192 lines
32 KiB
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192 lines
32 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.128.1804" ID-PMC="PMC3175138" ID-Pensoft-Pub="1313-2970-128-15" ID-Pensoft-UUID="6157FFBE34536B410C79CB5C11170069" ID-PubMed="21998550" ID-Zenodo-Dep="577176" ID-ZooBank="837E761ADDC84351B1080F9B9CDC7ED1" ModsDocID="1313-2970-128-15" checkinTime="1451249919653" checkinUser="pensoft" docAuthor="Valim, Ye, Kavanaugh, David H., Shi, Hongliang & Liang, Hongbin" docDate="2011" docId="3A0C181636E554D69F88152B76C4CE96" docLanguage="en" docName="ZooKeys 128: 15-52" docOrigin="ZooKeys 128" docPubDate="2011-09-09" docSource="http://dx.doi.org/10.3897/zookeys.128.1804" docTitle="Chlaenius chuanqianensis Liu & Liang 2011, sp. n." docType="treatment" docUuid="2DA7CCAE-4DD2-4CF0-A722-88CA003C691A" docUuidSource="ZooBank" docVersion="4" id="6157FFBE34536B410C79CB5C11170069" lastPageNumber="20" masterDocId="6157FFBE34536B410C79CB5C11170069" masterDocTitle="A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species" masterLastPageNumber="52" masterPageNumber="15" pageNumber="19" updateTime="1668152074611" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Valim, Ye</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China & Key Laboratory for Plant Pests Management of Mountainous Region, Institute of Entomology, Guizhou, University, Guiyang 550025, China</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kavanaugh, David H.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Science, San Francisco, California 94118, U. S. A.</mods:affiliation>
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<mods:nameIdentifier type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Shi, Hongliang</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liang, Hongbin</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2011-09-09</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>128</mods:number>
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<mods:extent unit="page">
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<mods:start>15</mods:start>
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<mods:end>52</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.128.1804</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.128.1804</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-128-15</mods:identifier>
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<mods:identifier type="ZooBank">837E761ADDC84351B1080F9B9CDC7ED1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">6157FFBE34536B410C79CB5C11170069</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577176</mods:identifier>
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<treatment ID-GBIF-Taxon="182236337" LSID="urn:lsid:zoobank.org:act:2DA7CCAE-4DD2-4CF0-A722-88CA003C691A" httpUri="http://treatment.plazi.org/id/3A0C181636E554D69F88152B76C4CE96" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
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<subSubSection pageId="4" pageNumber="19" type="nomenclature">
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<paragraph pageId="4" pageNumber="19">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:2DA7CCAE-4DD2-4CF0-A722-88CA003C691A" authority="Liu & Liang" authorityName="Liu & Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chlaenius chuanqianensis" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="chuanqianensis" status="sp. n.">Chlaenius Lithochlaenius chuanqianensis Liu & Liang</taxonomicName>
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<taxonomicNameLabel pageId="4" pageNumber="19">sp. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 16–21" captionStartId="F2" captionText="Figures 16 - 21. Habitus of Chlaenius (Lithochlaenius) spp 16 Chlaenius chuanqianensis sp. n., holotype, male, dorsal view 17 Chlaenius chuanqianensis sp. n., holotype, male, ventral view 18 Chlaenius linwensini sp. n., holotype, male, dorsal view 19 Chlaenius linwensini sp. n., holotype, male, ventral view 20 Chlaenius propeagilis sp. n., holotype, male, dorsal view 21 Chlaenius propeagilis sp. n., holotype, male, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17279" pageId="4" pageNumber="19">Figs 16-17</figureCitation>
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<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="4" pageNumber="19">52</figureCitation>
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<figureCitation captionStart="Figures 66–79" captionStartId="F9" captionText="Figures 66 - 79. Humeral region of elytra of Chlaenius (Lithochlaenius) spp 66 Chlaenius chuanqianensis sp. n., holotype 67 Chlaenius linwensini sp. n., holotype 68 Chlaenius propeagilis sp. n., holotype 69 Chlaenius agilis Chaudoir, holotype 70 Chlaenius formosensis Lorenz, CCCC 71 Chlaenius leishanensis Kirschenhofer, IZCAS 72 Chlaenius noguchii Bates, IZCAS 73 Chlaenius wrasei Kirschenhofer, IZCAS 74 Chlaenius agiloides Jedlicka, IZCAS 75 Chlaenius rambouseki Lutshnik from Ussuri region, ZCAS 76 Chlaenius anchomenoides Bates, paratype 77 Chlaenius nuristanus Jedlicka, paratype 78 Chlaenius nuristanus Jedlicka, paratype 79 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17286" pageId="4" pageNumber="19">66</figureCitation>
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<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="4" pageNumber="19">80</figureCitation>
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<figureCitation captionStart="Figures 94–116" captionStartId="F11" captionText="Figures 94 - 116. Abdominal sternum VII (ventral aspect) of Chlaenius (Lithochlaenius) species 94 Chlaenius chuanqianensis sp. n., male, holotype 95 Chlaenius chuanqianensis sp. n., female, paratype 96 Chlaenius linwensini sp. n., male, holotype 97 Chlaenius linwensini sp. n., female, paratype 98 Chlaenius propeagilis sp. n., male, holotype 99 Chlaenius propeagilis sp. n., female, paratype 100 Chlaenius agilis Chaudoir, male, holotype 101 Chlaenius formosensis Lorenz, male 102 Chlaenius formosensis Lorenz, female 103 Chlaenius leishanensis Kirschenhofer, male 104 Chlaenius leishanensis Kirschenhofer, female 105 Chlaenius noguchii Bates, male 106 Chlaenius noguchii Bates, female 107 Chlaenius wrasei Kirschenhofer, male 108 Chlaenius wrasei Kirschenhofer, female 109 Chlaenius agiloides Jedlicka, male 110 Chlaenius agiloides Jedlicka, female 111 Chlaenius rambouseki Lutshnik, from Ussuri region, male 112 Chlaenius rambouseki Lutshnik from Ussuri region, female 113 Chlaenius anchomenoides Bates, male, paratype 114 Chlaenius nuristanus Jedlicka, male, paratype 115 Chlaenius nuristanus Jedlicka, female, paratype 116 Chlaenius formosanus Jedlicka from Taiwan, male. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17288" pageId="4" pageNumber="19">94-95</figureCitation>
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<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">117-118</figureCitation>
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<figureCitation captionStart="Figures 146–151" captionStartId="F14" captionText="Figures 146 - 151. Photographs of habitats for Chlaenius (Lithochlaenius) spp. 146 Chlaenius chuanqianensis sp. n., Dabaitang, Xishui county, North Guizhou, China 147 Chlaenius anchomenoides Bates, Goorais valley, Pakistan (Provided by Dr. Muhammad Abbas in Pakistan Museum of Natural History) 148 Chlaenius propeagilis sp. n., Gaoligongshan, Yunnan, China 149 Chlaenius leishanensis Kirschenhofer, Xiaodanjiang, Leigongshan, Leishan county, Southeast Guizhou, China 150 Chlaenius chuanqianensis sp. n. and Chlaenius agiloides Jedlicka, Jinshajiang, Sichuan 151 Chlaenius rambouseki Lutshnik, Ussri river, Heilongjiang, China." httpUri="https://binary.pensoft.net/fig/17291" pageId="4" pageNumber="19">146, 150</figureCitation>
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<figureCitation captionStart="Figure 158" captionStartId="F16" captionText="Figure 158. Map showing the known geographical distributions of Chlaenius (Lithochlaenius) spp." httpUri="https://binary.pensoft.net/fig/17293" pageId="4" pageNumber="19">158</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="19" type="types">
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<paragraph pageId="4" pageNumber="19">Types.</paragraph>
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<paragraph pageId="4" pageNumber="19">
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Holotype: male (IZCAS), "China, Guizhou, Xishui County, Dabaitang, 600m"/ "IOZ & Guizhou Univ. Joint Expedition, 2000.9.24, Liang H.B" / ""Holotype,
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<taxonomicName authorityName="Liu & Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="chuanqianensis" subGenus="Lithochlaenius">
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<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius (Lithochlaenius) chuanqianensis</emphasis>
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</taxonomicName>
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Liu & Liang sp. n."" [red label]. Paratypes: Total 14 specimens. 1 female (IZCAS), "China, Guizhou, Xishui County, Dabaitang, 550m"/ "IOZ & Guizhou Univ Joint Expedition, 2000.9.27, Liang H. B."; 1 female (HBUM), "China, Guizhou, Xishui County, Dabaitang"/ "2000. IX. 25-29, collector Ren G. D."; 1 male (IZCAS), "China, Guizhou, Xishui County, Chengzhai, Hongqi village,
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<geoCoordinate degrees="28.419033" direction="north" orientation="latitude" precision="1" value="28.419033">28.419033</geoCoordinate>
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,
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<geoCoordinate degrees="106.273766" direction="east" orientation="longitude" precision="1" value="106.273766">106.273766</geoCoordinate>
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"/ "2009.10.8, Liu Y. & Shi H. L."; 1 female (IZCAS), "Sichuan, Ya-an"; / ""1990.VII.3, Xie Weiping collector"; 1 male (IZCAS), "Sichuan, Xinjin" / ""1981.VI.13, Liu Hongjiang collector"; 1 female (IZCAS), "Sichuan, Anxian"; 1 male (IZCAS), "Sichuan, Shehong" / ""1980.7.7"; 3 males and 3 females (IZCAS), "Sichuan, Yajiang, Hekou Town, Shanbeihou, Yalongjiang, N30.00020, E101.01526" / "2009.5.28 N, Liang Hongbin coll."; 1 female (IZCAS), "Sichuan, Yajiang, Hekou Town, Shanbeihou, Yalongjiang, E30.00020, N101.01526"/ "2009.5.27, Day, Liang Hongbin coll.". Each paratype with an additional yellow label: ""Paratype,
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<taxonomicName authorityName="Liu & Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="chuanqianensis" subGenus="Lithochlaenius">
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<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius (Lithochlaenius) chuanqianensis</emphasis>
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</taxonomicName>
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Liu & Liang sp. n."".
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="19" type="diagnosis">
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<paragraph pageId="4" pageNumber="19">Diagnosis.</paragraph>
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<paragraph pageId="4" pageNumber="19">
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Antennomere 1 coniform (
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<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="4" pageNumber="19">Fig. 52</figureCitation>
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); color of antennomere 1 yellow or brown; intervals 1-5 glabrous medially, with one row of pubescence laterally (
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<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="4" pageNumber="19">Fig. 80</figureCitation>
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); aedeagus stout, apical lamella round (
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<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Fig. 117</figureCitation>
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), thin and straight in lateral view (
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<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Fig. 118</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="20" type="description">
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<paragraph pageId="5" pageNumber="20">
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<pageBreakToken pageId="5" pageNumber="20" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="5" pageNumber="20">Total length = 16.0-17.0 mm (mean = 16.8), width = 5.60-6.13 mm (mean = 5.86); HW = 2.60-2.95 mm (mean = 2.82), EYL = 0.95-1.10 mm (mean = 1.03), ratio Ant3/Ant1 = 1.56-1.72 (mean = 1.64), PL/PW = 0.76-0.84 (mean = 0.81), EL/EW = 1.68-1.91 (mean = 1.79), EL/ PL = 1.37-1.49 (mean = 1.42).</paragraph>
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<paragraph pageId="5" pageNumber="20">Head and pronotum black with green or coppery metallic luster; elytra black; ventral surface black; mandibles and trochanters dark brown; antennomere 1, femora, tibiae yellow to brown; antennomeres 2-11, palpi and tarsi dark brown to nearly black.</paragraph>
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<paragraph pageId="5" pageNumber="20">
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Head with vertex smooth medially, coarsely punctate behind eyes; labrum slightly emarginate at apex; antennomere 1 coniform (
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<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="5" pageNumber="20">Fig. 52</figureCitation>
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). Pronotum with disk smooth, very sparsely punctate along midline in a few specimens; lateral furrow sparsely pubescent and coarsely punctate; medial longitudinal furrow deep, impunctate; basal foveae narrow, deep, sparsely pubescent and punctate. Elytral intervals convex, intervals 1-5 glabrous medially, with a row of setae laterally (
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<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="5" pageNumber="20">Fig. 80</figureCitation>
|
||
), pubescent in apical one-fifth in a few specimens, intervals 6-9 pubescent throughout; striae deep, punctate; humeral angles obtuse (
|
||
<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="5" pageNumber="20">Fig. 66</figureCitation>
|
||
). Abdominal sterna IV-VI sparsely pubescent medially, densely pubescent laterally; sternum VII narrowly rounded apically in both sexes (
|
||
<figureCitation captionStart="Figures 94–116" captionStartId="F11" captionText="Figures 94 - 116. Abdominal sternum VII (ventral aspect) of Chlaenius (Lithochlaenius) species 94 Chlaenius chuanqianensis sp. n., male, holotype 95 Chlaenius chuanqianensis sp. n., female, paratype 96 Chlaenius linwensini sp. n., male, holotype 97 Chlaenius linwensini sp. n., female, paratype 98 Chlaenius propeagilis sp. n., male, holotype 99 Chlaenius propeagilis sp. n., female, paratype 100 Chlaenius agilis Chaudoir, male, holotype 101 Chlaenius formosensis Lorenz, male 102 Chlaenius formosensis Lorenz, female 103 Chlaenius leishanensis Kirschenhofer, male 104 Chlaenius leishanensis Kirschenhofer, female 105 Chlaenius noguchii Bates, male 106 Chlaenius noguchii Bates, female 107 Chlaenius wrasei Kirschenhofer, male 108 Chlaenius wrasei Kirschenhofer, female 109 Chlaenius agiloides Jedlicka, male 110 Chlaenius agiloides Jedlicka, female 111 Chlaenius rambouseki Lutshnik, from Ussuri region, male 112 Chlaenius rambouseki Lutshnik from Ussuri region, female 113 Chlaenius anchomenoides Bates, male, paratype 114 Chlaenius nuristanus Jedlicka, male, paratype 115 Chlaenius nuristanus Jedlicka, female, paratype 116 Chlaenius formosanus Jedlicka from Taiwan, male. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17288" pageId="5" pageNumber="20">Figs 94-95</figureCitation>
