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<mods:title id="C8459FAEE264F031E07DB13BD6A4A9D8">Redescription and taxonomic status of Dipturus chilensis (Guichenot, 1848), and description of Dipturus lamillai sp. nov. (Rajiformes: Rajidae), a new species of long-snout skate from the Falkland Islands</mods:title>
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<paragraph id="977D36BFFF84FFB0A782274CFE60F8DF" blockId="4.[151,472,1824,1851]" box="[151,472,1824,1851]" pageId="4" pageNumber="505">
<heading id="CC3581D3FF84FFB0A782274CFE60F8DF" bold="true" box="[151,472,1824,1851]" fontSize="11" level="1" pageId="4" pageNumber="505" reason="1">
<taxonomicName id="50C24D3CFF84FFB0A782274CFE60F8DF" ID-CoL="632SL" ID-ENA="117858" authority="Rafinesque 1810" authorityName="Rafinesque" authorityYear="1810" box="[151,472,1824,1851]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="505" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF84FFB0A782274CFE60F8DF" bold="true" box="[151,472,1824,1851]" pageId="4" pageNumber="505">
<emphasis id="A5B6EAADFF84FFB0A782274CFEBBF8DE" bold="true" box="[151,259,1824,1850]" italics="true" pageId="4" pageNumber="505">Dipturus</emphasis>
<bibRefCitation id="F3534B4EFF84FFB0A61F274DFE60F8DF" author="Rafinesque, C." box="[266,472,1825,1851]" pageId="4" pageNumber="505" pagination="1 - 20" refId="ref14213" refString="Rafinesque, C. (1810) Caratteri di alcuni nuovi generi e nuove specie di animali e piante della Sicilia, con varie osservazioni sopra i medisimi. Sanfilippo, Palermo. (Part 1 involves fishes, pp. [i-iv] 3 - 69, Part 2 with slightly different title, pp. ia-iva + 71 - 105). Pls. 1 - 20." type="book chapter" year="1810">Rafinesque 1810</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="977D36BFFF84FFB0A782270BFC92F899" blockId="4.[151,1024,1895,1948]" box="[151,810,1895,1918]" pageId="4" pageNumber="505">
<emphasis id="A5B6EAADFF84FFB0A782270BFE92F899" bold="true" box="[151,298,1895,1917]" pageId="4" pageNumber="505">
<typeStatus id="4879881DFF84FFB0A782270BFF74F899" box="[151,204,1895,1917]" pageId="4" pageNumber="505">Type</typeStatus>
Species:
</emphasis>
<taxonomicName id="50C24D3CFF84FFB0A6252705FD8BF899" authority="Linnaeus 1758" authorityName="Linnaeus" authorityYear="1758" box="[304,563,1895,1918]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="505" phylum="Chordata" rank="species" species="batis">
<emphasis id="A5B6EAADFF84FFB0A6252705FE2FF899" box="[304,407,1895,1918]" italics="true" pageId="4" pageNumber="505">Raja batis</emphasis>
Linnaeus 1758
</taxonomicName>
by original designation.
</paragraph>
<paragraph id="977D36BFFF84FFB0A78227EAFBB8F878" blockId="4.[151,1024,1895,1948]" box="[151,1024,1926,1948]" pageId="4" pageNumber="505">
<emphasis id="A5B6EAADFF84FFB0A78227EAFEB7F878" bold="true" box="[151,271,1926,1948]" pageId="4" pageNumber="505">Synonyms:</emphasis>
<taxonomicName id="50C24D3CFF84FFB0A60127EBFE0EF878" authority="" box="[276,438,1926,1948]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="505" phylum="Chordata" rank="subGenus" subGenus="Dipturus">
<emphasis id="A5B6EAADFF84FFB0A60127EBFEFCF878" box="[276,324,1927,1948]" italics="true" pageId="4" pageNumber="505">Raja</emphasis>
(
<emphasis id="A5B6EAADFF84FFB0A64627EBFE15F878" box="[339,429,1927,1948]" italics="true" pageId="4" pageNumber="505">Dipturus</emphasis>
)
</taxonomicName>
as subgenus by Stehmann (1970);
<taxonomicName id="50C24D3CFF84FFB0A40227EBFC44F878" authority="Whitley 1939" authorityName="Whitley" authorityYear="1939" box="[791,1020,1926,1948]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="505" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF84FFB0A40227EBFCDFF878" box="[791,871,1927,1948]" italics="true" pageId="4" pageNumber="505">Zearaja</emphasis>
<bibRefCitation id="F3534B4EFF84FFB0A47B27EAFC44F878" author="Whitley, G." box="[878,1020,1926,1948]" pageId="4" pageNumber="505" pagination="227 - 262" refId="ref14478" refString="Whitley, G. (1939) Taxonomic notes on sharks and rays. Australian Zoologist, 9 (3), 227 - 262." type="journal article" year="1939">Whitley 1939</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DFD86534FF84FFB1A78227A5FE01FCF0" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505" type="description">
<paragraph id="977D36BFFF84FFB0A78227A5FA24F805" blockId="4.[151,1436,1993,2018]" box="[151,1436,1993,2018]" pageId="4" pageNumber="505">
<emphasis id="A5B6EAADFF84FFB0A78227A5FE3EF806" bold="true" box="[151,390,1993,2018]" pageId="4" pageNumber="505">Diagnostic features.</emphasis>
Modified from
<bibRefCitation id="F3534B4EFF84FFB0A52A27A5FD48F805" author="Ishihara H." box="[575,752,1993,2018]" pageId="4" pageNumber="505" pagination="241 - 285" refId="ref13031" refString="Ishihara H. (1987) Revision of the Western North Pacific species of the genus Raja. Japanese Journal of Ichthyology, 34 (3), 241 - 285. https: // doi. org / 10.1007 / BF 03380117" type="journal article" year="1987">Ishihara (1987)</bibRefCitation>
,
<bibRefCitation id="F3534B4EFF84FFB0A5EB27A5FC23F805" author="Leible, M." box="[766,923,1993,2018]" pageId="4" pageNumber="505" pagination="131 - 176" refId="ref13519" refString="Leible, M. (1987) Taxonomic description of juveniles and adults of Raja (Dipturus) flavirostris Philippi, 1892 (Rajiformes: Rajidae) captured in front of the coast off central Chile. Gayana Zoologia, 51, 131 - 176." type="journal article" year="1987">Leible (1987)</bibRefCitation>
,
<bibRefCitation id="F3534B4EFF84FFB0A4BF27A5FB50F805" author="McEachran, J. &amp; Dunn, K." box="[938,1256,1993,2018]" pageId="4" pageNumber="505" pagination="271 - 290" refId="ref13782" refString="McEachran, J. &amp; Dunn, K. (1998) Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia, 1998 (2), 271 - 290. https: // doi. org / 10.2307 / 1447424" type="journal article" year="1998">McEachran &amp; Dunn (1998)</bibRefCitation>
, and
<bibRefCitation id="F3534B4EFF84FFB0A23D27A6FA24F805" author="Last, P. R. &amp; White, W. T. &amp; Carvalho, M. R. de &amp; Seret, B. &amp; Stehmann, M. F. W. &amp; Naylor, G. J. P." box="[1320,1436,1993,2017]" pageId="4" pageNumber="505" refId="ref13352" refString="Last, P. R., White, W. T., Carvalho, M. R. de, Seret, B., Stehmann, M. F. W. &amp; Naylor, G. J. P. (Eds.) (2016) Rays of the World. CSIRO Publishing, Melbourne, 790 pp." type="book" year="2016">
Last
<emphasis id="A5B6EAADFF84FFB0A27727A6FA2DF805" box="[1378,1429,1993,2017]" italics="true" pageId="4" pageNumber="505">et al</emphasis>
.
</bibRefCitation>
</paragraph>
<paragraph id="977D36BFFF85FFB1A78220FBFE01FCF0" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="506">
(2016). Disc broad, rhombic-shaped, outer corners sharply rounded to angular; margins deeply concave anteriorly and produced posteriorly; brownish to dark grey on both dorsal and ventral surfaces; with numerous mucus and sensory pores of irregular pattern visible as black dots and short streaks, mostly on ventral surface; snout elongated, acutely angled, with slightly translucent area flanking rostral cartilage; rostral cartilage long, its length from tip to axils of nasal capsules usually more than 60% of dorsal head length; rostral appendices attached over all or most of length to rostral cartilage; pre-cerebral fontanelle elongated, narrow, extending anteriorly into the rostral cartilage well anterior to leading edge of nasal capsules; nasal capsules without basal fenestrae; inter-nasal plate broad; internarial width usually exceeding 60% of pre-nasal length; pre-orbital processes developed; scapulocoracoid without anterior bridge; distance between pro- and meso- less than distance between meso- and meta-condyles, with postventral fenestra; propterygium of pectoral girdle falling short of middle length of rostral shaft; claspers enlarged, broad at shield, with pointed or spatulated distal lobe; shield long or short, capable of rotation; clasper glans expanded, with cleft, sentina, pseudorhipidion, and rhipidion; spike concealed by integument or exposed; shield with pleated lamellae on its inner surface; sentinel may be absent; funnel absent; clasper skeleton with 3 dorsal terminal cartilages on dorsal aspect of clasper; 1 or 2 accessory terminal cartilages; pectoral radials 7786; trunk vertebrae 2534, pre-dorsal caudal vertebrae 5264; tail with two subterminal dorsal fins; caudal fin small to rudimentary; electrocytes in caudal electric organs with cortical processes; egg capsules broad, convex dorsally, concave or flattened ventrally, short horns, length of egg capsule excluding horns, more than
<quantity id="503A9B5AFF85FFB1A3A822BBFAB7FD0B" box="[1213,1295,727,752]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="506" unit="mm" value="65.0">65 mm</quantity>
, with lateral keels and posterior apron.
</paragraph>
</subSubSection>
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<paragraph id="977D36BFFF85FFB1A7D22373FF45FAB1" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="506">
<emphasis id="A5B6EAADFF85FFB1A7D22373FE83FCDC" bold="true" box="[199,315,799,824]" pageId="5" pageNumber="506">Remarks.</emphasis>
The genus
<taxonomicName id="50C24D3CFF85FFB1A6D6234CFD9FFCD3" authorityName="Rafinesque" authorityYear="1810" box="[451,551,800,823]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A6D6234CFD9FFCD3" box="[451,551,800,823]" italics="true" pageId="5" pageNumber="506">Dipturus</emphasis>
</taxonomicName>
was established by
<bibRefCitation id="F3534B4EFF85FFB1A4052373FC63FCDC" author="Rafinesque, C." box="[784,987,799,824]" pageId="5" pageNumber="506" pagination="1 - 20" refId="ref14213" refString="Rafinesque, C. (1810) Caratteri di alcuni nuovi generi e nuove specie di animali e piante della Sicilia, con varie osservazioni sopra i medisimi. Sanfilippo, Palermo. (Part 1 involves fishes, pp. [i-iv] 3 - 69, Part 2 with slightly different title, pp. ia-iva + 71 - 105). Pls. 1 - 20." type="book chapter" year="1810">Rafinesque (1810</bibRefCitation>
; page 16) as: “Due ale dorsali sopra la coda, nessun ala caudale alla sua estremità—Oss. La
<taxonomicName id="50C24D3CFF85FFB1A5EA2329FC4AFCB9" authority="Batio di Linneo" authorityName="Batio di Linneo" box="[767,1010,836,861]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A5EA2329FCC2FCB8" box="[767,890,837,860]" italics="true" pageId="5" pageNumber="506">Raja Batio</emphasis>
di Linneo
</taxonomicName>
compone sola questo genere” (“Two dorsal fins on the tail, without a caudal fin at the extremity—the
<taxonomicName id="50C24D3CFF85FFB1A47E2304FC5CFC9B" authority="Batio" authorityName="Batio" box="[875,996,872,895]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A47E2304FC5CFC9B" box="[875,996,872,895]" italics="true" pageId="5" pageNumber="506">Raja Batio</emphasis>
</taxonomicName>
of Linnaeus is the only one composing this genus”). Despite its brevity and lack of detail, the diagnosis has been widely accepted and, according to
<bibRefCitation id="F3534B4EFF85FFB1A24423E1FEB6FC2C" author="Last, P. R. &amp; White, W. T. &amp; Carvalho, M. R. de &amp; Seret, B. &amp; Stehmann, M. F. W. &amp; Naylor, G. J. P." pageId="5" pageNumber="506" refId="ref13352" refString="Last, P. R., White, W. T., Carvalho, M. R. de, Seret, B., Stehmann, M. F. W. &amp; Naylor, G. J. P. (Eds.) (2016) Rays of the World. CSIRO Publishing, Melbourne, 790 pp." type="book" year="2016">
Last
<emphasis id="A5B6EAADFF85FFB1A29D23E1FF0CFC2C" italics="true" pageId="5" pageNumber="506">et al.</emphasis>
(2016)
</bibRefCitation>
, the genus now comprises 38 species. However, molecular work suggests that the monophyly of
<taxonomicName id="50C24D3CFF85FFB1A78223B9FF42FC08" authorityName="Rafinesque" authorityYear="1810" box="[151,250,981,1004]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A78223B9FF42FC08" box="[151,250,981,1004]" italics="true" pageId="5" pageNumber="506">Dipturus</emphasis>
</taxonomicName>
is compromised by the recognition of
<taxonomicName id="50C24D3CFF85FFB1A5BE23B9FCBCFC08" authorityName="Whitley" authorityYear="1939" box="[683,772,981,1004]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A5BE23B9FCBCFC08" box="[683,772,981,1004]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
as a valid genus because the latter constitutes a small clade deeply nested within a clade of
<taxonomicName id="50C24D3CFF85FFB1A5152394FDDCFBEB" authorityName="Rafinesque" authorityYear="1810" box="[512,612,1016,1039]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A5152394FDDCFBEB" box="[512,612,1016,1039]" italics="true" pageId="5" pageNumber="506">Dipturus</emphasis>
</taxonomicName>
species (e.g., Naylor
<emphasis id="A5B6EAADFF85FFB1A4492395FC2EFBF4" box="[860,918,1016,1040]" italics="true" pageId="5" pageNumber="506">et al.</emphasis>
2012b; Vargas-Caro
<emphasis id="A5B6EAADFF85FFB1A3922395FB79FBF4" box="[1159,1217,1016,1040]" italics="true" pageId="5" pageNumber="506">et al.</emphasis>
2016b). The genus
<taxonomicName id="50C24D3CFF85FFB1A7822471FF48FBD0" authorityName="Whitley" authorityYear="1939" box="[151,240,1053,1076]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A7822471FF48FBD0" box="[151,240,1053,1076]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
was established by
<bibRefCitation id="F3534B4EFF85FFB1A6C42470FDCFFBD0" author="Whitley, G." box="[465,631,1052,1077]" pageId="5" pageNumber="506" pagination="227 - 262" refId="ref14478" refString="Whitley, G. (1939) Taxonomic notes on sharks and rays. Australian Zoologist, 9 (3), 227 - 262." type="journal article" year="1939">Whitley (1939</bibRefCitation>
; page 254) for
<taxonomicName id="50C24D3CFF85FFB1A4362471FC34FBD0" baseAuthorityName="Müller &amp; Henle" baseAuthorityYear="1841" box="[803,908,1053,1076]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="A5B6EAADFF85FFB1A4362471FC34FBD0" box="[803,908,1053,1076]" italics="true" pageId="5" pageNumber="506">Z. nasuta</emphasis>
</taxonomicName>
. Beyond noting that the species “has produced snout, shouldered pectorals, surface of disc rough above and with blackish pits below. A median row of strong spines along tail, one on middle of back, and a few near the eyes”,
<bibRefCitation id="F3534B4EFF85FFB1A4992408FB80FB98" author="Whitley, G." box="[908,1080,1124,1149]" pageId="5" pageNumber="506" pagination="227 - 262" refId="ref14478" refString="Whitley, G. (1939) Taxonomic notes on sharks and rays. Australian Zoologist, 9 (3), 227 - 262." type="journal article" year="1939">Whitley (1939)</bibRefCitation>
provided no diagnosis. Despite this,
<taxonomicName id="50C24D3CFF85FFB1A7DA24E4FE90FB7B" authorityName="Whitley" authorityYear="1939" box="[207,296,1160,1183]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A7DA24E4FE90FB7B" box="[207,296,1160,1183]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
was recognized at the subgeneric rank by
<bibRefCitation id="F3534B4EFF85FFB1A41B24E5FBA1FB44" author="Last, P. R. &amp; Yearsley, G." box="[782,1049,1159,1184]" pageId="5" pageNumber="506" pagination="1627 - 1641" refId="ref13300" refString="Last, P. R. &amp; Yearsley, G. (2002) Zoogeography and relationships of Australasian skates (Chondrichthyes: Rajidae). Journal of Biogeography, 29, 1627 - 1641. https: // doi. org / 10.1046 / j. 1365 - 2699.2002.00793. x" type="journal article" year="2002">Last &amp; Yearsley (2002)</bibRefCitation>
and was subsequently returned to generic status by
<bibRefCitation id="F3534B4EFF85FFB1A64E24C1FDD9FB20" author="Last, P. R. &amp; Gledhill, D. C." box="[347,609,1196,1221]" pageId="5" pageNumber="506" pagination="45 - 65" refId="ref13251" refString="Last, P. R. &amp; Gledhill, D. C. (2007) The Maugean skate, Zearaja maugeana sp. nov. (Rajiformes: Rajidae) - a micro-endemic, Gondwanan relict from Tasmanian estuaries. Zootaxa, 1494, 45 - 65." type="journal article" year="2007">Last &amp; Gledhill (2007)</bibRefCitation>
. The latter authors provided a diagnosis and considered this genus to also include the Maugean skate (
<taxonomicName id="50C24D3CFF85FFB1A6C524BDFCE1FB0C" authority="Last &amp; Gledhill 2007" authorityName="Last &amp; Gledhill" authorityYear="2007" box="[464,857,1231,1256]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="maugeana">
<emphasis id="A5B6EAADFF85FFB1A6C524BDFDDAFB0C" box="[464,610,1233,1256]" italics="true" pageId="5" pageNumber="506">Z. maugeana</emphasis>
<bibRefCitation id="F3534B4EFF85FFB1A57D24BDFCE1FB0C" author="Last, P. R. &amp; Gledhill, D. C." box="[616,857,1231,1256]" pageId="5" pageNumber="506" pagination="45 - 65" refId="ref13251" refString="Last, P. R. &amp; Gledhill, D. C. (2007) The Maugean skate, Zearaja maugeana sp. nov. (Rajiformes: Rajidae) - a micro-endemic, Gondwanan relict from Tasmanian estuaries. Zootaxa, 1494, 45 - 65." type="journal article" year="2007">Last &amp; Gledhill 2007</bibRefCitation>
</taxonomicName>
), the
<collectingCountry id="EFD5762FFF85FFB1A48124BDFB91FB0C" box="[916,1065,1231,1256]" name="New Zealand" pageId="5" pageNumber="506">New Zealand</collectingCountry>
Rough-skate (
<taxonomicName id="50C24D3CFF85FFB1A3D924BDFA8CFB0C" baseAuthorityName="Müller &amp; Henle" baseAuthorityYear="1841" box="[1228,1332,1233,1256]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="A5B6EAADFF85FFB1A3D924BDFA8CFB0C" box="[1228,1332,1233,1256]" italics="true" pageId="5" pageNumber="506">Z. nasuta</emphasis>
</taxonomicName>
), and the Yellow-nose skate (
<taxonomicName id="50C24D3CFF85FFB1A6612499FE4BFAE8" baseAuthorityName="Guichenot" baseAuthorityYear="1848" box="[372,499,1268,1292]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="chilensis">
<emphasis id="A5B6EAADFF85FFB1A6612499FE4BFAE8" box="[372,499,1268,1292]" italics="true" pageId="5" pageNumber="506">Z. chilensis</emphasis>
</taxonomicName>
). Additionally,
<bibRefCitation id="F3534B4EFF85FFB1A5B02498FC08FAE8" author="Gabbanelli, V. &amp; Diaz de Astarloa, J. &amp; Gonzalez-Castro, M. &amp; Vazquez, D. &amp; Mabragana, E." box="[677,944,1268,1293]" pageId="5" pageNumber="506" pagination="454 - 470" refId="ref12754" refString="Gabbanelli, V., Diaz de Astarloa, J., Gonzalez-Castro, M., Vazquez, D. &amp; Mabragana, E. (2018) Almost a century of oblivion: Integrative taxonomy allows the resurrection of the longnose skate Zearaja brevicaudata (Marini, 1933) (Rajiformes; Rajidae). Comptes Rendus Biologies, 341 (9 - 10), 454 - 470. https: // doi. org / 10.1016 / j. crvi. 2018.10.002" type="journal article" year="2018">
Gabbanelli
<emphasis id="A5B6EAADFF85FFB1A4322499FCE0FAE8" box="[807,856,1268,1292]" italics="true" pageId="5" pageNumber="506">et al</emphasis>
. (2018)
</bibRefCitation>
resurrected
<taxonomicName id="50C24D3CFF85FFB1A32E2499FA24FAE8" authority="Marini 1933" authorityName="Marini" authorityYear="1933" box="[1083,1436,1268,1293]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="brevicaudata">
<emphasis id="A5B6EAADFF85FFB1A32E2499FAB2FAE8" box="[1083,1290,1268,1292]" italics="true" pageId="5" pageNumber="506">Raja brevicaudata</emphasis>
<bibRefCitation id="F3534B4EFF85FFB1A2042498FA24FAE8" author="Marini, T." box="[1297,1436,1268,1293]" pageId="5" pageNumber="506" pagination="328 - 332" refId="ref13760" refString="Marini, T. (1933) Rectificando errores ictiologicos. Physis, 11 (1933), 328 - 332." type="journal article" year="1933">Marini 1933</bibRefCitation>
</taxonomicName>
from Argentinean waters, and tentatively transferred this species to the genus
<taxonomicName id="50C24D3CFF85FFB1A3372574FBC3FACB" authorityName="Whitley" authorityYear="1939" box="[1058,1147,1304,1327]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A3372574FBC3FACB" box="[1058,1147,1304,1327]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
based largely on clasper anatomy.
</paragraph>
<paragraph id="977D36BFFF85FFB1A7D22533FD70F8E1" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="506">
The most compelling morphological feature presented by
<bibRefCitation id="F3534B4EFF85FFB1A440250DFBE1FA9C" author="Last, P. R. &amp; Gledhill, D. C." box="[853,1113,1375,1400]" pageId="5" pageNumber="506" pagination="45 - 65" refId="ref13251" refString="Last, P. R. &amp; Gledhill, D. C. (2007) The Maugean skate, Zearaja maugeana sp. nov. (Rajiformes: Rajidae) - a micro-endemic, Gondwanan relict from Tasmanian estuaries. Zootaxa, 1494, 45 - 65." type="journal article" year="2007">Last &amp; Gledhill (2007)</bibRefCitation>
that both unites members of
<taxonomicName id="50C24D3CFF85FFB1A78225E9FF48FA78" authorityName="Whitley" authorityYear="1939" box="[151,240,1413,1436]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A78225E9FF48FA78" box="[151,240,1413,1436]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
and distinguishes them from species of
<taxonomicName id="50C24D3CFF85FFB1A5A325E9FCA2FA78" authorityName="Rafinesque" authorityYear="1810" box="[694,794,1413,1436]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A5A325E9FCA2FA78" box="[694,794,1413,1436]" italics="true" pageId="5" pageNumber="506">Dipturus</emphasis>
</taxonomicName>
is their relatively simple clasper morphology. Compared with
<taxonomicName id="50C24D3CFF85FFB1A7C425C4FE8DFA5B" authorityName="Rafinesque" authorityYear="1810" box="[209,309,1448,1471]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A7C425C4FE8DFA5B" box="[209,309,1448,1471]" italics="true" pageId="5" pageNumber="506">Dipturus</emphasis>
</taxonomicName>
, the claspers of
<taxonomicName id="50C24D3CFF85FFB1A6E425C4FDF2FA5B" authorityName="Whitley" authorityYear="1939" box="[497,586,1448,1471]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A6E425C4FDF2FA5B" box="[497,586,1448,1471]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
have shorter ventral terminal and dorsal terminal 2 cartilages, a spatulated distal lobe, lack both accessory terminal cartilage 1 and the sentinel, and they represent 2432% of total length. However,
<bibRefCitation id="F3534B4EFF85FFB1A61F2583FE6EF9EC" author="Ebert, D. &amp; Compagno, L. &amp; Cowley, P." box="[266,470,1519,1544]" pageId="5" pageNumber="506" pagination="81 - 102" refId="ref12621" refString="Ebert, D., Compagno, L. &amp; Cowley, P. (2008) Aspects of the reproductive biology of skates (Chondrichthyes: Rajiformes: Rajoidei) from southern Africa. ICES Journal of Marine Science, 65, 81 - 102. https: // doi. org / 10.1093 / icesjms / fsm 169" type="journal article" year="2008">
Ebert
<emphasis id="A5B6EAADFF85FFB1A658259DFEC6F9EC" box="[333,382,1520,1544]" italics="true" pageId="5" pageNumber="506">et al</emphasis>
. (2008)
</bibRefCitation>
reported claspers of
<taxonomicName id="50C24D3CFF85FFB1A5AA259DFBBBF9EC" authority="(Wallace 1967)" baseAuthorityName="Wallace" baseAuthorityYear="1967" box="[703,1027,1519,1544]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="campbelli">
<emphasis id="A5B6EAADFF85FFB1A5AA259DFCF7F9EC" box="[703,847,1520,1544]" italics="true" pageId="5" pageNumber="506">D. campbelli</emphasis>
(Wallace 1967)
</taxonomicName>
,
<taxonomicName id="50C24D3CFF85FFB1A31B259DFB6EF9EC" baseAuthorityName="Smith" baseAuthorityYear="1964" box="[1038,1238,1520,1544]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="pullopunctatus">
<emphasis id="A5B6EAADFF85FFB1A31B259DFB6EF9EC" box="[1038,1238,1520,1544]" italics="true" pageId="5" pageNumber="506">D. pullopunctatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="50C24D3CFF85FFB1A206259DFEFEF9C9" authority="(Wallace 1967)" baseAuthorityName="Wallace" baseAuthorityYear="1967" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="springeri">
<emphasis id="A5B6EAADFF85FFB1A206259DFA24F9E3" box="[1299,1436,1520,1544]" italics="true" pageId="5" pageNumber="506">D. springeri</emphasis>
(Wallace 1967)
</taxonomicName>
to range between 20% and 32% of total length. Furthermore,
<bibRefCitation id="F3534B4EFF85FFB1A3302679FA89F9C8" author="Last, P. R. &amp; Gledhill, D. C." box="[1061,1329,1556,1581]" pageId="5" pageNumber="506" pagination="45 - 65" refId="ref13251" refString="Last, P. R. &amp; Gledhill, D. C. (2007) The Maugean skate, Zearaja maugeana sp. nov. (Rajiformes: Rajidae) - a micro-endemic, Gondwanan relict from Tasmanian estuaries. Zootaxa, 1494, 45 - 65." type="journal article" year="2007">Last &amp; Gledhill (2007)</bibRefCitation>
included several additional details of clasper morphology in their diagnosis of
<taxonomicName id="50C24D3CFF85FFB1A4B62654FC44F9AB" authorityName="Whitley" authorityYear="1939" box="[931,1020,1592,1615]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A4B62654FC44F9AB" box="[931,1020,1592,1615]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
that are also present in some species of
<taxonomicName id="50C24D3CFF85FFB1A7A92631FE98F990" authorityName="Rafinesque" authorityYear="1810" box="[188,288,1629,1652]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A7A92631FE98F990" box="[188,288,1629,1652]" italics="true" pageId="5" pageNumber="506">Dipturus</emphasis>
</taxonomicName>
. For instance, a spatulate distal end of the axial cartilage in
<taxonomicName id="50C24D3CFF85FFB1A3002631FBD6F990" authorityName="Whitley" authorityYear="1939" box="[1045,1134,1629,1652]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A3002631FBD6F990" box="[1045,1134,1629,1652]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
was also reported in
<taxonomicName id="50C24D3CFF85FFB1A2942631FE85F97C" baseAuthorityName="Smith" baseAuthorityYear="1964" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="pullopunctatus">
<emphasis id="A5B6EAADFF85FFB1A2942631FE85F97C" italics="true" pageId="5" pageNumber="506">D. pullopunctatus</emphasis>
</taxonomicName>
and
<taxonomicName id="50C24D3CFF85FFB1A69326EDFC9DF97C" authority="(Wallace 1967)" baseAuthorityName="Wallace" baseAuthorityYear="1967" box="[390,805,1663,1688]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="lanceorostratus">
<emphasis id="A5B6EAADFF85FFB1A69326EDFDD9F97C" box="[390,609,1664,1688]" italics="true" pageId="5" pageNumber="506">D. lanceorostratus</emphasis>
(Wallace 1967)
</taxonomicName>
by
<bibRefCitation id="F3534B4EFF85FFB1A4772613FBBAF97C" author="Hulley, P." box="[866,1026,1663,1688]" pageId="5" pageNumber="506" pagination="151 - 220" refId="ref12954" refString="Hulley, P. (1970) An investigation of the Rajidae of the west and south coasts of southern Africa. Annals of the South African Museum, 55 (4), 151 - 220." type="journal article" year="1970">Hulley (1970</bibRefCitation>
;
<bibRefCitation id="F3534B4EFF85FFB1A30D26ECFBECF97C" author="Hulley, P." box="[1048,1108,1664,1688]" pageId="5" pageNumber="506" pagination="1 - 103" refId="ref12992" refString="Hulley, P. (1972) The origin, interrelationship and distribution of Southern African Rajidae (Chondrichthyes, Batoidei). Annals of the South African Museum, 60 (1), 1 - 103." type="journal article" year="1972">1972</bibRefCitation>
). Moreover,
<bibRefCitation id="F3534B4EFF85FFB1A3E02613FA24F97C" author="Hulley, P." box="[1269,1436,1663,1688]" pageId="5" pageNumber="506" pagination="1 - 103" refId="ref12992" refString="Hulley, P. (1972) The origin, interrelationship and distribution of Southern African Rajidae (Chondrichthyes, Batoidei). Annals of the South African Museum, 60 (1), 1 - 103." type="journal article" year="1972">Hulley (1972)</bibRefCitation>
highlighted the absence of a sentinel in
<taxonomicName id="50C24D3CFF85FFB1A55B26C9FCAEF958" baseAuthorityName="Smith" baseAuthorityYear="1964" box="[590,790,1700,1724]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="pullopunctatus">
<emphasis id="A5B6EAADFF85FFB1A55B26C9FCAEF958" box="[590,790,1700,1724]" italics="true" pageId="5" pageNumber="506">D. pullopunctatus</emphasis>
</taxonomicName>
, despite the presence of an accessory terminal 1 cartilage. Finally, although not necessarily of generic importance, the eurybathic depth range of
<taxonomicName id="50C24D3CFF85FFB1A39526A4FB61F93B" authorityName="Whitley" authorityYear="1939" box="[1152,1241,1736,1759]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A39526A4FB61F93B" box="[1152,1241,1736,1759]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
overlaps with at least 17 species of
<taxonomicName id="50C24D3CFF85FFB1A67F2681FE75F8E0" authorityName="Rafinesque" authorityYear="1810" box="[362,461,1773,1796]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A67F2681FE75F8E0" box="[362,461,1773,1796]" italics="true" pageId="5" pageNumber="506">Dipturus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="F3534B4EFF85FFB1A51C2681FD04F8E1" author="Last, P. R. &amp; White, W. T. &amp; Carvalho, M. R. de &amp; Seret, B. &amp; Stehmann, M. F. W. &amp; Naylor, G. J. P." box="[521,700,1772,1797]" pageId="5" pageNumber="506" refId="ref13352" refString="Last, P. R., White, W. T., Carvalho, M. R. de, Seret, B., Stehmann, M. F. W. &amp; Naylor, G. J. P. (Eds.) (2016) Rays of the World. CSIRO Publishing, Melbourne, 790 pp." type="book" year="2016">
Last
<emphasis id="A5B6EAADFF85FFB1A5552681FDC9F8E0" box="[576,625,1772,1796]" italics="true" pageId="5" pageNumber="506">et al</emphasis>
. 2016
</bibRefCitation>
).
</paragraph>
<paragraph id="977D36BFFF85FFB1A7D22763FA23F838" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="506">
Given the otherwise substantial morphological similarity between the two genera, these results suggest that the simple clasper morphology seen in species assigned to
<taxonomicName id="50C24D3CFF85FFB1A4062759FCD4F8A8" authorityName="Whitley" authorityYear="1939" box="[787,876,1845,1868]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A4062759FCD4F8A8" box="[787,876,1845,1868]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
unites a subset of
<taxonomicName id="50C24D3CFF85FFB1A3522759FB13F8A8" authorityName="Rafinesque" authorityYear="1810" box="[1095,1195,1845,1868]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A3522759FB13F8A8" box="[1095,1195,1845,1868]" italics="true" pageId="5" pageNumber="506">Dipturus</emphasis>
</taxonomicName>
species but does not justify recognition of
<taxonomicName id="50C24D3CFF85FFB1A6862734FE54F88B" authorityName="Whitley" authorityYear="1939" box="[403,492,1880,1903]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A6862734FE54F88B" box="[403,492,1880,1903]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
as a genus independent of
<taxonomicName id="50C24D3CFF85FFB1A4392734FC28F88B" authorityName="Rafinesque" authorityYear="1810" box="[812,912,1880,1903]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A4392734FC28F88B" box="[812,912,1880,1903]" italics="true" pageId="5" pageNumber="506">Dipturus</emphasis>
</taxonomicName>
if the monophyly of the latter genus is to be maintained. As a consequence,
<taxonomicName id="50C24D3CFF85FFB1A6EB2711FDEFF870" authorityName="Whitley" authorityYear="1939" box="[510,599,1917,1940]" class="Elasmobranchii" family="Rajidae" genus="Zearaja" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A6EB2711FDEFF870" box="[510,599,1917,1940]" italics="true" pageId="5" pageNumber="506">Zearaja</emphasis>
</taxonomicName>
is here considered a junior synonym of
<taxonomicName id="50C24D3CFF85FFB1A3332711FB32F870" authorityName="Rafinesque" authorityYear="1810" box="[1062,1162,1917,1940]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A3332711FB32F870" box="[1062,1162,1917,1940]" italics="true" pageId="5" pageNumber="506">Dipturus</emphasis>
</taxonomicName>
. In terms of the species assigned to the genus by
<bibRefCitation id="F3534B4EFF85FFB1A6DD27CDFD63F85C" author="Last, P. R. &amp; Gledhill, D. C." box="[456,731,1952,1977]" pageId="5" pageNumber="506" pagination="45 - 65" refId="ref13251" refString="Last, P. R. &amp; Gledhill, D. C. (2007) The Maugean skate, Zearaja maugeana sp. nov. (Rajiformes: Rajidae) - a micro-endemic, Gondwanan relict from Tasmanian estuaries. Zootaxa, 1494, 45 - 65." type="journal article" year="2007">Last &amp; Gledhill (2007)</bibRefCitation>
,
<bibRefCitation id="F3534B4EFF85FFB1A5FE27CDFC05F85D" author="Last, P. R. &amp; White, W. T. &amp; Carvalho, M. R. de &amp; Seret, B. &amp; Stehmann, M. F. W. &amp; Naylor, G. J. P." box="[747,957,1952,1977]" pageId="5" pageNumber="506" refId="ref13352" refString="Last, P. R., White, W. T., Carvalho, M. R. de, Seret, B., Stehmann, M. F. W. &amp; Naylor, G. J. P. (Eds.) (2016) Rays of the World. CSIRO Publishing, Melbourne, 790 pp." type="book" year="2016">
Last
<emphasis id="A5B6EAADFF85FFB1A43227CDFCE5F85C" box="[807,861,1952,1976]" italics="true" pageId="5" pageNumber="506">et al</emphasis>
. (2016)
</bibRefCitation>
, and
<bibRefCitation id="F3534B4EFF85FFB1A31727CCFAA7F85C" author="Gabbanelli, V. &amp; Diaz de Astarloa, J. &amp; Gonzalez-Castro, M. &amp; Vazquez, D. &amp; Mabragana, E." box="[1026,1311,1952,1977]" pageId="5" pageNumber="506" pagination="454 - 470" refId="ref12754" refString="Gabbanelli, V., Diaz de Astarloa, J., Gonzalez-Castro, M., Vazquez, D. &amp; Mabragana, E. (2018) Almost a century of oblivion: Integrative taxonomy allows the resurrection of the longnose skate Zearaja brevicaudata (Marini, 1933) (Rajiformes; Rajidae). Comptes Rendus Biologies, 341 (9 - 10), 454 - 470. https: // doi. org / 10.1016 / j. crvi. 2018.10.002" type="journal article" year="2018">
Gabbanelli
<emphasis id="A5B6EAADFF85FFB1A39C27CDFB07F85C" box="[1161,1215,1952,1976]" italics="true" pageId="5" pageNumber="506">et al</emphasis>
. (2018)
</bibRefCitation>
, following
<bibRefCitation id="F3534B4EFF85FFB1A78227A8FE6AF838" author="McEachran, J. &amp; Dunn, K." box="[151,466,1988,2013]" pageId="5" pageNumber="506" pagination="271 - 290" refId="ref13782" refString="McEachran, J. &amp; Dunn, K. (1998) Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia, 1998 (2), 271 - 290. https: // doi. org / 10.2307 / 1447424" type="journal article" year="1998">McEachran &amp; Dunn (1998)</bibRefCitation>
,
<taxonomicName id="50C24D3CFF85FFB1A6CA27A9FDE0F838" baseAuthorityName="Müller &amp; Henle" baseAuthorityYear="1841" box="[479,600,1989,2012]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="nasutus">
<emphasis id="A5B6EAADFF85FFB1A6CA27A9FDE0F838" box="[479,600,1989,2012]" italics="true" pageId="5" pageNumber="506">D. nasutus</emphasis>
</taxonomicName>
and
<taxonomicName id="50C24D3CFF85FFB1A58727A9FCA0F838" baseAuthorityName="Guichenot" baseAuthorityYear="1848" box="[658,792,1988,2012]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="chilensis">
<emphasis id="A5B6EAADFF85FFB1A58727A9FD1EF838" box="[658,678,1989,2012]" italics="true" pageId="5" pageNumber="506">D</emphasis>
.
<emphasis id="A5B6EAADFF85FFB1A5A327A8FCA0F838" box="[694,792,1988,2012]" italics="true" pageId="5" pageNumber="506">chilensis</emphasis>
</taxonomicName>
are considered members of
<taxonomicName id="50C24D3CFF85FFB1A34927A9FB78F838" authorityName="Rafinesque" authorityYear="1810" box="[1116,1216,1989,2012]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="A5B6EAADFF85FFB1A34927A9FB78F838" box="[1116,1216,1989,2012]" italics="true" pageId="5" pageNumber="506">Dipturus</emphasis>
</taxonomicName>
and
<taxonomicName id="50C24D3CFF85FFB1A3EC27A9FA23F838" box="[1273,1435,1989,2012]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="maugeanus">
<emphasis id="A5B6EAADFF85FFB1A3EC27A9FA23F838" box="[1273,1435,1989,2012]" italics="true" pageId="5" pageNumber="506">D. maugeanus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="977D36BFFF86FFB2A78220F4FB30FF31" blockId="6.[151,1436,151,213]" pageId="6" pageNumber="507">
(
<bibRefCitation id="F3534B4EFF86FFB2A78B20F4FE27FF54" author="Last, P. R. &amp; Gledhill, D. C." box="[158,415,151,176]" pageId="6" pageNumber="507" pagination="45 - 65" refId="ref13251" refString="Last, P. R. &amp; Gledhill, D. C. (2007) The Maugean skate, Zearaja maugeana sp. nov. (Rajiformes: Rajidae) - a micro-endemic, Gondwanan relict from Tasmanian estuaries. Zootaxa, 1494, 45 - 65." type="journal article" year="2007">Last &amp; Gledhill, 2007</bibRefCitation>
)
<emphasis id="A5B6EAADFF86FFB2A6BA20FBFDAFFF54" bold="true" box="[431,535,151,176]" pageId="6" pageNumber="507">n. comb.</emphasis>
,
<taxonomicName id="50C24D3CFF86FFB2A53220F5FAC4FF54" authority="(Diaz de Astarloa, Mabragana, Hanner &amp; Figueroa 2008)" authorityName="Concha &amp; Caira &amp; Ebert &amp; Pompert" authorityYear="2019" baseAuthorityName="Diaz de Astarloa, Mabragana, Hanner &amp; Figueroa" baseAuthorityYear="2008" box="[551,1404,151,176]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="6" pageNumber="507" phylum="Chordata" rank="species" species="argentinensis" status="comb. nov.">
<emphasis id="A5B6EAADFF86FFB2A53220F5FD5AFF4B" box="[551,738,152,176]" italics="true" pageId="6" pageNumber="507">D. argentinensis</emphasis>
(
<bibRefCitation id="F3534B4EFF86FFB2A5E720F4FACDFF54" author="Diaz de Astarloa, J. &amp; Mabragana, E. &amp; Hanner, R. &amp; Figueroa, D." box="[754,1397,151,176]" pageId="6" pageNumber="507" pagination="35 - 46" refId="ref12510" refString="Diaz de Astarloa, J., Mabragana, E., Hanner, R. &amp; Figueroa, D. (2008) Morphological and molecular evidence for a new species of longnose skate (Rajiformes: Rajidae: Dipturus) from Argentinean waters based on DNA barcoding. Zootaxa, 1291, 35 - 46." type="journal article" year="2008">Díaz de Astarloa, Mabragaña, Hanner &amp; Figueroa 2008</bibRefCitation>
)
</taxonomicName>
<emphasis id="A5B6EAADFF86FFB2A29020FBFF67FF31" bold="true" pageId="6" pageNumber="507">
<taxonomicNameLabel id="BE8557D6FF86FFB2A29020FBFF67FF31" pageId="6" pageNumber="507" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="50C24D3CFF86FFB2A60920D1FD39FF30" authority="(Marini 1933)" authorityName="Concha &amp; Caira &amp; Ebert &amp; Pompert" authorityYear="2019" baseAuthorityName="Marini" baseAuthorityYear="1933" box="[284,641,188,213]" class="Elasmobranchii" family="Rajidae" genus="Dipturus" kingdom="Animalia" order="Rajiformes" pageId="6" pageNumber="507" phylum="Chordata" rank="species" species="brevicaudatus" status="comb. nov.">
<emphasis id="A5B6EAADFF86FFB2A60920D1FE64FF30" box="[284,476,188,212]" italics="true" pageId="6" pageNumber="507">D. brevicaudatus</emphasis>
(
<bibRefCitation id="F3534B4EFF86FFB2A6FF20D0FDC1FF30" author="Marini, T." box="[490,633,188,213]" pageId="6" pageNumber="507" pagination="328 - 332" refId="ref13760" refString="Marini, T. (1933) Rectificando errores ictiologicos. Physis, 11 (1933), 328 - 332." type="journal article" year="1933">Marini 1933</bibRefCitation>
)
</taxonomicName>
<emphasis id="A5B6EAADFF86FFB2A59D20D0FD56FF31" bold="true" box="[648,750,188,213]" pageId="6" pageNumber="507">
<taxonomicNameLabel id="BE8557D6FF86FFB2A59D20D0FD56FF31" box="[648,750,188,213]" pageId="6" pageNumber="507" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
are hereby transferred to that genus.
</paragraph>
</subSubSection>
</treatment>
</document>