84D4F1137FFF8BA54FEC6FC7689D6" box="[859,984,368,392]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="419" phylum="Arthropoda" rank="species" species="darvishi">
|
||
<emphasis id="80CCEADE1137FFF8BA54FEC6FC7689D6" box="[859,984,368,392]" italics="true" pageId="18" pageNumber="419">N. darvishi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |