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<document id="38ABA5D7728EF798D789E5637605C7D5" ID-CLB-Dataset="24105" ID-DOI="10.5852/ejt.2019.585" ID-GBIF-Dataset="dca608e8-fccf-4c1c-b8df-ef0c28e1d518" ID-Zenodo-Dep="3660259" ID-ZooBank="181B6FBA-ED75-4BB4-84C4-FB512B794749" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581251975682" checkinUser="plazi" docAuthor="Ebersole, Jun A., Cicimurri, David J. &amp; Stringer, Gary L." docDate="2019" docId="18174D41FF35FF1EFD7E9D154CF00D42" docLanguage="en" docName="ejt-585_ebersole_cicimurri_stinger.pdf.imf" docOrigin="European Journal of Taxonomy 585" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gymnosarda Gill 1862" docType="treatment" docVersion="19" lastPageNumber="178" masterDocId="E42E3539FF85FFAFFFC69F344D4B0F37" masterDocTitle="Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths" masterLastPageNumber="274" masterPageNumber="1" pageNumber="177" updateTime="1698783570407" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2107223BA7E9C6AEACBBD62D4C12A395">Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths</mods:title>
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<subSubSection id="D8A4AFDCFF35FF1FFD7E9D154E300D0B" box="[696,891,545,572]" pageId="176" pageNumber="177" type="nomenclature">
<paragraph id="9001FC57FF35FF1FFD7E9D154E300D0B" blockId="176.[696,891,545,607]" box="[696,891,545,572]" pageId="176" pageNumber="177">
<heading id="CB494B3BFF35FF1FFD7E9D154E300D0B" box="[696,891,545,572]" centered="true" fontSize="11" level="2" pageId="176" pageNumber="177" reason="2">
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<emphasis id="A2CA2045FF35FF1FFD7E9D154E180D0B" bold="true" box="[696,851,545,572]" italics="true" pageId="176" pageNumber="177">Gymnosarda</emphasis>
</taxonomicName>
sp.
</heading>
</paragraph>
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<subSubSection id="D8A4AFDCFF35FF1FFD369D704E080D68" box="[752,835,580,607]" pageId="176" pageNumber="177" type="description">
<paragraph id="9001FC57FF35FF1FFD369D704E080D68" blockId="176.[696,891,545,607]" box="[752,835,580,607]" pageId="176" pageNumber="177">
<figureCitation id="0885E0D2FF35FF1FFD369D704E080D68" box="[752,835,580,607]" captionStart="Fig" captionStartId="176.[189,232,1964,1990]" captionTargetBox="[202,1315,1404,1924]" captionTargetId="figure@176.[189,1398,1400,1927]" captionTargetPageId="176" captionText="Fig. 65. Gymnosarda sp., hypural plate. AC. MSC 37271, basal Lisbon Formation. A. Right lateral view. B. Anterior view. C. Dorsal view. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3660385" httpUri="https://zenodo.org/record/3660385/files/figure.png" pageId="176" pageNumber="177">Fig. 65</figureCitation>
</paragraph>
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<subSubSection id="D8A4AFDCFF35FF1FFF7B9DB248630DF8" pageId="176" pageNumber="177" type="materials_examined">
<paragraph id="9001FC57FF35FF1FFF7B9DB24CE10D97" blockId="176.[189,426,646,672]" box="[189,426,646,672]" pageId="176" pageNumber="177">
<heading id="CB494B3BFF35FF1FFF7B9DB24CE10D97" bold="true" box="[189,426,646,672]" fontSize="11" level="3" pageId="176" pageNumber="177" reason="3">
<emphasis id="A2CA2045FF35FF1FFF7B9DB24CE10D97" bold="true" box="[189,426,646,672]" pageId="176" pageNumber="177">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FF35FF1FFF7B9D8148630DF8" blockId="176.[189,1320,692,719]" box="[189,1320,692,719]" pageId="176" pageNumber="177">
<materialsCitation id="20D6F60AFF35FF1FFF7B9D81486F0DF8" ID-GBIF-Occurrence="2564277958" box="[189,1316,692,719]" collectionCode="MSC" country="United States of America" location="Claiborne Group" pageId="176" pageNumber="177" specimenCount="1" stateProvince="Alabama">
<collectingCountry id="E8A9BCC7FF35FF1FFF7B9D814F1B0DF8" box="[189,592,693,719]" name="United States of America" pageId="176" pageNumber="177">UNITED STATES OF AMERICA</collectingCountry>
<emphasis id="A2CA2045FF35FF1FFDAB9D804F940DF9" bold="true" box="[621,735,692,718]" pageId="176" pageNumber="177">
<collectingRegion id="527A32B5FF35FF1FFDAB9D804F940DF9" box="[621,735,692,718]" country="United States of America" name="Alabama" pageId="176" pageNumber="177">Alabama</collectingRegion>
</emphasis>
• 1 isolated tooth;
<location id="9561AA8CFF35FF1FFC7A9D8049CD0DF8" LSID="urn:lsid:plazi:treatment:18174D41FF35FF1EFD7E9D154CF00D42:9561AA8CFF35FF1FFC7A9D8049CD0DF8" box="[956,1158,692,719]" country="United States of America" name="Claiborne Group" pageId="176" pageNumber="177" stateProvince="Alabama">Claiborne Group</location>
;
<collectionCode id="F6AF6492FF35FF1FFB549D81499B0DF8" box="[1170,1232,693,719]" pageId="176" pageNumber="177">MSC</collectionCode>
37271
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.
</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFF35FF1FFF7B9DC2490E0C9E" pageId="176" pageNumber="177" type="description">
<paragraph id="9001FC57FF35FF1FFF7B9DC24C050C27" blockId="176.[189,334,758,784]" box="[189,334,758,784]" pageId="176" pageNumber="177">
<heading id="CB494B3BFF35FF1FFF7B9DC24C050C27" bold="true" box="[189,334,758,784]" fontSize="11" level="3" pageId="176" pageNumber="177" reason="3">
<emphasis id="A2CA2045FF35FF1FFF7B9DC24C050C27" bold="true" box="[189,334,758,784]" pageId="176" pageNumber="177">Description</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FF35FF1FFF7B9C10490E0C9E" blockId="176.[189,1399,804,937]" pageId="176" pageNumber="177">Hypural plate roughly triangular in outline, consisting of fused hypurals, parahypural, and urostyle. Dorsal and ventral edges flat, straight, of equal length, wedge-shaped in dorsal and ventral views. Lateral faces flat; hypural sutures worn but visible; posterior margin slightly concave; basal notch nearly closed. Uroneural projecting dorsally from urostyle; face of urostyle ovoid, concave.</paragraph>
</subSubSection>
<subSubSection id="D8A4AFDCFF35FF1EFF7B9CE44CF00D42" lastPageId="177" lastPageNumber="178" pageId="176" pageNumber="177" type="discussion">
<paragraph id="9001FC57FF35FF1FFF7B9CE44C640CDD" blockId="176.[189,303,976,1002]" box="[189,303,976,1002]" pageId="176" pageNumber="177">
<heading id="CB494B3BFF35FF1FFF7B9CE44C640CDD" bold="true" box="[189,303,976,1002]" fontSize="11" level="3" pageId="176" pageNumber="177" reason="3">
<emphasis id="A2CA2045FF35FF1FFF7B9CE44C640CDD" bold="true" box="[189,303,976,1002]" pageId="176" pageNumber="177">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FF35FF1EFF7B9CCA493E0EEA" blockId="176.[189,1399,1022,1365]" lastBlockId="177.[189,1399,267,477]" lastPageId="177" lastPageNumber="178" pageId="176" pageNumber="177">
A diagnostic characteristic of the members of the
<taxonomicName id="57BE87D4FF35FF1FFD2A9CCA4E3E0B2F" authorityName="Rafinesque" authorityYear="1815" box="[748,885,1022,1048]" class="Actinopterygii" family="Scombridae" kingdom="Animalia" order="Perciformes" pageId="176" pageNumber="177" phylum="Chordata" rank="family">Scombridae</taxonomicName>
is the fusion of the hypurals and urostyle into a unique hypural plate (
<bibRefCitation id="F42F81A6FF35FF1FFE149B154F3C0B0C" author="Johnson G. D." box="[466,631,1057,1083]" pageId="176" pageNumber="177" pagination="1 - 41" refId="ref143857" refString="Johnson G. D. 1986. Scombroid phylogeny, an alternative hypothesis. Bulletin of Marine Science 39 (1): 1 - 41." type="journal article" year="1986">Johnson 1986</bibRefCitation>
;
<bibRefCitation id="F42F81A6FF35FF1FFD429B154E630B0B" author="Monsch K. A." box="[644,808,1057,1084]" pageId="176" pageNumber="177" pagination="445 - 489" refId="ref146740" refString="Monsch K. A. 2005. Revision of the scombroid fishes from the Cenozoic of England. Transactions of the Royal Society of Edinburgh, Earth Sciences 95: 445 - 489. https: // doi. org / 10.1017 / S 0263593300001164" type="journal article" year="2005">Monsch 2005</bibRefCitation>
). The hypural plate identified in our sample (MSC 37271;
<figureCitation id="0885E0D2FF35FF1FFED59B704C2F0B68" box="[275,356,1092,1119]" captionStart="Fig" captionStartId="176.[189,232,1964,1990]" captionTargetBox="[202,1315,1404,1924]" captionTargetId="figure@176.[189,1398,1400,1927]" captionTargetPageId="176" captionText="Fig. 65. Gymnosarda sp., hypural plate. AC. MSC 37271, basal Lisbon Formation. A. Right lateral view. B. Anterior view. C. Dorsal view. Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3660385" httpUri="https://zenodo.org/record/3660385/files/figure.png" pageId="176" pageNumber="177">Fig. 65</figureCitation>
) also possesses a fused parahypural and has a basal notch that is nearly closed. According to
<bibRefCitation id="F42F81A6FF35FF1FFF7B9B5C4C2D0BB5" author="Monsch K. A." box="[189,358,1128,1154]" pageId="176" pageNumber="177" pagination="75 - 80" refId="ref146693" refString="Monsch K. A. 2000. A new fossil bonito (Sardini, Teleostei) from the Eocene of England and the Caucasus, and evolution of tail region characters of its recent relatives. Paleontological Research 4 (1): 75 - 80." type="journal article" year="2000">Monsch (2000</bibRefCitation>
,
<bibRefCitation id="F42F81A6FF35FF1FFEB69B534CE60BB5" author="Monsch K. A." box="[368,429,1127,1154]" pageId="176" pageNumber="177" pagination="445 - 489" refId="ref146740" refString="Monsch K. A. 2005. Revision of the scombroid fishes from the Cenozoic of England. Transactions of the Royal Society of Edinburgh, Earth Sciences 95: 445 - 489. https: // doi. org / 10.1017 / S 0263593300001164" type="journal article" year="2005">2005</bibRefCitation>
),
<taxonomicName id="57BE87D4FF35FF1FFE069B5C4F380BB5" authorityName="Gill" authorityYear="1862" box="[448,627,1128,1154]" class="Actinopterygii" family="Scombridae" genus="Acanthocybium" kingdom="Animalia" order="Perciformes" pageId="176" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FF35FF1FFE069B5C4F380BB5" box="[448,627,1128,1154]" italics="true" pageId="176" pageNumber="177">Acanthocybium</emphasis>
</taxonomicName>
and the members of the
<taxonomicName id="57BE87D4FF35FF1FFC4E9B5C49100BB5" baseAuthorityName="Starks" baseAuthorityYear="1910" box="[904,1115,1128,1154]" class="Actinopterygii" family="Scombridae" kingdom="Animalia" order="Perciformes" pageId="176" pageNumber="177" phylum="Chordata" rank="subFamily" subFamily="Scomberomorinae">Scomberomorinae</taxonomicName>
have hypural plates with a basal notch that is large and distinct. However, a caudal notch is completely absent on the hypural plates of the members of the Tribe
<taxonomicName id="57BE87D4FF35FF1FFE3F9B9A4FB30BFF" authority="Starks, 1910" authorityName="Starks" authorityYear="1910" box="[505,760,1198,1224]" class="Actinopterygii" family="Scombridae" kingdom="Animalia" order="Perciformes" pageId="176" pageNumber="177" phylum="Chordata" rank="tribe" tribe="Thunnini">Thunnini Starks, 1910</taxonomicName>
, and nearly absent on those within the
<taxonomicName id="57BE87D4FF35FF1FFB6E9B9A4C0E0BDB" authority="Collette &amp; Chao, 1975" authorityName="Collette &amp; Chao" authorityYear="1975" class="Actinopterygii" family="Scombridae" kingdom="Animalia" order="Perciformes" pageId="176" pageNumber="177" phylum="Chordata" rank="tribe" tribe="Sardini">Sardini Collette &amp; Chao, 1975</taxonomicName>
. The presence of a reduced notch and fused parhypural on MSC 37271 are characteristic of a single member of
<taxonomicName id="57BE87D4FF35FF1FFE549BC04CAC0A39" authorityName="Collette &amp; Chao" authorityYear="1975" box="[402,487,1268,1294]" class="Actinopterygii" family="Scombridae" kingdom="Animalia" order="Perciformes" pageId="176" pageNumber="177" phylum="Chordata" rank="tribe" tribe="Sardini">Sardini</taxonomicName>
,
<taxonomicName id="57BE87D4FF35FF1FFE329BC04E4D0A39" authority="Gill, 1862" authorityName="Gill" authorityYear="1862" box="[500,774,1268,1294]" class="Actinopterygii" family="Scombridae" genus="Gymnosarda" kingdom="Animalia" order="Perciformes" pageId="176" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FF35FF1FFE329BC04FC30A39" box="[500,648,1268,1294]" italics="true" pageId="176" pageNumber="177">Gymnosarda</emphasis>
Gill, 1862
</taxonomicName>
(
<bibRefCitation id="F42F81A6FF35FF1FFCD19BC04EF10A38" author="Monsch K. A." box="[791,954,1268,1295]" pageId="176" pageNumber="177" pagination="75 - 80" refId="ref146693" refString="Monsch K. A. 2000. A new fossil bonito (Sardini, Teleostei) from the Eocene of England and the Caucasus, and evolution of tail region characters of its recent relatives. Paleontological Research 4 (1): 75 - 80." type="journal article" year="2000">Monsch 2000</bibRefCitation>
,
<bibRefCitation id="F42F81A6FF35FF1FFC019BC0494E0A38" author="Monsch K. A." box="[967,1029,1268,1295]" pageId="176" pageNumber="177" pagination="445 - 489" refId="ref146740" refString="Monsch K. A. 2005. Revision of the scombroid fishes from the Cenozoic of England. Transactions of the Royal Society of Edinburgh, Earth Sciences 95: 445 - 489. https: // doi. org / 10.1017 / S 0263593300001164" type="journal article" year="2005">2005</bibRefCitation>
). MSC 37271 is very similar to the hypural plate described on the extant
<taxonomicName id="57BE87D4FF35FF1FFD589A2C49100A05" authority="(Ruppell, 1838)" baseAuthorityName="Ruppell" baseAuthorityYear="1838" box="[670,1115,1304,1330]" class="Actinopterygii" family="Scombridae" genus="Gymnosarda" kingdom="Animalia" order="Perciformes" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="unicolor">
<emphasis id="A2CA2045FF35FF1FFD589A2C4ED70A05" box="[670,924,1304,1330]" italics="true" pageId="176" pageNumber="177">Gymnosarda unicolor</emphasis>
(Rüppell, 1838)
</taxonomicName>
and also very similar to the hypural of the only Eocene member of this genus,
<taxonomicName id="57BE87D4FF35FF1FFC859A0F49930A62" authority="Monsch, 2000" authorityName="Monsch" authorityYear="2000" box="[835,1240,1339,1365]" class="Actinopterygii" family="Scombridae" genus="Gymnosarda" kingdom="Animalia" order="Perciformes" pageId="176" pageNumber="177" phylum="Chordata" rank="species" species="prisca">
<emphasis id="A2CA2045FF35FF1FFC859A0F49630A62" box="[835,1064,1339,1365]" italics="true" pageId="176" pageNumber="177">Gymnosarda prisca</emphasis>
<bibRefCitation id="F42F81A6FF35FF1FFBF79A0F49930A62" author="Monsch K. A." box="[1073,1240,1339,1365]" pageId="176" pageNumber="177" pagination="75 - 80" refId="ref146693" refString="Monsch K. A. 2000. A new fossil bonito (Sardini, Teleostei) from the Eocene of England and the Caucasus, and evolution of tail region characters of its recent relatives. Paleontological Research 4 (1): 75 - 80." type="journal article" year="2000">Monsch, 2000</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="F42F81A6FF35FF1EFADF9A0F4DB00E12" author="Monsch K. A." lastPageId="177" lastPageNumber="178" pageId="176" pageNumber="177" pagination="75 - 80" refId="ref146693" refString="Monsch K. A. 2000. A new fossil bonito (Sardini, Teleostei) from the Eocene of England and the Caucasus, and evolution of tail region characters of its recent relatives. Paleontological Research 4 (1): 75 - 80." type="journal article" year="2000">Monsch 2000</bibRefCitation>
: fig. 1c, 2005: fig. 9c). However, the hypural plate of
<taxonomicName id="57BE87D4FF34FF1EFCA79E384E870E12" authorityName="Monsch" authorityYear="2000" box="[865,972,268,294]" class="Actinopterygii" family="Scombridae" genus="Gymnosarda" kingdom="Animalia" order="Perciformes" pageId="177" pageNumber="178" phylum="Chordata" rank="species" species="prisca">
<emphasis id="A2CA2045FF34FF1EFCA79E384E870E12" box="[865,972,268,294]" italics="true" pageId="177" pageNumber="178">G. prisca</emphasis>
</taxonomicName>
has a clear caudal notch, a purported plesiomorphic character within this genus.
<bibRefCitation id="F42F81A6FF34FF1EFD789E044E390E7D" author="Monsch K. A." box="[702,882,304,330]" pageId="177" pageNumber="178" pagination="75 - 80" refId="ref146693" refString="Monsch K. A. 2000. A new fossil bonito (Sardini, Teleostei) from the Eocene of England and the Caucasus, and evolution of tail region characters of its recent relatives. Paleontological Research 4 (1): 75 - 80." type="journal article" year="2000">Monsch (2000)</bibRefCitation>
described another Eocene
<taxonomicName id="57BE87D4FF34FF1EFB719E0448000E7D" authorityName="Gill" authorityYear="1862" box="[1207,1355,304,330]" class="Actinopterygii" family="Scombridae" genus="Gymnosarda" kingdom="Animalia" order="Perciformes" pageId="177" pageNumber="178" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FF34FF1EFB719E0448000E7D" box="[1207,1355,304,330]" italics="true" pageId="177" pageNumber="178">Gymnosarda</emphasis>
</taxonomicName>
sp. hypural plate that, although damaged along the posterior edge, exhibits a basal notch that appears nearly closed (see fig. 10) and is more elongated dorsoventrally than MSC 37271, suggesting the plates represent two different species. Although MSC 37271 likely represents an undescribed species within
<taxonomicName id="57BE87D4FF34FF1EFB1A9EAA483B0E8F" authorityName="Gill" authorityYear="1862" box="[1244,1392,414,440]" class="Actinopterygii" family="Scombridae" genus="Gymnosarda" kingdom="Animalia" order="Perciformes" pageId="177" pageNumber="178" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FF34FF1EFB1A9EAA483B0E8F" box="[1244,1392,414,440]" italics="true" pageId="177" pageNumber="178">Gymnosarda</emphasis>
</taxonomicName>
, we refrain from erecting one because our sample consist of only a single specimen.
</paragraph>
<caption id="C4C1ACDFFF35FF1FFF7B98984E2B08DE" ID-DOI="http://doi.org/10.5281/zenodo.3660385" ID-Zenodo-Dep="3660385" httpUri="https://zenodo.org/record/3660385/files/figure.png" pageId="176" pageNumber="177" startId="176.[189,232,1964,1990]" targetBox="[202,1315,1404,1924]" targetPageId="176">
<paragraph id="9001FC57FF35FF1FFF7B98984E2B08DE" blockId="176.[189,1398,1964,2025]" pageId="176" pageNumber="177">
<emphasis id="A2CA2045FF35FF1FFF7B98984C5708F1" bold="true" box="[189,284,1964,1990]" pageId="176" pageNumber="177">Fig. 65.</emphasis>
<taxonomicName id="57BE87D4FF35FF1FFEE398984CF008F1" authorityName="Gill" authorityYear="1862" box="[293,443,1964,1990]" class="Actinopterygii" family="Scombridae" genus="Gymnosarda" kingdom="Animalia" order="Perciformes" pageId="176" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="A2CA2045FF35FF1FFEE398984CF008F1" box="[293,443,1964,1990]" italics="true" pageId="176" pageNumber="177">Gymnosarda</emphasis>
</taxonomicName>
sp., hypural plate.
<emphasis id="A2CA2045FF35FF1FFD6298984F9608F1" bold="true" box="[676,733,1964,1990]" pageId="176" pageNumber="177">AC</emphasis>
.
<collectionCode id="F6AF6492FF35FF1FFD2898984E6708F1" box="[750,812,1964,1990]" pageId="176" pageNumber="177">MSC</collectionCode>
37271, basal Lisbon Formation.
<emphasis id="A2CA2045FF35FF1FFB7A9898499A08F1" bold="true" box="[1212,1233,1964,1990]" pageId="176" pageNumber="177">A</emphasis>
. Right lateral view.
<emphasis id="A2CA2045FF35FF1FFEC598FB4C5C08DE" bold="true" box="[259,279,1999,2025]" pageId="176" pageNumber="177">B</emphasis>
. Anterior view.
<emphasis id="A2CA2045FF35FF1FFE1298FB4CA208DE" bold="true" box="[468,489,1999,2025]" pageId="176" pageNumber="177">C</emphasis>
. Dorsal view. Scale bar = 1 cm.
</paragraph>
</caption>
<paragraph id="9001FC57FF34FF1EFF7B9D324E5F0D17" blockId="177.[189,788,518,544]" box="[189,788,518,544]" pageId="177" pageNumber="178">
<heading id="CB494B3BFF34FF1EFF7B9D324E5F0D17" bold="true" box="[189,788,518,544]" fontSize="11" level="3" pageId="177" pageNumber="178" reason="3">
<emphasis id="A2CA2045FF34FF1EFF7B9D324E5F0D17" bold="true" box="[189,788,518,544]" pageId="177" pageNumber="178">
Stratigraphic and geographic range in
<collectingRegion id="527A32B5FF34FF1EFD649D324E5F0D17" box="[674,788,518,544]" country="United States of America" name="Alabama" pageId="177" pageNumber="178">Alabama</collectingRegion>
</emphasis>
</heading>
</paragraph>
<paragraph id="9001FC57FF34FF1EFF7B9D024CF00D42" blockId="177.[189,1399,566,629]" pageId="177" pageNumber="178">
The lone specimen in our sample was collected from the basal
<collectingRegion id="527A32B5FF34FF1EFC569D024EAA0D67" box="[912,993,566,592]" country="Portugal" name="Lisboa" pageId="177" pageNumber="178">Lisbon</collectingRegion>
Formation at site ACov-11. Lower Lutetian, Zone NP15.
</paragraph>
</subSubSection>
</treatment>
</document>