|
||
). Lamella of aedeagus round at apex (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="5" pageNumber="20">Fig. 117</figureCitation>
|
||
), thin and straight in lateral view (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="5" pageNumber="20">Fig. 118</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="20" type="etymology">
|
||
<paragraph pageId="5" pageNumber="20">Etymology.</paragraph>
|
||
<paragraph pageId="5" pageNumber="20">
|
||
The Latinized name
|
||
<emphasis italics="true" pageId="5" pageNumber="20">chuanqianensis</emphasis>
|
||
refers to type localities of this new species in "chuanqian" regions, of which "chuan" refers to Sichuan Province and "qian" refers to Guizhou Province).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="20" type="distribution">
|
||
<paragraph pageId="5" pageNumber="20">Geographical distribution.</paragraph>
|
||
<paragraph pageId="5" pageNumber="20">
|
||
<figureCitation captionStart="Figure 158" captionStartId="F16" captionText="Figure 158. Map showing the known geographical distributions of Chlaenius (Lithochlaenius) spp." httpUri="https://binary.pensoft.net/fig/17293" pageId="5" pageNumber="20">Fig. 158</figureCitation>
|
||
. Known only from Guizhou and Sichuan Provinces, China.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="20" type="remarks">
|
||
<paragraph pageId="5" pageNumber="20">Remarks.</paragraph>
|
||
<paragraph pageId="5" pageNumber="20">Mensural data cited in the description were obtained from the holotype and all paratypes.</paragraph>
|
||
<paragraph pageId="5" pageNumber="20">
|
||
Members of this species are most similar to those of
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="wrasei">
|
||
<emphasis italics="true" pageId="5" pageNumber="20">Chlaenius wrasei</emphasis>
|
||
</taxonomicName>
|
||
in shape of antennomere 1 and elytral pubescence, but differ from the latter in color of antennomere 1 (paler than antennomere 3 in
|
||
<taxonomicName authorityName="Liu & Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="chuanqianensis">
|
||
<emphasis italics="true" pageId="5" pageNumber="20">Chlaenius chuanqianensis</emphasis>
|
||
</taxonomicName>
|
||
, antennomeres 1 and 3 concolorous in
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="wrasei">
|
||
<emphasis italics="true" pageId="5" pageNumber="20">Chlaenius wrasei</emphasis>
|
||
</taxonomicName>
|
||
) and orientation of the apical lamella of the aedeagus of males (straight in
|
||
<taxonomicName authorityName="Liu & Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="chuanqianensis">
|
||
<emphasis italics="true" pageId="5" pageNumber="20">Chlaenius chuanqianensis</emphasis>
|
||
</taxonomicName>
|
||
, bent ventrally in
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="wrasei">
|
||
<emphasis italics="true" pageId="5" pageNumber="20">Chlaenius wrasei</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="5" pageNumber="20">
|
||
They are also similar to
|
||
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="leishanensis">
|
||
<emphasis italics="true" pageId="5" pageNumber="20">Chlaenius leishanensis</emphasis>
|
||
</taxonomicName>
|
||
members in pubescence of elytral intervals, but different from the latter in having yellow tibiae (black or dark brown in
|
||
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="leishanensis">
|
||
<emphasis italics="true" pageId="5" pageNumber="20">Chlaenius leishanensis</emphasis>
|
||
</taxonomicName>
|
||
), and males have a thin aedeagal lamella of the aedeagus (lamella thick in
|
||
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="leishanensis">
|
||
<emphasis italics="true" pageId="5" pageNumber="20">Chlaenius leishanensis</emphasis>
|
||
</taxonomicName>
|
||
males).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |