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<document ID-DOI="10.11646/zootaxa.5264.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836798" approvalRequired="1" approvalRequired_for_textStreams="1" checkinTime="1681728995984" checkinUser="plazi" docAuthor="Heppner, John B. &amp; Bae, Yang-Seop" docDate="2023" docId="3A0C87F6DD1EA71851D9FA1DFBD7FC8B" docLanguage="en" docName="zootaxa.5264.3.5.pdf" docOrigin="Zootaxa 5264 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vangatinea panghapha Heppner &amp; Bae 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="376" masterDocId="C635FF8EDD18A71F514EFFB4FFD3FF9E" masterDocTitle="New species of colorful tineids from Cambodia, Thailand, and Vietnam in the new genus Vangatinea (Lepidoptera: Tineidae: Erechthiinae)" masterLastPageNumber="380" masterPageNumber="369" pageNumber="375" updateTime="1681729545978" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species of colorful tineids from Cambodia, Thailand, and Vietnam in the new genus Vangatinea (Lepidoptera: Tineidae: Erechthiinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heppner, John B.</mods:namePart>
<mods:affiliation>McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA. &amp; jheppner @ flmnh. ufl. edu</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation>Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; io ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea. &amp; baeys @ inu. ac. kr; https: // orcid. org / 0000 - 0001 - 7356 - 5633</mods:affiliation>
<mods:nameIdentifier type="email">baeys@inu.ac.kr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5264</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>369</mods:start>
<mods:end>380</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.3.5</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5264.3.5</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7836798</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7836760" ID-Zenodo-Dep="7836760" LSID="urn:lsid:plazi:treatment:3A0C87F6DD1EA71851D9FA1DFBD7FC8B" httpUri="http://treatment.plazi.org/id/3A0C87F6DD1EA71851D9FA1DFBD7FC8B" lastPageId="7" lastPageNumber="376" pageId="6" pageNumber="375">
<subSubSection box="[151,745,1449,1476]" pageId="6" pageNumber="375" type="nomenclature">
<paragraph blockId="6.[151,974,1449,1548]" box="[151,745,1449,1476]" pageId="6" pageNumber="375">
<heading bold="true" box="[151,745,1449,1476]" fontSize="11" level="1" pageId="6" pageNumber="375" reason="1">
<emphasis bold="true" box="[151,745,1449,1476]" pageId="6" pageNumber="375">
<taxonomicName authority="Heppner &amp; Bae" authorityName="Heppner &amp; Bae" authorityYear="2023" box="[151,638,1449,1476]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="panghapha" status="sp. nov.">
<emphasis bold="true" box="[151,432,1449,1476]" italics="true" pageId="6" pageNumber="375">Vangatinea panghapha</emphasis>
Heppner &amp; Bae
</taxonomicName>
,
<taxonomicNameLabel box="[651,745,1449,1475]" pageId="6" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="375" type="description">
<paragraph blockId="6.[151,974,1449,1548]" box="[151,359,1485,1512]" pageId="6" pageNumber="375">
(
<figureCitation box="[159,239,1485,1511]" captionStart="FIGURES 35" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[199,1398,1421,1873]" captionTargetId="figure-539@4.[190,1398,1412,1875]" captionTargetPageId="4" captionText="FIGURES 35. Vangatinea species: 3) V. sontraensis sp. nov. (Vietnam: Danang), male holotpe (11.4mm wingspan) (MGCL); 4) V. panghapha sp. nov. (Thailand: Mai Hong Son), male holotype (13.5mm wingspan) (MGCL); 5) V. cambodiensis sp. nov. (Cambodia: Pursat), female holotype (16mm wingspan) (INUC)." figureDoi="http://doi.org/10.5281/zenodo.7836804" httpUri="https://zenodo.org/record/7836804/files/figure.png" pageId="6" pageNumber="375">Figs. 4</figureCitation>
,
<figureCitation box="[251,267,1486,1512]" captionStart="FIGURES 68" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[205,1382,181,1920]" captionTargetId="figure-17@5.[205,1382,181,1920]" captionTargetPageId="5" captionText="FIGURES 68. Head views of Vangatinea species (lateral and (a) dorsal views): 6) V. sontraensis sp. nov. (holotype male); 7) V. panghapha sp. nov. (holotype male); 8) V. cambodiensis sp. nov. (holotype female)." figureDoi="http://doi.org/10.5281/zenodo.7836810" httpUri="https://zenodo.org/record/7836810/files/figure.png" pageId="6" pageNumber="375">7</figureCitation>
,
<figureCitation box="[279,309,1485,1511]" captionStart="FIGURE 10" captionStartId="7.[151,250,1645,1669]" captionTargetBox="[155,1436,814,1621]" captionTargetId="figure-283@7.[151,1437,813,1621]" captionTargetPageId="7" captionText="FIGURE 10. Vangatinea panghapha sp. nov., holotype male genitalia (Thailand: Mai Hong Son), with details, a) aedeagus, b) juxta-anellus, and c) pre-genital segment of abdomen (gen. slide JBH-3197) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.7836818" httpUri="https://zenodo.org/record/7836818/files/figure.png" pageId="6" pageNumber="375">10</figureCitation>
,
<figureCitation box="[321,350,1485,1511]" captionStart="FIGURE 12" captionStartId="9.[151,250,1934,1958]" captionTargetBox="[311,1277,930,1911]" captionTargetId="figure-367@9.[311,1277,930,1911]" captionTargetPageId="9" captionText="FIGURE 12. Southeast Asia distribution map of Vangatinea species: V. cambodiensis sp. nov. (white square, Cambodia), V. panghapha sp. nov. (black star, Thailand), and V. sontraensis sp. nov. (black square, Vietnam) (base map courtesy of J. Bartholomew &amp; Son Maps, Edinburgh, Scotland; Times Atlas, 1988. London, UK)." figureDoi="http://doi.org/10.5281/zenodo.7836822" httpUri="https://zenodo.org/record/7836822/files/figure.png" pageId="6" pageNumber="375">12</figureCitation>
)
</paragraph>
<paragraph blockId="6.[151,974,1449,1548]" box="[151,974,1521,1548]" pageId="6" pageNumber="375">
<uri box="[151,974,1521,1548]" pageId="6" pageNumber="375">
urn:lsid:zoobank.org:act:
<uuid box="[428,974,1521,1548]" pageId="6" pageNumber="375">14B2A551-BD04-417A-93A5-3340CF1A9CBF</uuid>
</uri>
</paragraph>
<paragraph blockId="6.[151,1437,1593,2016]" box="[151,877,1593,1620]" pageId="6" pageNumber="375">
<materialsCitation box="[151,873,1593,1620]" country="Thailand" location="Pangha Pha Pass" municipality="Mai Hong Son Prov." pageId="6" pageNumber="375" specimenCount="1" typeStatus="holotype">
<typeStatus box="[151,207,1594,1620]" pageId="6" pageNumber="375">Type</typeStatus>
locality:
<collectingCountry box="[312,412,1593,1619]" name="Thailand" pageId="6" pageNumber="375">Thailand</collectingCountry>
(
<location LSID="urn:lsid:plazi:treatment:3A0C87F6DD1EA71851D9FA1DFBD7FC8B:B77A603BDD1EA71950E4F98DFDB9F9CA" box="[426,618,1593,1620]" country="Thailand" municipality="Mai Hong Son Prov." name="Pangha Pha Pass" pageId="6" pageNumber="375">Pangha Pha Pass</location>
,
<collectingMunicipality box="[630,865,1593,1620]" pageId="6" pageNumber="375">Mai Hong Son Prov.</collectingMunicipality>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="375" type="materials_examined">
<paragraph blockId="6.[151,1437,1593,2016]" pageId="6" pageNumber="375">
<emphasis bold="true" box="[199,383,1629,1655]" pageId="6" pageNumber="375">Type materials.</emphasis>
<materialsCitation collectingDate="2009-07-27" collectionCode="MGCL" collectorName="J. B. Heppner" country="Thailand" elevation="868" location="Pangha Pha Pass" municipality="Mai Hong Son Prov." pageId="6" pageNumber="375" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[389,498,1629,1655]" pageId="6" pageNumber="375">
<typeStatus box="[389,498,1629,1655]" pageId="6" pageNumber="375">Holotype</typeStatus>
</emphasis>
: ♁,
<location LSID="urn:lsid:plazi:treatment:3A0C87F6DD1EA71851D9FA1DFBD7FC8B:B77A603BDD1EA719536DF9E9FD0DF9E6" box="[547,734,1629,1656]" country="Thailand" municipality="Mai Hong Son Prov." name="Pangha Pha Pass" pageId="6" pageNumber="375">Pangha Pha Pass</location>
(
<quantity box="[748,815,1629,1655]" metricMagnitude="2" metricUnit="m" metricValue="8.68" pageId="6" pageNumber="375" unit="m" value="868.0">
<elevation box="[748,815,1629,1655]" metricMagnitude="2" metricUnit="m" metricValue="8.68" pageId="6" pageNumber="375" unit="m" value="868.0">868m</elevation>
</quantity>
) [near Soppong],
<collectingMunicipality box="[1015,1247,1629,1656]" pageId="6" pageNumber="375">Mai Hong Son Prov.</collectingMunicipality>
,
<collectingCountry box="[1256,1358,1629,1655]" name="Thailand" pageId="6" pageNumber="375">Thailand</collectingCountry>
,
<date pageId="6" pageNumber="375" value="2009-07-27">
<collectingDate pageId="6" pageNumber="375" value="2009-07-27">27 Jul 2009</collectingDate>
</date>
,
<collectorName box="[221,375,1666,1692]" pageId="6" pageNumber="375">J. B. Heppner</collectorName>
(gen. slide JBH3197; adult photo 12222) (
<collectionCode box="[868,952,1665,1691]" pageId="6" pageNumber="375">MGCL</collectionCode>
) (deposited at
<collectionCode box="[1118,1204,1665,1691]" pageId="6" pageNumber="375">MGCL</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="375" type="diagnosis">
<paragraph blockId="6.[151,1437,1593,2016]" pageId="6" pageNumber="375">
<emphasis bold="true" box="[199,314,1701,1727]" pageId="6" pageNumber="375">Diagnosis</emphasis>
. This species is distinctive in the forewing having a basal orange mark besides the midwing orange fascia, the latter also without an extension to the apex (with apical finger-like extension in
<taxonomicName authority="Heppner &amp; Bae, 2023" authorityName="Heppner &amp; Bae" authorityYear="2023" box="[1147,1325,1737,1763]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="375" phylum="Arthropoda" rank="species" species="cambodiensis" status="sp. nov.">
<emphasis box="[1147,1325,1737,1763]" italics="true" pageId="6" pageNumber="375">V. cambodiensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1332,1420,1737,1763]" pageId="6" pageNumber="375">
<taxonomicNameLabel box="[1332,1420,1737,1763]" pageId="6" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Male genitalia with basal appendage on valvae.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="376" pageId="6" pageNumber="375" type="description">
<paragraph blockId="6.[151,1437,1593,2016]" lastBlockId="7.[151,1437,150,789]" lastPageId="7" lastPageNumber="376" pageId="6" pageNumber="375">
<emphasis bold="true" box="[199,337,1809,1835]" pageId="6" pageNumber="375">Description</emphasis>
. Wing expanse:
<quantity box="[527,628,1809,1836]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="375" unit="mm" value="13.5">13.5 mm</quantity>
male (n = 1). Male (
<figureCitation box="[864,935,1809,1835]" captionStart="FIGURES 35" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[199,1398,1421,1873]" captionTargetId="figure-539@4.[190,1398,1412,1875]" captionTargetPageId="4" captionText="FIGURES 35. Vangatinea species: 3) V. sontraensis sp. nov. (Vietnam: Danang), male holotpe (11.4mm wingspan) (MGCL); 4) V. panghapha sp. nov. (Thailand: Mai Hong Son), male holotype (13.5mm wingspan) (MGCL); 5) V. cambodiensis sp. nov. (Cambodia: Pursat), female holotype (16mm wingspan) (INUC)." figureDoi="http://doi.org/10.5281/zenodo.7836804" httpUri="https://zenodo.org/record/7836804/files/figure.png" pageId="6" pageNumber="375">Fig. 4</figureCitation>
).
<emphasis box="[956,1017,1809,1835]" italics="true" pageId="6" pageNumber="375">Head</emphasis>
(
<figureCitation box="[1033,1103,1809,1835]" captionStart="FIGURES 68" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[205,1382,181,1920]" captionTargetId="figure-17@5.[205,1382,181,1920]" captionTargetPageId="5" captionText="FIGURES 68. Head views of Vangatinea species (lateral and (a) dorsal views): 6) V. sontraensis sp. nov. (holotype male); 7) V. panghapha sp. nov. (holotype male); 8) V. cambodiensis sp. nov. (holotype female)." figureDoi="http://doi.org/10.5281/zenodo.7836810" httpUri="https://zenodo.org/record/7836810/files/figure.png" pageId="6" pageNumber="375">Fig. 7</figureCitation>
): vertex orange; frons yellow and laterally; caudal eye margin yellow; labial palpus yellow-white, with distal segment lighter and dark brown on basal segment, and bristles dark brown; antenna purple-brown, with light yellow segments on basal 5 segments and at 4/
<quantity box="[249,291,1917,1944]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="375" unit="in" value="5.0">5 in</quantity>
short section; scape yellow.
<emphasis box="[614,695,1917,1943]" italics="true" pageId="6" pageNumber="375">Thorax</emphasis>
: golden-yellow to orange; tegula purple-brown to golden distally; venter purple-brown; forelegs purple-brown except yellow coxae, with remainder of legs golden except purplebrown markings at joints and dorsally on tarsi.
<emphasis box="[671,779,1990,2015]" italics="true" pageId="6" pageNumber="375">Forewing</emphasis>
(
<figureCitation box="[793,862,1989,2015]" captionStart="FIGURES 35" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[199,1398,1421,1873]" captionTargetId="figure-539@4.[190,1398,1412,1875]" captionTargetPageId="4" captionText="FIGURES 35. Vangatinea species: 3) V. sontraensis sp. nov. (Vietnam: Danang), male holotpe (11.4mm wingspan) (MGCL); 4) V. panghapha sp. nov. (Thailand: Mai Hong Son), male holotype (13.5mm wingspan) (MGCL); 5) V. cambodiensis sp. nov. (Cambodia: Pursat), female holotype (16mm wingspan) (INUC)." figureDoi="http://doi.org/10.5281/zenodo.7836804" httpUri="https://zenodo.org/record/7836804/files/figure.png" pageId="6" pageNumber="375">Fig. 4</figureCitation>
): lustrous dark brown, with dark orange fascia from costa at 2/5 and recurved to wing base, and wide orange fascia at midwing and with concave emargination by tornus; fringe purple-brown and long from before apex to tornus; venter lustrous dark brown, with light yellow patch on tornus.
<emphasis box="[235,345,223,249]" italics="true" pageId="7" pageNumber="376">Hindwing</emphasis>
(
<figureCitation box="[360,429,223,249]" captionStart="FIGURES 35" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[199,1398,1421,1873]" captionTargetId="figure-539@4.[190,1398,1412,1875]" captionTargetPageId="4" captionText="FIGURES 35. Vangatinea species: 3) V. sontraensis sp. nov. (Vietnam: Danang), male holotpe (11.4mm wingspan) (MGCL); 4) V. panghapha sp. nov. (Thailand: Mai Hong Son), male holotype (13.5mm wingspan) (MGCL); 5) V. cambodiensis sp. nov. (Cambodia: Pursat), female holotype (16mm wingspan) (INUC)." figureDoi="http://doi.org/10.5281/zenodo.7836804" httpUri="https://zenodo.org/record/7836804/files/figure.png" pageId="7" pageNumber="376">Fig. 4</figureCitation>
): pale bronze-brown, with purple-brown near apex; fringe purple-brown from before apex to tornus; venter lustrous dark brown.
<emphasis box="[589,695,259,285]" italics="true" pageId="7" pageNumber="376">Abdomen</emphasis>
: dark brown, with metallic bronze iridescence, with paler caudal marginal scales per tergite; venter light brown with bronze iridescence, with segmental caudal margins creamwhite; genital tufts brown with bronze iridescence; S6 caudally convex, medially conical.
<emphasis box="[1160,1324,331,357]" italics="true" pageId="7" pageNumber="376">Male genitalia</emphasis>
(
<figureCitation box="[1339,1423,331,357]" captionStart="FIGURE 10" captionStartId="7.[151,250,1645,1669]" captionTargetBox="[155,1436,814,1621]" captionTargetId="figure-283@7.[151,1437,813,1621]" captionTargetPageId="7" captionText="FIGURE 10. Vangatinea panghapha sp. nov., holotype male genitalia (Thailand: Mai Hong Son), with details, a) aedeagus, b) juxta-anellus, and c) pre-genital segment of abdomen (gen. slide JBH-3197) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.7836818" httpUri="https://zenodo.org/record/7836818/files/figure.png" pageId="7" pageNumber="376">Fig. 10</figureCitation>
): tegumen-vinculum as fused cylinder, with strong dorsal transtilla-like semi-circular ring; uncus undeveloped and membranous; medial linear subscaphium on diaphragma; valva quadratic and setose, with saccular margin strongly developed and densely setose and basally abruptly convex before basal joint; valval base with small triangular apically acute setose appendage; juxta-anellus complex as dorsal convex collar with lateral extended flattened arms to lateral semi-circular flattened ends; short oblong basal stylus plate; saccus short; aedeagus narrow and long, cornutus a single long thin tubule (subequal to aedeagus length and width).
<emphasis box="[998,1082,547,573]" italics="true" pageId="7" pageNumber="376">Female</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="376" type="etymology">
<paragraph blockId="7.[151,1437,150,789]" pageId="7" pageNumber="376">
<emphasis bold="true" box="[199,328,583,609]" pageId="7" pageNumber="376">Etymology</emphasis>
. The species is named for the
<typeStatus box="[671,719,583,609]" pageId="7" pageNumber="376">type</typeStatus>
locality, Pangha Pha Pass, northwestern
<collectingCountry box="[1179,1279,583,609]" name="Thailand" pageId="7" pageNumber="376">Thailand</collectingCountry>
(northwest of
<collectingRegion box="[151,289,619,645]" country="Thailand" name="Chiang Mai" pageId="7" pageNumber="376">Chiang Mai</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection box="[199,422,654,681]" pageId="7" pageNumber="376" type="biology_ecology">
<paragraph blockId="7.[151,1437,150,789]" box="[199,422,654,681]" pageId="7" pageNumber="376">
<emphasis bold="true" box="[199,289,654,680]" pageId="7" pageNumber="376">Biology</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection box="[199,1216,690,717]" pageId="7" pageNumber="376" type="distribution">
<paragraph blockId="7.[151,1437,150,789]" box="[199,1216,690,717]" pageId="7" pageNumber="376">
<emphasis bold="true" box="[199,345,690,716]" pageId="7" pageNumber="376">Distribution</emphasis>
. Known only from far northwestern
<collectingCountry box="[758,858,691,717]" name="Thailand" pageId="7" pageNumber="376">Thailand</collectingCountry>
(Mai Hong Son) (
<figureCitation box="[1063,1144,691,717]" captionStart="FIGURE 12" captionStartId="9.[151,250,1934,1958]" captionTargetBox="[311,1277,930,1911]" captionTargetId="figure-367@9.[311,1277,930,1911]" captionTargetPageId="9" captionText="FIGURE 12. Southeast Asia distribution map of Vangatinea species: V. cambodiensis sp. nov. (white square, Cambodia), V. panghapha sp. nov. (black star, Thailand), and V. sontraensis sp. nov. (black square, Vietnam) (base map courtesy of J. Bartholomew &amp; Son Maps, Edinburgh, Scotland; Times Atlas, 1988. London, UK)." figureDoi="http://doi.org/10.5281/zenodo.7836822" httpUri="https://zenodo.org/record/7836822/files/figure.png" pageId="7" pageNumber="376">Fig. 12</figureCitation>
map).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="376" type="discussion">
<paragraph blockId="7.[151,1437,150,789]" pageId="7" pageNumber="376">
<emphasis bold="true" box="[199,325,726,752]" pageId="7" pageNumber="376">Discussion</emphasis>
. The new species is closest to
<taxonomicName authority="Heppner &amp; Bae, 2023" authorityName="Heppner &amp; Bae" authorityYear="2023" box="[683,864,727,753]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="376" phylum="Arthropoda" rank="species" species="cambodiensis" status="sp. nov.">
<emphasis box="[683,864,727,753]" italics="true" pageId="7" pageNumber="376">V. cambodiensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[874,965,727,753]" pageId="7" pageNumber="376">
<taxonomicNameLabel box="[874,965,727,753]" pageId="7" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, known only from the female, and could possibly be the male of the same species, but it seems to be a separate species.
</paragraph>
</subSubSection>
</treatment>
</document>