216 lines
31 KiB
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216 lines
31 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.79.e62735" ID-GBIF-Dataset="2fae9b0a-ea4d-4715-9ee6-93a215740e7c" ID-GBIF-Taxon="182473841" ID-Pensoft-Pub="1864-8312-79-151" ID-Pensoft-UUID="8842F15397AF5A1BBD35D4ECB5A874CE" ID-ZooBank="414052252BB34048A62903AF1569C616" LSID="urn:lsid:zoobank.org:act:DFE52BBC-7153-4EA3-89B5-403D069BF5EE" ModsDocID="1864-8312-79-151" checkinTime="1622492420826" checkinUser="pensoft" docAuthor="Molineri, Carlos, Dias, Lucimar G. & Zuniga, Maria del Carmen" docDate="2021" docId="19FAC2E7F4AA5F7FAC79CF7EFE9A2E05" docLanguage="en" docName="ArthropodSystPhyl 79: 151-170" docOrigin="Arthropod Systematics & amp; Phylogeny 79" docPubDate="2021-05-31" docSource="http://dx.doi.org/10.3897/asp.79.e62735" docTitle="Tortopsis andaki Molineri & Dias & Zúñiga 2021, sp. nov." docType="treatment" docUuid="DFE52BBC-7153-4EA3-89B5-403D069BF5EE" docUuidSource="ZooBank" docVersion="3" id="8842F15397AF5A1BBD35D4ECB5A874CE" lastPageNumber="151" masterDocId="8842F15397AF5A1BBD35D4ECB5A874CE" masterDocTitle="New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species" masterLastPageNumber="170" masterPageNumber="151" pageId="0" pageNumber="151" updateTime="1643426787942" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>New insights into the phylogeny of Tortopus Needham and Murphy and Tortopsis Molineri (Ephemeroptera, Polymitarcyidae) with description of three new species</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Molineri, Carlos</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2662-624X</mods:nameIdentifier>
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<mods:affiliation>Instituto de Biodiversidad Neotropical, CONICET (National Council of Scientific Research), Universidad Nacional de Tucuman, Ciudad Universitaria Horco Molle, CP 4000, Argentina; Carlos Molineri [carlosmolineri @ gmail. com]</mods:affiliation>
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<mods:nameIdentifier type="email">carlosmolineri@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dias, Lucimar G.</mods:namePart>
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<mods:affiliation>Facultad de Ciencias Exactas y Naturales, Grupo de Investigacion BIONAT, Universidad de Caldas, Caldas, Colombia; Lucimar G. Dias [lucimar. dias @ ucaldas. edu. co]</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zuniga, Maria del Carmen</mods:namePart>
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<mods:affiliation>Grupo de Investigaciones Entomologicas, Departamento de Biologia, Universidad del Valle, Cali, Colombia; Maria del Carmen Zuniga [maczuniga @ gmail. com]</mods:affiliation>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-05-31</mods:number>
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<mods:detail type="volume">
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<mods:number>79</mods:number>
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<mods:start>151</mods:start>
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<mods:url>http://dx.doi.org/10.3897/asp.79.e62735</mods:url>
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<mods:classification>journal article</mods:classification>
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<treatment ID-GBIF-Taxon="182473841" LSID="urn:lsid:zoobank.org:act:DFE52BBC-7153-4EA3-89B5-403D069BF5EE" httpUri="http://treatment.plazi.org/id/19FAC2E7F4AA5F7FAC79CF7EFE9A2E05" lastPageNumber="151" pageId="0" pageNumber="151">
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<subSubSection pageId="0" pageNumber="151" type="nomenclature">
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<paragraph pageId="0" pageNumber="151">
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<taxonomicName LSID="https://zoobank.org/DFE52BBC-7153-4EA3-89B5-403D069BF5EE" authority="Molineri & Dias & Zúñiga, 2021" authorityName="Molineri & Dias & Zúñiga" authorityYear="2021" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortopsis andaki" order="Ephemeroptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="andaki" status="sp. nov.">Tortopsis andaki</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="151">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">Figs 27-32</figureCitation>
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<figureCitation captionStart="Figures 33–40" captionStartId="F6" captionText="Figures 33 - 40. Tortopsis andaki sp. nov., imago. Male: 33, male genitalia, ventral view; 34, genitalia, lateral external view; 35, same, inner view; 36, penis detail, dorsal view; 37, penis, detail, ventral view; 38, details of penis apex, dorsal view above, ventral view below. Female: 39, abdominal sternum VIII, lateral view; 40, same, ventral view." figureDoi="10.3897/asp.79.e62735.figures33-40" httpUri="https://binary.pensoft.net/fig/550222" pageId="0" pageNumber="151">, 33-40</figureCitation>
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<figureCitation captionStart="Figures 41–52" captionStartId="F7" captionText="Figures 41 - 52. Tortopsis andaki sp. nov., female nymph. 41, labrum, dorsal view; 42, left mandible, ventral view; 43, right mandible, ventral view; 44, same, detail (arrows indicate first and last spine of the inner marginal row); 45, hypopharynx, ventral view; 46, labium, ventral view; 47, right maxilla, ventral view (arrow indicates membranous gill); 48, left maxilla, ventral view; 49, fore leg, dorsal view; 50, detail of fore tarsal claw, dorsal view; 51, middle leg, dorsal view; 53, hind leg, dorsal view." figureDoi="10.3897/asp.79.e62735.figures41-52" httpUri="https://binary.pensoft.net/fig/550223" pageId="0" pageNumber="151">, 41-52</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="151" type="type material">
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<paragraph pageId="0" pageNumber="151">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="151">
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Holotype male imago (CEBUC) from Colombia,
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<normalizedToken originalValue="Caquetá">Caqueta</normalizedToken>
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, Curillo,
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<normalizedToken originalValue="río">rio</normalizedToken>
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<normalizedToken originalValue="Caquetá">Caqueta</normalizedToken>
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, 220 m,
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<geoCoordinate degrees="01" direction="north" minutes="01" orientation="latitude" precision="15" seconds="44" value="1.0288888">
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01°01
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<normalizedToken originalValue="’44”">'44"</normalizedToken>
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N
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</geoCoordinate>
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;
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<geoCoordinate degrees="75" direction="west" minutes="55" orientation="longitude" precision="15" seconds="9" value="-75.91917">
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75°55
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<normalizedToken originalValue="’9”">'9"</normalizedToken>
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W
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</geoCoordinate>
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, 20.ix.2018, LG Dias, MC
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<normalizedToken originalValue="Zúñiga">Zuniga</normalizedToken>
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, B Toro, JP Chaux and C Molineri cols. Paratypes: 14 male and 5 female imagos (IBN). Paratypes: 10 male 5 female imagos and 1 nymph (CEBUC); 10 male and 2 female imagos (MUSENUV), same data as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="151" type="diagnosis">
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<paragraph pageId="0" pageNumber="151">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="151">
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<taxonomicName authorityName="Molineri & Dias & Zúñiga" authorityYear="2021" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortopsis andaki" order="Ephemeroptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="andaki">
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<emphasis italics="true" pageId="0" pageNumber="151">Tortopsis andaki</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="151">sp. nov.</emphasis>
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, known from imagos of both sexes and nymph, can be distinguished from all other species of the genus by: In adults, 1) fore wing length 9.2-9.6 mm (male), 9.5-9.7 mm (female); 2) wings hyaline slightly tinted with purplish gray, veins purplish gray; 3) parastylus curved dorsally, more markedly on apical third, with a longitudinal ventral furrow (Figs
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<figureCitation captionStart="Figures 33–40" captionStartId="F6" captionText="Figures 33 - 40. Tortopsis andaki sp. nov., imago. Male: 33, male genitalia, ventral view; 34, genitalia, lateral external view; 35, same, inner view; 36, penis detail, dorsal view; 37, penis, detail, ventral view; 38, details of penis apex, dorsal view above, ventral view below. Female: 39, abdominal sternum VIII, lateral view; 40, same, ventral view." figureDoi="10.3897/asp.79.e62735.figures33-40" httpUri="https://binary.pensoft.net/fig/550222" pageId="0" pageNumber="151">33-34</figureCitation>
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); 4) penis slender, apical spine rounded and flattened (Figs
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<figureCitation captionStart="Figures 33–40" captionStartId="F6" captionText="Figures 33 - 40. Tortopsis andaki sp. nov., imago. Male: 33, male genitalia, ventral view; 34, genitalia, lateral external view; 35, same, inner view; 36, penis detail, dorsal view; 37, penis, detail, ventral view; 38, details of penis apex, dorsal view above, ventral view below. Female: 39, abdominal sternum VIII, lateral view; 40, same, ventral view." figureDoi="10.3897/asp.79.e62735.figures33-40" httpUri="https://binary.pensoft.net/fig/550222" pageId="0" pageNumber="151">36-38</figureCitation>
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); 5) parastyli receptors on sternum VIII with V-shaped sockets (Figs
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<figureCitation captionStart="Figures 33–40" captionStartId="F6" captionText="Figures 33 - 40. Tortopsis andaki sp. nov., imago. Male: 33, male genitalia, ventral view; 34, genitalia, lateral external view; 35, same, inner view; 36, penis detail, dorsal view; 37, penis, detail, ventral view; 38, details of penis apex, dorsal view above, ventral view below. Female: 39, abdominal sternum VIII, lateral view; 40, same, ventral view." figureDoi="10.3897/asp.79.e62735.figures33-40" httpUri="https://binary.pensoft.net/fig/550222" pageId="0" pageNumber="151">39-40</figureCitation>
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); 6) head shaded gray among ocelli, occiput with small gray marks (Fig.
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<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">27</figureCitation>
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). The nymphs are characterized by: 1) mandibles with 7 stout small spines on inner margin of tusk see arrows in Fig.
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<figureCitation captionStart="Figures 41–52" captionStartId="F7" captionText="Figures 41 - 52. Tortopsis andaki sp. nov., female nymph. 41, labrum, dorsal view; 42, left mandible, ventral view; 43, right mandible, ventral view; 44, same, detail (arrows indicate first and last spine of the inner marginal row); 45, hypopharynx, ventral view; 46, labium, ventral view; 47, right maxilla, ventral view (arrow indicates membranous gill); 48, left maxilla, ventral view; 49, fore leg, dorsal view; 50, detail of fore tarsal claw, dorsal view; 51, middle leg, dorsal view; 53, hind leg, dorsal view." figureDoi="10.3897/asp.79.e62735.figures41-52" httpUri="https://binary.pensoft.net/fig/550223" pageId="0" pageNumber="151">44</figureCitation>
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;
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<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Tortopsis toro sp. nov. Male imago: 1, general view, dorsal; 2, abdominal segments VIII-X, lateral view; 3, genitalia, ventral view. Nymph: 4, mature female nymph; 5, mature male nymph; 6, detail of head and pronotal coloration of not fully-grown male nymph." figureDoi="10.3897/asp.79.e62735.figures1-6" httpUri="https://binary.pensoft.net/fig/550217" pageId="0" pageNumber="151">2</figureCitation>
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) occiput whitish without gray markings (Fig.
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<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">31</figureCitation>
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); 3) wingbuds completely whitish (Fig.
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<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">32</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="151">
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<emphasis italics="true" pageId="0" pageNumber="151">Male imago</emphasis>
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. Length (mm): body, 10.8-11.0; fore wing, 9.2-9.6; hind wing, 4.0; foreleg, 4.9-5.0; cerci, 25.0-26.0. General coloration whitish shaded dorsally with light purplish gray (Figs
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<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">27</figureCitation>
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). Head. Whitish shaded with light brownish gray among ocelli and on scape and pedicel, flagellum hyaline. Thorax. Pronotum hyaline shaded with purplish gray dorsally as in Fig.
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<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">27</figureCitation>
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. Mesonotum whitish yellow shaded with purplish gray along medioparapsidal sutures and contiguous area, medial zone paler. Metanotum whitish yellow shaded gray dorsally. Thoracic pleura and sterna whitish. Legs whitish shaded with light purplish gray on fore tibia and fore tarsi. Wings: Membrane hyaline very lightly tinted with purplish gray, longitudinal and cross veins purplish gray. Abdomen. Translucent white, shaded slightly but extensively with purplish gray on terga, including medial area; some markings are darker (Fig.
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<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">27</figureCitation>
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) and a thin medial dark line is present along most terga; terga VIII-X shaded with darker gray. Abdominal sterna whitish; sternum IX shaded very slightly with gray on median area. Genitalia (Figs
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<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">28-29</figureCitation>
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,
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<figureCitation captionStart="Figures 33–40" captionStartId="F6" captionText="Figures 33 - 40. Tortopsis andaki sp. nov., imago. Male: 33, male genitalia, ventral view; 34, genitalia, lateral external view; 35, same, inner view; 36, penis detail, dorsal view; 37, penis, detail, ventral view; 38, details of penis apex, dorsal view above, ventral view below. Female: 39, abdominal sternum VIII, lateral view; 40, same, ventral view." figureDoi="10.3897/asp.79.e62735.figures33-40" httpUri="https://binary.pensoft.net/fig/550222" pageId="0" pageNumber="151">33-38</figureCitation>
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): whitish except apex of parastylus yellowish; parastylus curved dorsally more markedly on apical third (in lateral view, Figs
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<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">28</figureCitation>
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and
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<figureCitation captionStart="Figures 33–40" captionStartId="F6" captionText="Figures 33 - 40. Tortopsis andaki sp. nov., imago. Male: 33, male genitalia, ventral view; 34, genitalia, lateral external view; 35, same, inner view; 36, penis detail, dorsal view; 37, penis, detail, ventral view; 38, details of penis apex, dorsal view above, ventral view below. Female: 39, abdominal sternum VIII, lateral view; 40, same, ventral view." figureDoi="10.3897/asp.79.e62735.figures33-40" httpUri="https://binary.pensoft.net/fig/550222" pageId="0" pageNumber="151">34-35</figureCitation>
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) and with a longitudinal ventral furrow along its entire length (Fig.
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<figureCitation captionStart="Figures 33–40" captionStartId="F6" captionText="Figures 33 - 40. Tortopsis andaki sp. nov., imago. Male: 33, male genitalia, ventral view; 34, genitalia, lateral external view; 35, same, inner view; 36, penis detail, dorsal view; 37, penis, detail, ventral view; 38, details of penis apex, dorsal view above, ventral view below. Female: 39, abdominal sternum VIII, lateral view; 40, same, ventral view." figureDoi="10.3897/asp.79.e62735.figures33-40" httpUri="https://binary.pensoft.net/fig/550222" pageId="0" pageNumber="151">33</figureCitation>
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); forceps and penis translucent white, shaded very slightly gray on forceps; penis slender, spine at apex of penis rounded and flattened (Figs
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<figureCitation captionStart="Figures 33–40" captionStartId="F6" captionText="Figures 33 - 40. Tortopsis andaki sp. nov., imago. Male: 33, male genitalia, ventral view; 34, genitalia, lateral external view; 35, same, inner view; 36, penis detail, dorsal view; 37, penis, detail, ventral view; 38, details of penis apex, dorsal view above, ventral view below. Female: 39, abdominal sternum VIII, lateral view; 40, same, ventral view." figureDoi="10.3897/asp.79.e62735.figures33-40" httpUri="https://binary.pensoft.net/fig/550222" pageId="0" pageNumber="151">36-38</figureCitation>
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). Caudal filament whitish translucent.
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</paragraph>
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<caption doi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151" start="Figures 27–32" startId="F5">
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<paragraph pageId="0" pageNumber="151">
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<emphasis bold="true" pageId="0" pageNumber="151">Figures 27-32.</emphasis>
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<taxonomicName authorityName="Molineri & Dias & Zúñiga" authorityYear="2021" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortopsis andaki" order="Ephemeroptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="andaki">
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<emphasis italics="true" pageId="0" pageNumber="151">Tortopsis andaki</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="151">sp. nov.</emphasis>
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, male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="151">
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Female imago. Length (mm): body, 10.0-10.1; fore wing, 9.5-9.7; hind wing, 3.9-4.0; cerci, 3.0. General coloration as in male but shading more strongly marked. Head black between ocelli, with small light gray marks on occiput. Wings membrane hyaline slightly tinged with whitish yellow, veins brownish. Abdomen shaded with gray on terga. Parastyli receptors on sternum VIII with sockets (Figs
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<figureCitation captionStart="Figures 33–40" captionStartId="F6" captionText="Figures 33 - 40. Tortopsis andaki sp. nov., imago. Male: 33, male genitalia, ventral view; 34, genitalia, lateral external view; 35, same, inner view; 36, penis detail, dorsal view; 37, penis, detail, ventral view; 38, details of penis apex, dorsal view above, ventral view below. Female: 39, abdominal sternum VIII, lateral view; 40, same, ventral view." figureDoi="10.3897/asp.79.e62735.figures33-40" httpUri="https://binary.pensoft.net/fig/550222" pageId="0" pageNumber="151">39-40</figureCitation>
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) with sinuous lateral margin and acute distal corner (V-shaped).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="151">
|
||
Nymph (mature female). Length: body, 17.0 mm. General coloration yellowish white shaded with gray dorsally (Fig.
|
||
<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">30</figureCitation>
|
||
). Head. Whitish shaded with gray among ocelli, occiput completely white without marks (Fig.
|
||
<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">31</figureCitation>
|
||
). Antennae and mouthparts whitish except apex of tusks brownish, and spines yellowish. Finger-like gill present near base of maxillae (Figs
|
||
<figureCitation captionStart="Figures 41–52" captionStartId="F7" captionText="Figures 41 - 52. Tortopsis andaki sp. nov., female nymph. 41, labrum, dorsal view; 42, left mandible, ventral view; 43, right mandible, ventral view; 44, same, detail (arrows indicate first and last spine of the inner marginal row); 45, hypopharynx, ventral view; 46, labium, ventral view; 47, right maxilla, ventral view (arrow indicates membranous gill); 48, left maxilla, ventral view; 49, fore leg, dorsal view; 50, detail of fore tarsal claw, dorsal view; 51, middle leg, dorsal view; 53, hind leg, dorsal view." figureDoi="10.3897/asp.79.e62735.figures41-52" httpUri="https://binary.pensoft.net/fig/550223" pageId="0" pageNumber="151">47-48</figureCitation>
|
||
). Mandibular tusks with 7 small stout spines on inner margin, basal to subdistal tubercle (see arrows in Fig.
|
||
<figureCitation captionStart="Figures 41–52" captionStartId="F7" captionText="Figures 41 - 52. Tortopsis andaki sp. nov., female nymph. 41, labrum, dorsal view; 42, left mandible, ventral view; 43, right mandible, ventral view; 44, same, detail (arrows indicate first and last spine of the inner marginal row); 45, hypopharynx, ventral view; 46, labium, ventral view; 47, right maxilla, ventral view (arrow indicates membranous gill); 48, left maxilla, ventral view; 49, fore leg, dorsal view; 50, detail of fore tarsal claw, dorsal view; 51, middle leg, dorsal view; 53, hind leg, dorsal view." figureDoi="10.3897/asp.79.e62735.figures41-52" httpUri="https://binary.pensoft.net/fig/550223" pageId="0" pageNumber="151">44</figureCitation>
|
||
); outer margin of tusk with row of 16-17 stout spines (Figs
|
||
<figureCitation captionStart="Figures 41–52" captionStartId="F7" captionText="Figures 41 - 52. Tortopsis andaki sp. nov., female nymph. 41, labrum, dorsal view; 42, left mandible, ventral view; 43, right mandible, ventral view; 44, same, detail (arrows indicate first and last spine of the inner marginal row); 45, hypopharynx, ventral view; 46, labium, ventral view; 47, right maxilla, ventral view (arrow indicates membranous gill); 48, left maxilla, ventral view; 49, fore leg, dorsal view; 50, detail of fore tarsal claw, dorsal view; 51, middle leg, dorsal view; 53, hind leg, dorsal view." figureDoi="10.3897/asp.79.e62735.figures41-52" httpUri="https://binary.pensoft.net/fig/550223" pageId="0" pageNumber="151">42-44</figureCitation>
|
||
). Thorax. Pronotum, anterior ring shaded gray widely including the anterolateral pointed projections, posterior ring as in Fig.
|
||
<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">30</figureCitation>
|
||
, both with a median pale line. Meso- and metanotum with gray and brown marks, lighter on median area; wing buds whitish, longitudinal veins hyaline, except at the basal region, with the anal and costal margins brownish. Thoracic pleura and sterna whitish. Legs whitish, with yellowish setae and apex of tarsal claws yellowish (Figs
|
||
<figureCitation captionStart="Figures 41–52" captionStartId="F7" captionText="Figures 41 - 52. Tortopsis andaki sp. nov., female nymph. 41, labrum, dorsal view; 42, left mandible, ventral view; 43, right mandible, ventral view; 44, same, detail (arrows indicate first and last spine of the inner marginal row); 45, hypopharynx, ventral view; 46, labium, ventral view; 47, right maxilla, ventral view (arrow indicates membranous gill); 48, left maxilla, ventral view; 49, fore leg, dorsal view; 50, detail of fore tarsal claw, dorsal view; 51, middle leg, dorsal view; 53, hind leg, dorsal view." figureDoi="10.3897/asp.79.e62735.figures41-52" httpUri="https://binary.pensoft.net/fig/550223" pageId="0" pageNumber="151">49-52</figureCitation>
|
||
). Abdomen. Abdominal color pattern as described for the adults (Fig.
|
||
<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">30</figureCitation>
|
||
). Gills: vestigial gills I translucent; gills II-VII well developed, formed by a pair of large whitish lamellae, the outer (dorsal) lamellae of each pair is shaded with gray on a medio longitudinal band (Fig.
|
||
<figureCitation captionStart="Figures 27–32" captionStartId="F5" captionText="Figures 27 - 32. Tortopsis andaki sp. nov., male imago. 27, general dorsal view; 28, genitalia, lateral view; 29, genitalia, ventral view. Nymph (female): 30, general view; 31, head and pronotum; 32, part of mesonotum and wingbuds." figureDoi="10.3897/asp.79.e62735.figures27-32" httpUri="https://binary.pensoft.net/fig/550221" pageId="0" pageNumber="151">30</figureCitation>
|
||
), the inner (ventral) lamellae of each pair present only a thin brownish line along trachea. Caudal filaments yellowish white.
|
||
</paragraph>
|
||
<caption doi="10.3897/asp.79.e62735.figures33-40" httpUri="https://binary.pensoft.net/fig/550222" pageId="0" pageNumber="151" start="Figures 33–40" startId="F6">
|
||
<paragraph pageId="0" pageNumber="151">
|
||
<emphasis bold="true" pageId="0" pageNumber="151">Figures 33-40.</emphasis>
|
||
<taxonomicName authorityName="Molineri & Dias & Zúñiga" authorityYear="2021" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortopsis andaki" order="Ephemeroptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="andaki">
|
||
<emphasis italics="true" pageId="0" pageNumber="151">Tortopsis andaki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="151">sp. nov.</emphasis>
|
||
, imago. Male: 33, male genitalia, ventral view; 34, genitalia, lateral external view; 35, same, inner view; 36, penis detail, dorsal view; 37, penis, detail, ventral view; 38, details of penis apex, dorsal view above, ventral view below. Female: 39, abdominal sternum VIII, lateral view; 40, same, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="151">
|
||
Egg. Subcircular, white. Length, 330-350
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
; width, 280-300
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="151" type="etymology">
|
||
<paragraph pageId="0" pageNumber="151">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="151">
|
||
This species is dedicated to the Andaki indigenous people, an American ethnic group that inhabited the upper
|
||
<normalizedToken originalValue="Caquetá">Caqueta</normalizedToken>
|
||
River basin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="151" type="distribution">
|
||
<paragraph pageId="0" pageNumber="151">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="151">Known only from the type locality.</paragraph>
|
||
<caption doi="10.3897/asp.79.e62735.figures41-52" httpUri="https://binary.pensoft.net/fig/550223" pageId="0" pageNumber="151" start="Figures 41–52" startId="F7">
|
||
<paragraph pageId="0" pageNumber="151">
|
||
<emphasis bold="true" pageId="0" pageNumber="151">Figures 41-52.</emphasis>
|
||
<taxonomicName authorityName="Molineri & Dias & Zúñiga" authorityYear="2021" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortopsis andaki" order="Ephemeroptera" pageId="0" pageNumber="151" phylum="Arthropoda" rank="species" species="andaki">
|
||
<emphasis italics="true" pageId="0" pageNumber="151">Tortopsis andaki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="151">sp. nov.</emphasis>
|
||
, female nymph. 41, labrum, dorsal view; 42, left mandible, ventral view; 43, right mandible, ventral view; 44, same, detail (arrows indicate first and last spine of the inner marginal row); 45, hypopharynx, ventral view; 46, labium, ventral view; 47, right maxilla, ventral view (arrow indicates membranous gill); 48, left maxilla, ventral view; 49, fore leg, dorsal view; 50, detail of fore tarsal claw, dorsal view; 51, middle leg, dorsal view; 53, hind leg, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |