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<document ID-DOI="10.1206/352.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="5405335" approvalRequired="503" approvalRequired_for_taxonomicNames="62" approvalRequired_for_textStreams="429" approvalRequired_for_treatments="12" checkinTime="1630345222680" checkinUser="tatiana" docAuthor="Nesbitt, Sterling J." docDate="2011" docId="357D771BFFBEFFB5EDABFED0FCF8FADC" docLanguage="en" docName="B352.pdf" docOrigin="Bulletin of the American Museum of Natural History 2011 (352)" docSource="http://www.bioone.org/doi/abs/10.1206/352.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.3:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="3" docTitle="Riojasuchus tenuisceps Bonaparte 1967" docType="treatment" docVersion="7" lastPageNumber="22" masterDocId="C9440F63FFABFFA0EF17FFF9FFBFFF8E" masterDocTitle="The Early Evolution Of Archosaurs: Relationships And The Origin Of Major Clades" masterLastPageNumber="292" masterPageNumber="1" pageNumber="22" updateTime="1631052925311" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>The Early Evolution Of Archosaurs: Relationships And The Origin Of Major Clades</mods:title>
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<mods:namePart>Nesbitt, Sterling J.</mods:namePart>
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<mods:date>2011</mods:date>
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<mods:number>2011-04-29</mods:number>
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<mods:number>2011</mods:number>
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<mods:number>352</mods:number>
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<mods:identifier type="DOI">10.1206/352.1</mods:identifier>
<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5476372" ID-Zenodo-Dep="5476372" LSID="urn:lsid:plazi:treatment:357D771BFFBEFFB5EDABFED0FCF8FADC" httpUri="http://treatment.plazi.org/id/357D771BFFBEFFB5EDABFED0FCF8FADC" lastPageNumber="22" pageId="21" pageNumber="22">
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<paragraph blockId="21.[700,1160,297,349]" box="[700,1160,297,320]" pageId="21" pageNumber="22">
<heading box="[700,1160,297,320]" centered="true" fontSize="9" level="2" pageId="21" pageNumber="22" reason="2">
<taxonomicName authority="Bonaparte, 1967" authorityName="Bonaparte" authorityYear="1967" box="[700,1160,297,320]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="tenuisceps">
<emphasis box="[700,958,297,320]" italics="true" pageId="21" pageNumber="22">Riojasuchus tenuisceps</emphasis>
<bibRefCitation author="Bonaparte, J. F." box="[967,1160,298,320]" pageId="21" pageNumber="22" refId="ref184820" refString="Bonaparte, J. F. 1967. Dos nuevas '' faunas' ' de reptiles Triasicos de Argentina. Gondwana Stratigraphy. I. U. G. S. Symposium: 283 - 306. Paris: UNESCO." type="proceedings" year="1967">Bonaparte, 1967</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="21.[700,1160,297,349]" box="[864,996,327,349]" pageId="21" pageNumber="22">
<heading box="[864,996,327,349]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="22" reason="4">
(
<figureCitation box="[872,988,327,349]" captionStart="Fig" captionStartId="20.[152,190,1552,1571]" captionText="Fig. 8. Skull reconstructions of pseudosuchian archosaur terminal taxa: A, Riojasuchus tenuisceps in lateral and B, dorsal views; redrawn from Sereno (1991a); C, Gracilisuchus stipanicicorum in lateral and D, dorsal views; redrawn from Romer (1971b); E, Arizonasaurus babbitti in lateral view; redrawn from Nesbitt (2005); F, Stagonolepis robertsoni in lateral and G, dorsal views; redrawn from Walker (1961); H, Revueltosaurus callenderi in lateral view; based on PEFO 34561. I, Effigia okeeffeae in lateral view; redrawn from Nesbitt and Norell (2006); J, Xilousuchus sapingensis in lateral view; based on IVPP V 6026. Shaded areas indicate incomplete preservation. See appendix for anatomical abbreviations. Scale bars 5 1 cm." pageId="21" pageNumber="22">fig. 8AB</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="description">
<paragraph blockId="21.[667,1193,371,1362]" pageId="21" pageNumber="22">
AGE: Norian-?Rhaetian, Late Triassic (
<bibRefCitation author="Arcucci, A. B. &amp; C. A. Marsicano &amp; A. T. Caselli" box="[674,909,402,424]" pageId="21" pageNumber="22" pagination="557 - 568" refId="ref183936" refString="Arcucci, A. B., C. A. Marsicano, and A. T. Caselli. 2004. Tetrapod association and palaeoenvironment of the Los Colorados Formation (Argentina): a significant sample from western Gondwana at the end of the Triassic. Geobios 37: 557 - 568." type="journal article" year="2004">Arcucci et al., 2004</bibRefCitation>
).
</paragraph>
<paragraph blockId="21.[667,1193,371,1362]" pageId="21" pageNumber="22">
OCCURRENCE: Los Colorados Formation, El Salto,
<collectingCountry box="[778,896,461,483]" name="Argentina" pageId="21" pageNumber="22">Argentina</collectingCountry>
(
<bibRefCitation author="Arcucci, A. B. &amp; C. A. Marsicano &amp; A. T. Caselli" box="[912,1147,461,483]" pageId="21" pageNumber="22" pagination="557 - 568" refId="ref183936" refString="Arcucci, A. B., C. A. Marsicano, and A. T. Caselli. 2004. Tetrapod association and palaeoenvironment of the Los Colorados Formation (Argentina): a significant sample from western Gondwana at the end of the Triassic. Geobios 37: 557 - 568." type="journal article" year="2004">Arcucci et al., 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="materials_examined">
<paragraph blockId="21.[667,1193,371,1362]" pageId="21" pageNumber="22">
<typeStatus box="[696,824,489,513]" pageId="21" pageNumber="22">HOLOTYPE</typeStatus>
: PVL 3827, complete skull, cervical, dorsal, sacral and caudal vertebrae, scapula, coracoid, humerus, distal portion of the radius and ulna, partial manus, ilium, pubis, femur, tibia, fibula, nearly complete pes.
</paragraph>
<paragraph blockId="21.[667,1193,371,1362]" pageId="21" pageNumber="22">REFERRED MATERIAL: PVL 3828, nearly complete skull, cervical, dorsal, sacral and caudal vertebrae, scapula, coracoid, humerus, ulna, radius, pubis, ischium, ilium, femur, tibia, fibula, calcaneum; PVL 2826 cervical, dorsal, sacral and caudal vertebrae, coracoids, scapula fragments, humerus, ulna, radius, ilium, femur, and tibia; PVL 3814 vertebrae, humerus, and tibia.</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="discussion">
<paragraph blockId="21.[667,1193,371,1362]" pageId="21" pageNumber="22">
REMARKS:
<taxonomicName box="[837,972,900,923]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis box="[837,972,900,923]" italics="true" pageId="21" pageNumber="22">Riojasuchus</emphasis>
</taxonomicName>
is represented by nearly all portions of the skeleton. The wellpreserved complete skull was well described by
<bibRefCitation author="Bonaparte, J. F." box="[713,927,989,1011]" pageId="21" pageNumber="22" pagination="1 - 183" refId="ref184891" refString="Bonaparte, J. F. 1971. Los tetrapodos del sector superior de la Formacion Los Colorados, La Rioja, Argentina. (Triasico Superior) I parte. Opera Lilloana 22: 1 - 183." type="journal article" year="1971">Bonaparte (1971)</bibRefCitation>
, followed by
<bibRefCitation author="Sereno, P. C." pageId="21" pageNumber="22" pagination="1 - 53" refId="ref200036" refString="Sereno, P. C. 1991 a. Basal archosaurs: phylogenetic relationships and functional implications. Journal of Vertebrate Paleontology 10 (supplement to 3): 1 - 53." type="journal article" year="1991">Sereno (1991a)</bibRefCitation>
. It is clearly the youngest member of the
<taxonomicName box="[718,911,1047,1069]" class="Reptilia" family="Ornithosuchidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="family">Ornithosuchidae</taxonomicName>
and apparently represents a late surviving member of the clade.
</paragraph>
<paragraph blockId="21.[667,1193,371,1362]" pageId="21" pageNumber="22">
<bibRefCitation author="Sereno, P. C." box="[696,872,1106,1128]" pageId="21" pageNumber="22" pagination="1 - 53" refId="ref200036" refString="Sereno, P. C. 1991 a. Basal archosaurs: phylogenetic relationships and functional implications. Journal of Vertebrate Paleontology 10 (supplement to 3): 1 - 53." type="journal article" year="1991">Sereno (1991a)</bibRefCitation>
provided a list of autapomorphies that all differentiate
<taxonomicName box="[1058,1193,1135,1158]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis box="[1058,1193,1135,1158]" italics="true" pageId="21" pageNumber="22">Riojasuchus</emphasis>
</taxonomicName>
from
<taxonomicName baseAuthorityName="BMNH R" baseAuthorityYear="2409" box="[740,902,1164,1187]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis box="[740,902,1164,1187]" italics="true" pageId="21" pageNumber="22">Ornithosuchus</emphasis>
</taxonomicName>
. However, autapomorphies I, K, and L have a wider distribution. For example, aetosaurs (e.g.,
<taxonomicName authority=", SMNS" authorityName="SMNS" class="Reptilia" family="Stagonolepididae" genus="Aetosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis box="[1059,1185,1224,1245]" italics="true" pageId="21" pageNumber="22">Aetosaurus</emphasis>
, SMNS
</taxonomicName>
5770) possess a sloping occiput much like that of
<taxonomicName box="[807,942,1281,1304]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecodontia" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis box="[807,942,1281,1304]" italics="true" pageId="21" pageNumber="22">Riojasuchus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="key">
<paragraph blockId="21.[667,1193,371,1362]" pageId="21" pageNumber="22">
KEY REFERENCES:
<bibRefCitation author="Bonaparte, J. F." box="[922,1118,1311,1333]" pageId="21" pageNumber="22" refId="ref184820" refString="Bonaparte, J. F. 1967. Dos nuevas '' faunas' ' de reptiles Triasicos de Argentina. Gondwana Stratigraphy. I. U. G. S. Symposium: 283 - 306. Paris: UNESCO." type="proceedings" year="1967">Bonaparte, 1967</bibRefCitation>
;
<bibRefCitation author="Bonaparte, J. F." box="[1131,1188,1311,1332]" pageId="21" pageNumber="22" pagination="1 - 183" refId="ref184891" refString="Bonaparte, J. F. 1971. Los tetrapodos del sector superior de la Formacion Los Colorados, La Rioja, Argentina. (Triasico Superior) I parte. Opera Lilloana 22: 1 - 183." type="journal article" year="1971">1971</bibRefCitation>
;
<bibRefCitation author="Sereno, P. C." box="[667,834,1341,1362]" pageId="21" pageNumber="22" pagination="1 - 53" refId="ref200036" refString="Sereno, P. C. 1991 a. Basal archosaurs: phylogenetic relationships and functional implications. Journal of Vertebrate Paleontology 10 (supplement to 3): 1 - 53." type="journal article" year="1991">Sereno, 1991a</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>